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<document id="522DDAC9054D31438C47638C81DA94F0" ID-CLB-Dataset="22901" ID-DOI="10.11646/zootaxa.4763.3.4" ID-GBIF-Dataset="68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4" ID-ISSN="1175-5334" ID-Zenodo-Dep="3759527" ID-ZooBank="urn:lsid:zoobank.org:pub:721665BA-2B50-407E-9561-FBAD1CA161F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587482723970" checkinUser="carolina" docAuthor="Daniel, Gimo M., Deschodt, Christian M., Davis, Adrian L. V. &amp; Sole, Catherine L." docDate="2020" docId="03EABB021574A14710D88D43FF2945F2" docLanguage="en" docName="zootaxa.4763.3.4.pdf.imf" docOrigin="Zootaxa 4763 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stiptopodius muellerae Daniel &amp; Deschodt &amp; Davis &amp; Sole 2020, new species" docType="treatment" docVersion="8" lastPageNumber="402" masterDocId="FFD3C37A1572A14F104F8F7DFFD84314" masterDocTitle="The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa" masterLastPageNumber="404" masterPageNumber="394" pageNumber="400" updateTime="1698834361793" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0893CF68E1042E0FDFADF59DC18B1594" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6CF02916962632F19976E89989DC48B">
<mods:title id="65F61F84D5B38C1F91DD57D03ECD0DD4">The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="1D8F05462F9909510B59B4024311E410" type="personal">
<mods:role id="8A9222FE5FB8BE2235141C08A7171EBF">
<mods:roleTerm id="38B8BFCD167AF3FB5E0E62EBCF8C935C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2FBF58ACB4447CC2F4F614D12A752789">Daniel, Gimo M.</mods:namePart>
<mods:nameIdentifier id="1861252ACBA0A4B6137FBF0AA6D88983" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
<mods:affiliation id="664226D4B4C5804AD3F4736B4861AFCB">Department of Terrestria Invertebrates, The National Museum South Africal</mods:affiliation>
<mods:nameIdentifier id="468CB7FA812E3A56F0003D33FA3F7BD8" type="URL">https://orcid.org/0000-0003-3602-5034</mods:nameIdentifier>
<mods:nameIdentifier id="2550855471E81ED1B8041C0DB815FD55" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name id="263D6C7332980769063DD3EF9697204F" type="personal">
<mods:role id="7B1B7BD4CC9FA7A5E867EF3CADEFB3A3">
<mods:roleTerm id="0B473825A13E0460D4260FFBBF21B753">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8A2EEC0EFF9EBBB46690A6264270814">Deschodt, Christian M.</mods:namePart>
<mods:affiliation id="60C9C6445615DA28F77C8C1D0E078D4F">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="D36609390A192435FD6FC495A9D109A5" type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="6195143EE878FB6CB0E029E63F3C1D48" type="personal">
<mods:role id="A2DB62F17859F5D9091717F5D30EE6C3">
<mods:roleTerm id="2EB5B0824F7B80AD2D95B669560A6D8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52F8795D79F83CB3B12C1D1D6E8ED217">Davis, Adrian L. V.</mods:namePart>
<mods:affiliation id="F6AE002C40925D5849C08D0050AEF3AA">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="73ADC4106B2D15379BCA4ABA9C7547F0" type="email">adrian.davis04@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="350D1B217D38B0535F3792401D9762B3" type="personal">
<mods:role id="2E3CAE854D04A50AD479A74CC9A79AB7">
<mods:roleTerm id="59178BDEEFF6960D65EA0A1505147D83">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4DC83AA379B4084A3712F758BB85144">Sole, Catherine L.</mods:namePart>
<mods:affiliation id="9F1EB93DB0D457BD2990FFC94BE7D561">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="44612893B8D67EA3EDCA9C76F0A9F13D" type="email">catherine.sole@up.ac.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CA9DFDBC63791F87FE60170AD709B215">text</mods:typeOfResource>
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<mods:titleInfo id="BEC6FFC384BE1DAEB59F29025B8E8E7B">
<mods:title id="C4710EAED3AC7A1311D71ADE2CB6D7C8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FE586D86978A15B0510B9211FFFB2B6D">
<mods:date id="E66597E31314782F45BD31912D9F14CF">2020</mods:date>
<mods:detail id="CF57712D4235A87F407A3EBA99B1D25C" type="pubDate">
<mods:number id="D5BEABCB2E056A16885930FCECCEA214">2020-04-14</mods:number>
</mods:detail>
<mods:detail id="9B87C2CB9B6D67460477CA8D1DD5DE7B" type="volume">
<mods:number id="A24C8DDEBEA04ECC8A844F6E7A2D42FA">4763</mods:number>
</mods:detail>
<mods:detail id="F246C14E95417A18BB2B0579A2A7A0BD" type="issue">
<mods:number id="E66A2E7A507A8DCE9245A7819E154206">3</mods:number>
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<mods:classification id="045105ECF9AC2865684AB0C10FC4933D">journal article</mods:classification>
<mods:identifier id="9BD22B31A9DAD250A7CEDFC4D0968B36" type="CLB-Dataset">22901</mods:identifier>
<mods:identifier id="F7C87FDAFD15BC1F831002D604802F36" type="DOI">10.11646/zootaxa.4763.3.4</mods:identifier>
<mods:identifier id="23B7C82801DE0FFF4EDD51A42594BFFB" type="GBIF-Dataset">68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4</mods:identifier>
<mods:identifier id="A8E1F4B6703392D7025F7794EA66EB59" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="1008D9CBF587465531EC9262EC9682FB" type="Zenodo-Dep">3759527</mods:identifier>
<mods:identifier id="88116D9746C7DA51FA6E334365AD5DC9" type="ZooBank">urn:lsid:zoobank.org:pub:721665BA-2B50-407E-9561-FBAD1CA161F1</mods:identifier>
</mods:mods>
<treatment id="03EABB021574A14710D88D43FF2945F2" ID-DOI="http://doi.org/10.5281/zenodo.3805327" ID-GBIF-Taxon="163726936" ID-Zenodo-Dep="3805327" LSID="urn:lsid:plazi:treatment:03EABB021574A14710D88D43FF2945F2" httpUri="http://treatment.plazi.org/id/03EABB021574A14710D88D43FF2945F2" lastPageId="8" lastPageNumber="402" pageId="6" pageNumber="400">
<subSubSection id="C359599F1574A14910D88D43FCE7414C" box="[151,831,574,601]" pageId="6" pageNumber="400" type="nomenclature">
<paragraph id="8BFC0A141574A14910D88D43FCE7414C" blockId="6.[151,831,574,636]" box="[151,831,574,601]" pageId="6" pageNumber="400">
<heading id="D0B4BD781574A14910D88D43FCE7414C" bold="true" box="[151,831,574,601]" fontSize="10" level="2" pageId="6" pageNumber="400" reason="2">
<emphasis id="B937D6061574A14910D88D43FCE7414C" bold="true" box="[151,831,574,601]" pageId="6" pageNumber="400">
<taxonomicName id="4C4371971574A14910D88D43FD7B414D" ID-CoL="6ZW2N" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[151,675,574,601]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D6061574A14910D88D43FE75414D" bold="true" box="[151,429,574,601]" italics="true" pageId="6" pageNumber="400">Stiptopodius muellerae</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D1574A14912E08D42FCE7414C" box="[687,831,574,601]" pageId="6" pageNumber="400" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910D88D1FFE0C4168" box="[151,468,610,636]" pageId="6" pageNumber="400" type="description">
<paragraph id="8BFC0A141574A14910D88D1FFE0C4168" blockId="6.[151,831,574,636]" box="[151,468,610,636]" pageId="6" pageNumber="400">
(
<figureCitation id="137816911574A14910D08D1FFED94168" box="[159,257,610,636]" captionStart="FIGURES 1" captionStartId="3.[151,264,1877,1902]" captionTargetBox="[163,1426,206,1839]" captionTargetId="figure@3.[151,1436,198,1849]" captionTargetPageId="3" captionText="FIGURES 1A–D. Habitus of Stiptopodius Harold, 1871 species, holotypes. A, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; B, Stiptopodius savuti Daniel &amp; Deschodt, new species; C, Stiptopodius muellerae Daniel &amp; Deschodt, new species; D, Stiptopodius singularis (Péringuey, 1901). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759529" httpUri="https://zenodo.org/record/3759529/files/figure.png" pageId="6" pageNumber="400">Figs. 1C</figureCitation>
,
<figureCitation id="137816911574A14911428D1FFEC34168" box="[269,283,610,636]" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="6" pageNumber="400">2</figureCitation>
A–B, 3A, 3C, 4)
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910D88DD7FD8D41D0" box="[151,597,681,708]" pageId="6" pageNumber="400" type="distribution">
<paragraph id="8BFC0A141574A14910D88DD7FD8D41D0" blockId="6.[151,1437,681,2005]" box="[151,597,681,708]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910D88DD7FEE141D0" bold="true" box="[151,313,681,708]" pageId="6" pageNumber="400">
<typeStatus id="54F8B4B61574A14910D88DD7FF0A41D0" box="[151,210,682,708]" pageId="6" pageNumber="400">Type</typeStatus>
locality:
</emphasis>
Caconda,
<collectingRegion id="4987C4F61574A14911FF8DD7FE2941D0" box="[432,497,682,708]" country="Angola" name="Huila" pageId="6" pageNumber="400">Huila</collectingRegion>
,
<collectingCountry id="F3544A841574A14911B48DD7FD8941D0" box="[507,593,682,708]" name="Angola" pageId="6" pageNumber="400">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910888DB3FB514024" pageId="6" pageNumber="400" type="materials_examined">
<paragraph id="8BFC0A141574A14910888DB3FB514024" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888DB3FEAF41FC" bold="true" box="[199,375,717,744]" pageId="6" pageNumber="400">Type material.</emphasis>
<materialsCitation id="3B2B00491574A14911CF8DB3FB514024" ID-GBIF-Occurrence="2603207387" collectingDate="2011-08-11" collectionCode="TMSA" country="South Africa" county="Angola" latitude="-13.24" location="Daniel &amp; Deschodt" longLatPrecision="774" longitude="15.28" municipality="Stiptopodius" pageId="6" pageNumber="400" specimenCode="TMSC13817" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F8B4B61574A14911CF8DB3FE3341FC" box="[384,491,718,744]" pageId="6" pageNumber="400" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D45C19D1574A14911B68DB3FDEC41FC" box="[505,564,718,744]" pageId="6" pageNumber="400" type="male">male</specimenCount>
:
<collectingCounty id="629D72981574A14912708DB3FD4C41FC" box="[575,660,718,744]" pageId="6" pageNumber="400">Angola</collectingCounty>
, Huila Prov. |
<locationDeviation id="9A4E13361574A14913748DB0FC4541FC" bearing="180.0" box="[827,925,717,744]" distance="75000.0" pageId="6" pageNumber="400">
<quantity id="4CBBA7F11574A14913748DB0FC5C41FC" box="[827,900,717,744]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="400" unit="km" value="75.0">75 km</quantity>
S
</locationDeviation>
Kacopnda [Caconda] |
<geoCoordinate id="EE776CD31574A14914E28DB3FADD41FC" box="[1197,1285,718,744]" degrees="13.24" direction="south" orientation="latitude" pageId="6" pageNumber="400" precision="555" value="-13.24">13.24 S</geoCoordinate>
–
<geoCoordinate id="EE776CD31574A149156A8DB0FA5041FD" box="[1317,1416,717,745]" degrees="15. 28" direction="east" orientation="longitude" pageId="6" pageNumber="400" precision="555" value="15.28">15. 28 E</geoCoordinate>
||
<date id="FFFD2CD41574A14910D88D8FFED14018" box="[151,265,754,780]" pageId="6" pageNumber="400" value="2011-08-11">
<collectingDate id="EFB9D53C1574A14910D88D8FFED14018" box="[151,265,754,780]" pageId="6" pageNumber="400" value="2011-08-11">8.11.2011</collectingDate>
</date>
;E-Y: 3916 | sandy forest,
<quantity id="4CBBA7F11574A14912798D8FFD5C4018" box="[566,644,754,780]" metricMagnitude="3" metricUnit="m" metricValue="1.643" pageId="6" pageNumber="400" unit="m" value="1643.0">1643m</quantity>
| leg. Ruth Müller || TM
<collectingCountry id="F3544A841574A14913E38D8FFBAE4018" box="[940,1142,754,780]" name="South Africa" pageId="6" pageNumber="400">SOUTH AFRICA</collectingCountry>
|
<specimenCode id="DBE5A26F1574A14914C18D8FFAF94018" box="[1166,1313,754,780]" pageId="6" pageNumber="400">TMSC13817</specimenCode>
||
<emphasis id="B937D6061574A14915708D8CFF334024" bold="true" pageId="6" pageNumber="400">HOLOTYPE:</emphasis>
<taxonomicName id="4C4371971574A14910BD8C6BFEA64024" authorityName="Harold" authorityYear="1871" box="[242,382,790,816]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="genus">
<emphasis id="B937D6061574A14910BD8C6BFEA64024" box="[242,382,790,816]" italics="true" pageId="6" pageNumber="400">
<collectingMunicipality id="6B98906E1574A14910BD8C6BFEA64024" box="[242,382,790,816]" pageId="6" pageNumber="400">Stiptopodius</collectingMunicipality>
</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4371971574A14911DD8C6BFD394024" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[402,737,790,816]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="species" species="muellerae">
<emphasis id="B937D6061574A14911DD8C6BFDDA4024" box="[402,514,790,816]" italics="true" pageId="6" pageNumber="400">muellerae</emphasis>
<location id="8E9C5CCF1574A14912468C6BFD394024" LSID="urn:lsid:plazi:treatment:03EABB021574A14710D88D43FF2945F2:8E9C5CCF1574A14912468C6BFD394024" box="[521,737,790,816]" country="South Africa" county="Angola" latitude="-13.24" longLatPrecision="774" longitude="15.28" municipality="Stiptopodius" name="Daniel &amp; Deschodt" pageId="6" pageNumber="400">Daniel &amp; Deschodt</location>
</taxonomicName>
| 2020 (printed on red card) (
<collectionCode id="ED5292D11574A14914668C6BFBA54024" box="[1065,1149,790,816]" collectionName="South Africa, Gauteng, Pretoria, Transvaal Museum" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="400">TMSA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910888C44FE424450" pageId="6" pageNumber="400" type="description">
<paragraph id="8BFC0A141574A14910888C44FC504041" blockId="6.[151,1437,681,2005]" box="[199,904,825,853]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888C44FE814040" bold="true" box="[199,345,825,852]" pageId="6" pageNumber="400">Description.</emphasis>
<typeStatus id="54F8B4B61574A149112F8C47FE134040" box="[352,459,826,852]" pageId="6" pageNumber="400" type="holotype">Holotype</typeStatus>
, male. Length:
<quantity id="4CBBA7F11574A14912378C47FD084041" box="[632,720,826,853]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="400" unit="mm" value="7.6">7.6 mm</quantity>
, width:
<quantity id="4CBBA7F11574A14913648C47FC5C4041" box="[811,900,826,853]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="400" unit="mm" value="3.8">3.8 mm</quantity>
.
</paragraph>
<paragraph id="8BFC0A141574A14910888C20FEA8406C" blockId="6.[151,1437,681,2005]" box="[199,368,861,888]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888C20FEF8406C" bold="true" box="[199,288,861,888]" pageId="6" pageNumber="400">Colour.</emphasis>
Black.
</paragraph>
<paragraph id="8BFC0A141574A14910888CFFFABF471D" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888CFFFED64088" bold="true" box="[199,270,898,924]" pageId="6" pageNumber="400">Head.</emphasis>
Incision between medial clypeal teeth deeply concave, lateral clypeal edges not sinuate; external protrusion of genae sharply triangular; surface of head with green sheen; punctation varying from fine to large; setae well-developed except on glabrous anterior clypeal edge; clypeo-frontal carina visible; clypeo-genal suture present; vertex with small medial carina. Antennae with eight antennomeres; labium bears a few well-developed setae.
</paragraph>
<paragraph id="8BFC0A141574A14910888B6FFE5E478C" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888B6FFE9F4738" bold="true" box="[199,327,1042,1068]" pageId="6" pageNumber="400">Pronotum.</emphasis>
Convex, wider than long; base of pronotum marginate; anterior margin wider than posterior; in lateral view, lateral edge strongly sinuate before postero-lateral angles, surface with green metallic sheen, ocellate punctation varying from fine to large, each side with small longitudinal tubercle; well-developed setae, except on glabrous central disc.
</paragraph>
<paragraph id="8BFC0A141574A14910888BDFFC2047F5" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888BDFFEC247A8" bold="true" box="[199,282,1186,1212]" pageId="6" pageNumber="400">Elytra.</emphasis>
Lateral margin straight, somewhat convex distally and concave proximally, well-developed humeral umbone; striae sparsely and finely punctate. Interstriae punctate, fully setose.
</paragraph>
<paragraph id="8BFC0A141574A14910888B94FCDC4610" blockId="6.[151,1437,681,2005]" box="[199,772,1257,1285]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888B94FE9A4610" bold="true" box="[199,322,1257,1284]" pageId="6" pageNumber="400">Pygidium.</emphasis>
Convex and setose, punctation ocellate.
</paragraph>
<paragraph id="8BFC0A141574A14910888A70FBAF4681" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888A70FE1E463C" bold="true" box="[199,454,1293,1320]" pageId="6" pageNumber="400">Abdominal ventrites.</emphasis>
Prosternum surface with well-developed setae, anterior angles of prosternum slanted forward and inward. Meso- and metaventrite dull, setose and densely punctate; metaventrite with long setae, surface impressed longitudinally at middle. Abdominal ventrites with ocellate punctures and strong setae arranged in rows laterally, glabrous medially; last abdominal ventrite narrowed medially (male character).
</paragraph>
<paragraph id="8BFC0A141574A14910888AE3FE6A45E8" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888AE3FEDE46AC" bold="true" box="[199,262,1438,1464]" pageId="6" pageNumber="400">Legs.</emphasis>
Profemora sparsely punctate, bearing well-developed setae, external edge strongly marginate, internal margin slightly marginate. Meso- and metafemora with external and internal edges sparsely setigerous and punctate; surface between external and internal margin with few setigerous punctures, strongly marginate in both edges (external and internal). Protibiae crenulate with four teeth on external margin, line of setae around teeth in dorsal view; in ventral view, with two carinae, internally punctate and pilose. Protibial spur long and inward curving. Meso- and metatibiae triangular and strongly widened apically. Protarsi with five tarsomeres and clearly visible pair of claws; fifth protarsomere at least 2x longer than others (each individually). Meso- and metatarsomeres I–IV triangular, longer than wide, with internal line of strong setae on tarsomere I, strongly concave medially, bilobate distally, external lobe strong, internal lobe weak; tarsomere V rectangular and longer than others, followed by pair of very tiny apical claws.
</paragraph>
<paragraph id="8BFC0A141574A14910888878FBD04434" blockId="6.[151,1437,681,2005]" box="[199,1032,1797,1825]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888878FE994434" bold="true" box="[199,321,1797,1824]" pageId="6" pageNumber="400">Aedeagus.</emphasis>
Parameres symmetrical, elongate with tips acute (
<figureCitation id="137816911574A149133A887BFC1A4434" box="[885,962,1798,1824]" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="6" pageNumber="400">Figs. 2</figureCitation>
A–B).
</paragraph>
<paragraph id="8BFC0A141574A14910888857FE424450" blockId="6.[151,1437,681,2005]" box="[199,410,1834,1860]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888857FEFD4450" bold="true" box="[199,293,1834,1860]" pageId="6" pageNumber="400">Female.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910888830FE354498" pageId="6" pageNumber="400" type="distribution">
<paragraph id="8BFC0A141574A14910888830FE354498" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888830FEB9447C" bold="true" box="[199,353,1869,1896]" pageId="6" pageNumber="400">Distribution.</emphasis>
The species is only known from the
<typeStatus id="54F8B4B61574A14913428832FCE5447D" box="[781,829,1871,1897]" pageId="6" pageNumber="400">type</typeStatus>
locality, Caconda, Province of
<collectingRegion id="4987C4F61574A14914E78833FB31447C" box="[1192,1257,1870,1896]" country="Angola" name="Huila" pageId="6" pageNumber="400">Huila</collectingRegion>
,
<collectingCountry id="F3544A841574A14914BB8833FA92447C" box="[1268,1354,1870,1896]" name="Angola" pageId="6" pageNumber="400">Angola</collectingCountry>
. It was hand collected in sandy forest.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A149108888EBFB3F44C0" pageId="6" pageNumber="400" type="etymology">
<paragraph id="8BFC0A141574A149108888EBFB3F44C0" blockId="6.[151,1437,681,2005]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A149108888EBFE9644A4" bold="true" box="[199,334,1942,1968]" pageId="6" pageNumber="400">Etymology.</emphasis>
The specific name honours Ruth Müller, a friend and curator in charge of
<taxonomicName id="4C4371971574A14914CE88EBFB2444A4" authorityName="Linnaeus" authorityYear="1758" box="[1153,1276,1942,1968]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
at the Ditsong National Museum of Natural History,
<collectingRegion id="4987C4F61574A149120E88C7FD7B44C0" box="[577,675,1978,2004]" country="South Africa" name="Gauteng" pageId="6" pageNumber="400">Gauteng</collectingRegion>
, Pretoria,
<collectingCountry id="F3544A841574A149135A88C7FC7D44C0" box="[789,933,1978,2004]" name="South Africa" pageId="6" pageNumber="400">South Africa</collectingCountry>
, who collected the
<typeStatus id="54F8B4B61574A149143288C7FB3B44C0" box="[1149,1251,1978,2004]" pageId="6" pageNumber="400" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF3C5A9C1575A14810D888E5FA9F44C0" ID-DOI="http://doi.org/10.5281/zenodo.3759533" ID-Zenodo-Dep="3759533" httpUri="https://zenodo.org/record/3759533/files/figure.png" pageId="7" pageNumber="401" startId="7.[151,264,1944,1969]" targetBox="[339,1231,187,1911]" targetPageId="7">
<paragraph id="8BFC0A141575A14810D888E5FA9F44C0" blockId="7.[151,1436,1944,2005]" pageId="7" pageNumber="401">
<emphasis id="B937D6061575A14810D888E5FE8B44A5" bold="true" box="[151,339,1944,1969]" pageId="7" pageNumber="401">FIGURES 3A–D.</emphasis>
Left middle (A, B) and left hind tibiae and tarsi (C, D) of
<taxonomicName id="4C4371971575A14813D188E4FB7144A5" authority="Harold, 1871" authorityName="Harold" authorityYear="1871" box="[926,1193,1944,1969]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="401" phylum="Arthropoda" rank="genus">
<emphasis id="B937D6061575A14813D188E4FBC544A5" box="[926,1053,1945,1969]" italics="true" pageId="7" pageNumber="401">Stiptopodius</emphasis>
<bibRefCitation id="EFD277E51575A148146C88E5FB7144A5" author="Harold" box="[1059,1193,1944,1969]" firstAuthor="Harold" pageId="7" pageNumber="401" pagination="1 - 28" refId="ref7328" refString="Harold, E. von (1871) Verzeichniss der von Dr Beccari in Bogos gesammelten Coprophagen Lamellicornien. Coleopterologische Hefte, 8, 1 - 28. https: // doi. org / 10.1002 / mmnd. 47918710229" type="journal article" year="1871">Harold, 1871</bibRefCitation>
</taxonomicName>
species. A, C,
<taxonomicName id="4C4371971575A148157088E4FE2144C1" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="401" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D6061575A148157088E4FEF544C1" italics="true" pageId="7" pageNumber="401">Stiptopodius muellerae</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D1575A148124B88C0FD5844C1" box="[516,640,1980,2005]" pageId="7" pageNumber="401" rank="species">new species</taxonomicNameLabel>
; B, D,
<taxonomicName id="4C4371971575A148128888C0FBA844C1" authority="(Peringuey, 1901)" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[711,1136,1980,2005]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="401" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D6061575A148128888C0FC6944C1" box="[711,945,1981,2005]" italics="true" pageId="7" pageNumber="401">Stiptopodius singularis</emphasis>
(
<bibRefCitation id="EFD277E51575A14813F088C1FBB044C1" author="Peringuey" box="[959,1128,1980,2005]" firstAuthor="Peringuey" pageId="7" pageNumber="401" pagination="1 - 920" refId="ref7395" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12, 1 - 920. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
)
</taxonomicName>
. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF3C5A9C157AA14710D88B38FB684796" ID-DOI="http://doi.org/10.5281/zenodo.3759535" ID-Zenodo-Dep="3759535" httpUri="https://zenodo.org/record/3759535/files/figure.png" pageId="8" pageNumber="402" startId="8.[151,250,1093,1118]" targetBox="[151,1412,151,1074]" targetPageId="8">
<paragraph id="8BFC0A14157AA14710D88B38FB684796" blockId="8.[151,1437,1093,1154]" pageId="8" pageNumber="402">
<emphasis id="B937D606157AA14710D88B38FED4474A" bold="true" box="[151,268,1093,1118]" pageId="8" pageNumber="402">FIGURE 4</emphasis>
. Map showing the type localities of
<taxonomicName id="4C437197157AA147123A8B3BFBFE474A" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[629,1062,1093,1118]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157AA147123A8B3BFC85474A" box="[629,861,1094,1118]" italics="true" pageId="8" pageNumber="402">Stiptopodius muellerae</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D157AA14714608B3AFB70474A" box="[1071,1192,1093,1118]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
(.),
<taxonomicName id="4C437197157AA14714938B3BFE854796" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D606157AA14714938B3BFA45474A" box="[1244,1437,1094,1118]" italics="true" pageId="8" pageNumber="402">Stiptopodius savuti</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D157AA14711288B16FE394796" box="[359,481,1129,1154]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
(
<emphasis id="B937D606157AA14711BF8B15FDD14794" bold="true" box="[496,521,1128,1152]" pageId="8" pageNumber="402">A</emphasis>
) and
<taxonomicName id="4C437197157AA147120D8B17FBD84796" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[578,1024,1129,1154]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D606157AA147120D8B17FCEB4796" box="[578,819,1130,1154]" italics="true" pageId="8" pageNumber="402">Stiptopodius peringueyi</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D157AA14714458B16FB5C4796" box="[1034,1156,1129,1154]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
(■).
</paragraph>
</caption>
<subSubSection id="C359599F157AA14710888BCDFC7546D2" pageId="8" pageNumber="402" type="diagnosis">
<paragraph id="8BFC0A14157AA14710888BCDFC7546D2" blockId="8.[151,1437,1200,1766]" pageId="8" pageNumber="402">
<emphasis id="B937D606157AA14710888BCDFE9A47DF" bold="true" box="[199,322,1200,1227]" pageId="8" pageNumber="402">Diagnosis.</emphasis>
The new species is somewhat similar to
<taxonomicName id="4C437197157AA147135B8BCCFC7947DE" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[788,929,1200,1227]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157AA147135B8BCCFC7947DE" box="[788,929,1200,1227]" italics="true" pageId="8" pageNumber="402">S. singularis</emphasis>
</taxonomicName>
by having the basal half of the pronotal disc glabrous and frontal carina obsolete. However,
<taxonomicName id="4C437197157AA14712F18BA8FC9547FA" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[702,845,1236,1263]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157AA14712F18BA8FC9547FA" box="[702,845,1236,1263]" italics="true" pageId="8" pageNumber="402">S. singularis</emphasis>
</taxonomicName>
bears uniform punctation on the pronotum, with a brown metallic sheen, whereas
<taxonomicName id="4C437197157AA14711B28B84FD534606" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[509,651,1272,1299]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157AA14711B28B84FD534606" box="[509,651,1272,1299]" italics="true" pageId="8" pageNumber="402">S. muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D157AA14712DC8B84FCC34606" box="[659,795,1272,1299]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
has a distinctive green metallic sheen with punctures of variable sizes ranging from fine to large. The elytra of this new species are fully setose, while those of
<taxonomicName id="4C437197157AA147155F8A60FA444622" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[1296,1436,1308,1335]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157AA147155F8A60FA444622" box="[1296,1436,1308,1335]" italics="true" pageId="8" pageNumber="402">S. singularis</emphasis>
</taxonomicName>
are slightly setose posteriorly and glabrous anteriorly. The mesotibiae are sinuate externally in
<taxonomicName id="4C437197157AA14714988A3CFABD464E" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[1239,1381,1344,1371]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157AA14714988A3CFABD464E" box="[1239,1381,1344,1371]" italics="true" pageId="8" pageNumber="402">S. muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D157AA14715218A3CFF30466A" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
but not in
<emphasis id="B937D606157AA14711138A18FE36466A" box="[348,494,1380,1407]" italics="true" pageId="8" pageNumber="402">
<taxonomicName id="4C437197157AA14711138A18FE33466A" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[348,491,1380,1407]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">S. singularis</taxonomicName>
.
</emphasis>
The male genitalia differ between the two species by having the tips of the parameres sharp and projecting forward in the new species (
<figureCitation id="13781691157AA14712FB8AF5FD2746B7" box="[692,767,1416,1443]" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="8" pageNumber="402">Figs. 2</figureCitation>
A-B), whereas in
<taxonomicName id="4C437197157AA147138C8AF4FB9546B6" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[963,1101,1416,1443]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D606157AA147138C8AF4FB9546B6" box="[963,1101,1416,1443]" italics="true" pageId="8" pageNumber="402">S. singularis</emphasis>
</taxonomicName>
they are short and blunt (
<figureCitation id="13781691157AA147152B8AF5FF7D46D3" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="8" pageNumber="402">Figs. 2</figureCitation>
E-F).
<taxonomicName id="4C437197157AA14710A88AD0FEAB46D2" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[231,371,1452,1479]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157AA14710A88AD0FEAB46D2" box="[231,371,1452,1479]" italics="true" pageId="8" pageNumber="402">S. muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D157AA14711368AD0FDD846D2" box="[377,512,1452,1479]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
is also much larger than
<emphasis id="B937D606157AA14713568AD0FC7546D2" box="[793,941,1452,1479]" italics="true" pageId="8" pageNumber="402">
<taxonomicName id="4C437197157AA14713568AD0FC7146D2" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[793,937,1452,1479]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="singularis">S. singularis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C359599F157AA14710888AACFE094542" pageId="8" pageNumber="402" type="discussion">
<paragraph id="8BFC0A14157AA14710888AACFE094542" blockId="8.[151,1437,1200,1766]" pageId="8" pageNumber="402">
<emphasis id="B937D606157AA14710888AACFEE346FF" bold="true" box="[199,315,1489,1515]" pageId="8" pageNumber="402">Remarks.</emphasis>
This is the first record of a
<taxonomicName id="4C437197157AA14712348AADFCDF46FE" authorityName="Harold" authorityYear="1871" box="[635,775,1488,1514]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B937D606157AA14712348AADFCDF46FE" box="[635,775,1488,1514]" italics="true" pageId="8" pageNumber="402">Stiptopodius</emphasis>
</taxonomicName>
species whose meso- and metatarsomeres I–IV are longer than wide (
<figureCitation id="13781691157AA14711588A89FE78451B" box="[279,416,1524,1551]" captionStart="FIGURES 3" captionStartId="7.[151,264,1944,1969]" captionTargetBox="[339,1231,187,1911]" captionTargetId="figure@7.[335,1251,181,1919]" captionTargetPageId="7" captionText="FIGURES 3A–D. Left middle (A, B) and left hind tibiae and tarsi (C, D) of Stiptopodius Harold, 1871 species. A, C, Stiptopodius muellerae Daniel &amp; Deschodt, new species; B, D, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759533" httpUri="https://zenodo.org/record/3759533/files/figure.png" pageId="8" pageNumber="402">Figs. 3A, C</figureCitation>
). The first tarsomere is the longest, which is in contrast to the other 5-articulated species, where tarsomere V is longer than others. Furthermore, in
<taxonomicName id="4C437197157AA14713578964FC454526" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[792,925,1560,1587]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="402" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B937D606157AA14713578964FC454526" box="[792,925,1560,1587]" italics="true" pageId="8" pageNumber="402">S muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D157AA14713EC8964FBF54526" box="[931,1069,1560,1587]" pageId="8" pageNumber="402" rank="species">new species</taxonomicNameLabel>
, meso- and metatarsomeres I–IV bear a weak lobe externally.
</paragraph>
</subSubSection>
<subSubSection id="C359599F157AA1471088891DFF2945F2" pageId="8" pageNumber="402" type="conservation">
<paragraph id="8BFC0A14157AA1471088891DFF2945F2" blockId="8.[151,1437,1200,1766]" pageId="8" pageNumber="402">
<emphasis id="B937D606157AA1471088891DFD67456F" bold="true" box="[199,703,1632,1659]" pageId="8" pageNumber="402">Conservation status and potential threats.</emphasis>
The forest at the
<typeStatus id="54F8B4B6157AA14713C7891CFC60456F" box="[904,952,1633,1659]" pageId="8" pageNumber="402">type</typeStatus>
locality is surrounded by lands supporting agricultural and pastoral farming activities (R. Müller, personal communication). It is therefore probably under severe pressure. As this species is only known from this locality, it may warrant the threat category of at least Vulnerable.
</paragraph>
</subSubSection>
</treatment>
</document>
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