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<document id="BAB5AAFBF3D3FE9E45C4E6112B95A349" ID-CLB-Dataset="22901" ID-DOI="10.11646/zootaxa.4763.3.4" ID-GBIF-Dataset="68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4" ID-ISSN="1175-5334" ID-Zenodo-Dep="3759527" ID-ZooBank="urn:lsid:zoobank.org:pub:721665BA-2B50-407E-9561-FBAD1CA161F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587482723970" checkinUser="carolina" docAuthor="Daniel, Gimo M., Deschodt, Christian M., Davis, Adrian L. V. &amp; Sole, Catherine L." docDate="2020" docId="03EABB021573A14D10D88930FCD14483" docLanguage="en" docName="zootaxa.4763.3.4.pdf.imf" docOrigin="Zootaxa 4763 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stiptopodius peringueyi Daniel &amp; Deschodt &amp; Davis &amp; Sole 2020, new species" docType="treatment" docVersion="7" lastPageNumber="396" masterDocId="FFD3C37A1572A14F104F8F7DFFD84314" masterDocTitle="The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa" masterLastPageNumber="404" masterPageNumber="394" pageNumber="395" updateTime="1698834361793" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="06383B72C551C8CAE7B3BA72420999AA">The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa</mods:title>
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<mods:namePart id="F58B19754925DE6BEDD7A83A103EDBD6">Daniel, Gimo M.</mods:namePart>
<mods:nameIdentifier id="792652B1B1F69FFACE4F6D98446A976C" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
<mods:affiliation id="8D60EDA18B4C8479B1BD533EC430597F">Department of Terrestria Invertebrates, The National Museum South Africal</mods:affiliation>
<mods:nameIdentifier id="DD962A86D84EDDFC093DE05228886020" type="URL">https://orcid.org/0000-0003-3602-5034</mods:nameIdentifier>
<mods:nameIdentifier id="29036444313A6C9A4066045BC758FC75" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name id="7BBF53169DD8F2B5C768895A09BFC756" type="personal">
<mods:role id="17E95EAA0A917AA163E2702C0B2DFDF3">
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<mods:namePart id="CFD4053FF4F7E74F149875AE68AED70A">Deschodt, Christian M.</mods:namePart>
<mods:affiliation id="B88BB1926DBF2B70F522CCF6073C20D2">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="FD86EEBDD45D5AEDD67924B5E61BC63F" type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
</mods:name>
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<mods:role id="BD6341451CF0BEEA0E8B63EF9648F17F">
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<mods:namePart id="8A3E30F2BF78CC1302501E5BD567A96C">Davis, Adrian L. V.</mods:namePart>
<mods:affiliation id="6E9EC69FF0501A129ED7AF9CF189EEA8">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="7A63BA47A7960748AB07E3E01CF35233" type="email">adrian.davis04@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="98DBD2B7250E55EC9B57BFF8AE4F0FA9" type="personal">
<mods:role id="C347F5BCC88E2BA787C65389E4D09854">
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<mods:namePart id="6368A6862DBCF2DFD2836DE3AB37ADD0">Sole, Catherine L.</mods:namePart>
<mods:affiliation id="0C7B645671964B059F58D25C8ED3D08A">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="61DF1DC992A90CA667810959D9AF515E" type="email">catherine.sole@up.ac.za</mods:nameIdentifier>
</mods:name>
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<mods:title id="0698C2CB5CCF4602726A4215865AB223">Zootaxa</mods:title>
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<mods:part id="DEB513C1E1DD8DE5B93A4305A54B90FE">
<mods:date id="1E7F65BB6A56A90EE43322FD142B6E14">2020</mods:date>
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<mods:number id="F9DA05E0710BB161F60E85408C48A98E">2020-04-14</mods:number>
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<mods:number id="D5220FDE73CEFD9B4CFE361E35A69EE7">4763</mods:number>
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<mods:classification id="5C8047F78B60A5C0A16FC545D2C55CD5">journal article</mods:classification>
<mods:identifier id="6CAD12D3D44216E51F417FF9D5D7BE3A" type="CLB-Dataset">22901</mods:identifier>
<mods:identifier id="E0EEF1C84A5F34107973C524675DADB9" type="DOI">10.11646/zootaxa.4763.3.4</mods:identifier>
<mods:identifier id="A657A3440DC22E26B419E45419B6B899" type="GBIF-Dataset">68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4</mods:identifier>
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<treatment id="03EABB021573A14D10D88930FCD14483" ID-GBIF-Taxon="163726939" LSID="urn:lsid:plazi:treatment:03EABB021573A14D10D88930FCD14483" httpUri="http://treatment.plazi.org/id/03EABB021573A14D10D88930FCD14483" lastPageId="2" lastPageNumber="396" pageId="1" pageNumber="395">
<subSubSection id="C359599F1573A14E10D88930FC9F4573" box="[151,839,1613,1640]" pageId="1" pageNumber="395" type="nomenclature">
<paragraph id="8BFC0A141573A14E10D88930FC9F4573" blockId="1.[151,839,1613,1675]" box="[151,839,1613,1640]" pageId="1" pageNumber="395">
<heading id="D0B4BD781573A14E10D88930FC9F4573" bold="true" box="[151,839,1613,1640]" fontSize="10" level="2" pageId="1" pageNumber="395" reason="2">
<emphasis id="B937D6061573A14E10D88930FC9F4573" bold="true" box="[151,839,1613,1640]" pageId="1" pageNumber="395">
<taxonomicName id="4C4371971573A14E10D88930FD734573" ID-CoL="6ZVQK" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[151,683,1613,1640]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D6061573A14E10D88930FE6D4573" bold="true" box="[151,437,1613,1640]" italics="true" pageId="1" pageNumber="395">Stiptopodius peringueyi</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D1573A14E12F78930FC9F4573" box="[696,839,1613,1639]" pageId="1" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359599F1573A14E10D8890CFEB6459F" box="[151,366,1649,1675]" pageId="1" pageNumber="395" type="description">
<paragraph id="8BFC0A141573A14E10D8890CFEB6459F" blockId="1.[151,839,1613,1675]" box="[151,366,1649,1675]" pageId="1" pageNumber="395">
(
<figureCitation id="137816911573A14E10D0890CFEDB459F" box="[159,259,1649,1675]" captionStart="FIGURES 1" captionStartId="3.[151,264,1877,1902]" captionTargetBox="[163,1426,206,1839]" captionTargetId="figure@3.[151,1436,198,1849]" captionTargetPageId="3" captionText="FIGURES 1A–D. Habitus of Stiptopodius Harold, 1871 species, holotypes. A, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; B, Stiptopodius savuti Daniel &amp; Deschodt, new species; C, Stiptopodius muellerae Daniel &amp; Deschodt, new species; D, Stiptopodius singularis (Péringuey, 1901). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759529" httpUri="https://zenodo.org/record/3759529/files/figure.png" pageId="1" pageNumber="395">Figs. 1A</figureCitation>
,
<figureCitation id="137816911573A14E1140890CFEC4459F" box="[271,284,1649,1675]" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="1" pageNumber="395">2</figureCitation>
E–F, 4)
</paragraph>
</subSubSection>
<subSubSection id="C359599F1573A14E10D889C4FCE645C7" box="[151,830,1720,1747]" pageId="1" pageNumber="395" type="distribution">
<paragraph id="8BFC0A141573A14E10D889C4FCE645C7" blockId="1.[151,1437,1720,2035]" box="[151,830,1720,1747]" pageId="1" pageNumber="395">
<emphasis id="B937D6061573A14E10D889C4FEE845C7" bold="true" box="[151,304,1720,1747]" pageId="1" pageNumber="395">
<typeStatus id="54F8B4B61573A14E10D889C4FF0A45C7" box="[151,210,1721,1747]" pageId="1" pageNumber="395">Type</typeStatus>
locality
</emphasis>
: Richard’s Bay,
<collectingRegion id="4987C4F61573A14E11A589C4FD4545C7" box="[490,669,1721,1747]" country="South Africa" name="KwaZulu-Natal" pageId="1" pageNumber="395">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="F3544A841573A14E12E789C4FCE245C7" box="[680,826,1721,1747]" name="South Africa" pageId="1" pageNumber="395">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1573A14E108889A0FA5844E7" pageId="1" pageNumber="395" type="materials_examined">
<paragraph id="8BFC0A141573A14E108889A0FA5844E7" blockId="1.[151,1437,1720,2035]" pageId="1" pageNumber="395">
<emphasis id="B937D6061573A14E108889A0FEAD45E3" bold="true" box="[199,373,1756,1783]" pageId="1" pageNumber="395">Type material.</emphasis>
<materialsCitation id="3B2B00491573A14E113289A0FAF44477" ID-GBIF-Occurrence="2603207389" collectedFrom="Open shrubland" collectingDate="2000-01-27" collectingMethod="dung-baited pitfall trap" collectionCode="KZN, SANC" collectorName="Davis &amp; Delport" country="South Africa" latitude="28.6160000°" location="Richard's Bay" longLatPrecision="1" longitude="032.2931667°" municipality="Bay" pageId="1" pageNumber="395" specimenCode="TYPH02476" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<typeStatus id="54F8B4B61573A14E113289A0FE3045E3" box="[381,488,1757,1783]" pageId="1" pageNumber="395" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D45C19D1573A14E11BC89A0FDF645E3" box="[499,558,1757,1783]" pageId="1" pageNumber="395" type="male">male</specimenCount>
:
<collectingCountry id="F3544A841573A14E127689A0FCDE45E3" box="[569,774,1757,1783]" name="South Africa" pageId="1" pageNumber="395">SOUTH AFRICA</collectingCountry>
,
<collectionCode id="ED5292D11573A14E135E89A0FC9345E3" box="[785,843,1757,1783]" collectionName="KZN" pageId="1" pageNumber="395">KZN</collectionCode>
[
<collectingRegion id="4987C4F61573A14E131689A1FBD745E3" box="[857,1039,1756,1783]" country="South Africa" name="KwaZulu-Natal" pageId="1" pageNumber="395">KwaZulu-Natal</collectingRegion>
] |
<location id="8E9C5CCF1573A14E146689A0FB1145E3" LSID="urn:lsid:plazi:treatment:03EABB021573A14D10D88930FCD14483:8E9C5CCF1573A14E146689A0FB1145E3" box="[1065,1225,1757,1783]" country="South Africa" latitude="28.6160000°" longLatPrecision="1" longitude="032.2931667°" municipality="Bay" name="Richard's Bay" pageId="1" pageNumber="395" stateProvince="KwaZulu-Natal">Richard’s Bay</location>
||
<geoCoordinate id="EE776CD31573A14E14AC89A1FA4845E3" box="[1251,1424,1756,1783]" degrees="28" direction="south" minutes="36" orientation="latitude" pageId="1" pageNumber="395" precision="1" seconds="57.6" value="-28.616001">28° 36’ 57.6” S</geoCoordinate>
|
<geoCoordinate id="EE776CD31573A14E10D8887DFE9C440F" box="[151,324,1792,1819]" degrees="32" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="395" precision="1" seconds="35.4" value="32.293167">32° 17’ 35.4” E</geoCoordinate>
|
<date id="FFFD2CD41573A14E1116887CFE1B440F" box="[345,451,1793,1819]" pageId="1" pageNumber="395" value="2000-01-27">
<collectingDate id="EFB9D53C1573A14E1116887CFE1B440F" box="[345,451,1793,1819]" pageId="1" pageNumber="395" value="2000-01-27">27.i.2000</collectingDate>
</date>
|
<collectorName id="26B66FC21573A14E1198887CFD4B440F" box="[471,659,1793,1819]" pageId="1" pageNumber="395">Davis &amp; Delport</collectorName>
||
<collectingMethod id="520272031573A14E12E3887CFC68440F" box="[684,944,1793,1819]" pageId="1" pageNumber="395">dung-baited pitfall trap</collectingMethod>
| CP13 (Cattle [&amp; pig] composite) | +/-6 yr, Stand 6, 1995 [year that the rehabilitation of the forest started] |
<collectedFrom id="385F49E91573A14E13168858FBD1442B" box="[857,1033,1829,1855]" pageId="1" pageNumber="395">Open shrubland</collectedFrom>
&amp; | thicket | Site 2, Trap 1 ||
<emphasis id="B937D6061573A14E15708859FF334477" bold="true" pageId="1" pageNumber="395">HOLOTYPE:</emphasis>
<taxonomicName id="4C4371971573A14E10BC8834FEA74477" authorityName="Harold" authorityYear="1871" box="[243,383,1865,1891]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="B937D6061573A14E10BC8834FEA74477" box="[243,383,1865,1891]" italics="true" pageId="1" pageNumber="395">Stiptopodius</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4371971573A14E11DB8837FD354477" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[404,749,1865,1891]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="B937D6061573A14E11DB8837FDD44477" box="[404,524,1866,1891]" italics="true" pageId="1" pageNumber="395">peringueyi</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
| 2020 [printed on red card]
<specimenCode id="DBE5A26F1573A14E147E8834FB184477" box="[1073,1216,1865,1891]" pageId="1" pageNumber="395">TYPH02476</specimenCode>
(
<collectionCode id="ED5292D11573A14E14808834FAF84477" box="[1231,1312,1865,1891]" collectionName="South Africa, Pretoria, South African National Collection of Insects" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="395" type="Museum">SANC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B2B00491573A14E157C8834FAA444E7" ID-GBIF-Occurrence="3396360301" collectingDate="1992" collectionCode="KZN, SANC" collectorName="M. Vogt" country="South Africa" determinerName="J. duG. Harrison" latitude="-28.633333" location="Richards Bay" longLatPrecision="1231" longitude="32.266666" municipality="Lake Nhlabane" pageId="1" pageNumber="395" specimenCode="COLS 35796" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<typeStatus id="54F8B4B61573A14E157C8834FA404477" box="[1331,1432,1865,1891]" pageId="1" pageNumber="395" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D45C19D1573A14E10D88810FF0A4493" box="[151,210,1901,1927]" pageId="1" pageNumber="395" type="male">male</specimenCount>
:
<collectingCountry id="F3544A841573A14E10928810FE724493" box="[221,426,1901,1927]" name="South Africa" pageId="1" pageNumber="395">SOUTH AFRICA</collectingCountry>
,
<collectionCode id="ED5292D11573A14E11FA8810FE374493" box="[437,495,1901,1927]" collectionName="KZN" pageId="1" pageNumber="395">KZN</collectionCode>
[
<collectingRegion id="4987C4F61573A14E11B38811FD6A4493" box="[508,690,1900,1927]" country="South Africa" name="KwaZulu-Natal" pageId="1" pageNumber="395">KwaZulu-Natal</collectingRegion>
] | near
<collectingMunicipality id="6B98906E1573A14E134B8810FC764493" box="[772,942,1901,1927]" pageId="1" pageNumber="395">Lake Nhlabane</collectingMunicipality>
|
<locationDeviation id="9A4E13361573A14E138D8811FB954493" bearing="45.0" box="[962,1101,1900,1927]" distance="25000.0" pageId="1" pageNumber="395">
<quantity id="4CBBA7F11573A14E138D8811FBDA4493" box="[962,1026,1900,1927]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="395" unit="km" value="25.0">25km</quantity>
NE of
</locationDeviation>
<location id="8E9C5CCF1573A14E141A8810FB354493" LSID="urn:lsid:plazi:treatment:03EABB021573A14D10D88930FCD14483:8E9C5CCF1573A14E141A8810FB354493" box="[1109,1261,1901,1927]" country="South Africa" latitude="-28.633333" longLatPrecision="1231" longitude="32.266666" municipality="Lake Nhlabane" name="Richards Bay" pageId="1" pageNumber="395" stateProvince="KwaZulu-Natal">Richards Bay</location>
|
<geoCoordinate id="EE776CD31573A14E154E8811FAB24493" box="[1281,1386,1900,1927]" degrees="28" direction="south" minutes="38" orientation="latitude" pageId="1" pageNumber="395" precision="925" value="-28.633333">28° 38’ S</geoCoordinate>
;
<geoCoordinate id="EE776CD31573A14E15388811FF0944BF" degrees="32" direction="east" minutes="16" orientation="longitude" pageId="1" pageNumber="395" precision="925" value="32.266666">32° 16’ E</geoCoordinate>
|
<collectingDate id="EFB9D53C1573A14E10AC88ECFEC644BF" box="[227,286,1937,1963]" pageId="1" pageNumber="395">1992</collectingDate>
,
<collectorName id="26B66FC21573A14E116788ECFE5944BF" box="[296,385,1937,1963]" pageId="1" pageNumber="395">M. Vogt</collectorName>
|| University of Pretoria Project | Richards Bay Minerals | Dune forest rehab. Survey || National Coll. of Insects | Pretoria, S. Africa || Dichotomini | det.
<determinerName id="7E20E6D01573A14E135588C8FC0E44DB" box="[794,982,1973,1999]" pageId="1" pageNumber="395">J. duG. Harrison</determinerName>
2013 || SANC-Pretoria | Database No: |
<specimenCode id="DBE5A26F1573A14E10D888A4FEF444E7" box="[151,300,2008,2035]" pageId="1" pageNumber="395">COLS 35796</specimenCode>
|
<emphasis id="B937D6061573A14E117088A5FE0544E7" bold="true" box="[319,477,2008,2035]" pageId="1" pageNumber="395">PARATYPE:</emphasis>
<taxonomicName id="4C4371971573A14E11AB88A4FDA844E7" authorityName="Harold" authorityYear="1871" box="[484,624,2009,2035]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="B937D6061573A14E11AB88A4FDA844E7" box="[484,624,2009,2035]" italics="true" pageId="1" pageNumber="395">Stiptopodius</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4371971573A14E12CB88A7FC0344E7" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[644,987,2009,2035]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="B937D6061573A14E12CB88A7FD2444E7" box="[644,764,2010,2035]" italics="true" pageId="1" pageNumber="395">peringueyi</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
| 2020 [printed on red card] (
<collectionCode id="ED5292D11573A14E156C88A4FAAC44E7" box="[1315,1396,2009,2035]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="395" type="Museum">SANC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D10888FE5FCC347B7" pageId="2" pageNumber="396" type="description">
<paragraph id="8BFC0A141570A14D10888FE5FC5043A7" blockId="2.[151,1437,152,1943]" box="[199,904,152,179]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888FE5FE8143A7" bold="true" box="[199,345,152,179]" pageId="2" pageNumber="396">Description.</emphasis>
<typeStatus id="54F8B4B61570A14D112F8FE4FE1343A7" box="[352,459,153,179]" pageId="2" pageNumber="396" type="holotype">Holotype</typeStatus>
, male. Length: 4.0 mm, width:
<quantity id="4CBBA7F11570A14D13648FE4FC5C43A7" box="[811,900,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="396" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<paragraph id="8BFC0A141570A14D10888FC1FD8F43C3" blockId="2.[151,1437,152,1943]" box="[199,599,188,215]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888FC1FEF843C3" bold="true" box="[199,288,188,215]" pageId="2" pageNumber="396">Colour.</emphasis>
Brown to very dark brown.
</paragraph>
<paragraph id="8BFC0A141570A14D10888F9CFA9C429F" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888F9CFED643EF" bold="true" box="[199,270,225,251]" pageId="2" pageNumber="396">Head.</emphasis>
Incision between medial clypeal teeth deeply concave; latero-clypeal edges not sinuate; genae with external protrusion sharply triangular; surface of head shiny with punctation varying from fine to large, ocellate and setiferous; punctation dense on genae and sparse on vertex; clypeo-frontal carina visible, clypeal-genae suture present, vertex bearing carina slightly sinuate medially and situated immediately behind the anterior edge of eyes, never forming two cephalic tubercles. Antennae with eight antennomeres, labium with few well-developed setae.
</paragraph>
<paragraph id="8BFC0A141570A14D10888EE8FDF542E3" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888EE8FE9F42BB" bold="true" box="[199,327,405,431]" pageId="2" pageNumber="396">Pronotum.</emphasis>
Convex, wider than long; base of pronotum marginate; anterior edge wider than posterior; in lateral view, weakly sinuate before postero-lateral angles; surface shiny metallic brown, punctation ocellate, very short setae laterally, central disc glabrous.
</paragraph>
<paragraph id="8BFC0A141570A14D10888D7CFE9E4177" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888D7CFEC2410F" bold="true" box="[199,282,513,539]" pageId="2" pageNumber="396">Elytra.</emphasis>
Lateral margin fairly straight, somewhat convex distally and concave proximally; sparsely and finely punctate in the last four striae with the rest apunctate; interstriae punctate, punctures on the side and posterior part sparsely setose.
</paragraph>
<paragraph id="8BFC0A141570A14D10888D11FC3C4193" blockId="2.[151,1437,152,1943]" box="[199,996,620,647]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888D11FE9A4193" bold="true" box="[199,322,620,647]" pageId="2" pageNumber="396">Pygidium.</emphasis>
Convex, punctation ocellate with setae emanating centrally.
</paragraph>
<paragraph id="8BFC0A141570A14D10888DEDFBF9400C" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888DEDFE1041BF" bold="true" box="[199,456,656,683]" pageId="2" pageNumber="396">Abdominal ventrites.</emphasis>
Prosternum with well-developed setae, anterior angles slanted forward and inward. Mesoventrite dull, setose and densely punctate. Metaventrite shiny with long setae anteriorly, glabrous and sparsely punctate mid-posteriorly. Abdominal ventrites shiny, setigerous with ocellate punctures, strong setae arranged in lateral rows, glabrous medially, last ventrite narrowed medially (male character).
</paragraph>
<paragraph id="8BFC0A141570A14D10888C5CFC92476B" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888C5CFEDE402F" bold="true" box="[199,262,801,827]" pageId="2" pageNumber="396">Legs.</emphasis>
Profemora sparsely punctate, bearing well-developed setae, external edge strongly marginate, internal margin slightly marginate. Meso- and metafemora with external and internal edges sparsely setigerous and punctate; surface between external and internal margin glabrous and apunctate, strongly marginate in both edges (external and internal). Protibiae crenulate with four teeth on external margin, a line of setae around teeth in dorsal view; in ventral view, with two carinae, internally punctate and pilose. Protibial spur long, curved inwardly. Meso- and metatibiae triangular and strongly widened apically. Protarsi 5-articulated, protarsomere V at least 2x longer than others (each one individually), connected to a clearly visible pair of claws at distal tip. Meso- and metatarsomeres I–IV triangular, wider than long, with line of strong setae marginally (external and internal); tarsomeres widened apically, strongly concave medially with external and internal edges bearing a lobe distally; metatarsomere V rectangular and longer than others, followed by pair of very tiny apical claws.
</paragraph>
<paragraph id="8BFC0A141570A14D10888BF5FCC347B7" blockId="2.[151,1437,152,1943]" box="[199,795,1160,1187]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888BF5FE9947B7" bold="true" box="[199,321,1160,1187]" pageId="2" pageNumber="396">Aedeagus.</emphasis>
Parameres symmetrical (see
<figureCitation id="137816911570A14D12C68BF4FD0E47B7" box="[649,726,1161,1187]" captionStart="FIGURES 2" captionStartId="4.[151,264,1944,1969]" captionTargetBox="[183,1394,196,1907]" captionTargetId="figure@4.[179,1407,181,1919]" captionTargetPageId="4" captionText="FIGURES 2A–F. Aedeagi of Stiptopodius Harold, 1871 species in dorsal (A, C, E) and lateral (B, D, F) views. A, B, Stiptopodius muellerae Daniel &amp; Deschodt, new species; C, D, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; E, F, Stiptopodius singularis (Péringuey, 1901). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759531" httpUri="https://zenodo.org/record/3759531/files/figure.png" pageId="2" pageNumber="396">Figs. 2</figureCitation>
C–D).
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D10888BD1FD0047FF" pageId="2" pageNumber="396" type="distribution">
<paragraph id="8BFC0A141570A14D10888BD1FD0047FF" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888BD1FEB947D3" bold="true" box="[199,353,1196,1223]" pageId="2" pageNumber="396">Distribution.</emphasis>
The species is only known from the
<typeStatus id="54F8B4B61570A14D12B68BD0FCF147D3" box="[761,809,1197,1223]" pageId="2" pageNumber="396">type</typeStatus>
locality, Richards Bay,
<collectingRegion id="4987C4F61570A14D147C8BD0FB3E47D3" box="[1075,1254,1197,1223]" country="South Africa" name="KwaZulu-Natal" pageId="2" pageNumber="396">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="F3544A841570A14D14BF8BD0FA5947D3" box="[1264,1409,1197,1223]" name="South Africa" pageId="2" pageNumber="396">South Africa</collectingCountry>
. It was collected in a six-year-old rehabilitating forest.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D10888B88FE3E4643" pageId="2" pageNumber="396" type="etymology">
<paragraph id="8BFC0A141570A14D10888B88FE3E4643" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888B88FE96461B" bold="true" box="[199,334,1269,1295]" pageId="2" pageNumber="396">Etymology.</emphasis>
The specific name is in honour of Dr. Louis Albert Péringuey (
<date id="FFFD2CD41570A14D14558B89FB45461B" box="[1050,1181,1268,1295]" pageId="2" pageNumber="396" value="1855" valueMax="1924">1855–1924</date>
), a French-South African entomologist and former director of the Iziko Museums of South African, who contributed immensely to the knowledge of African insects.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D10888A1DFE0B453B" pageId="2" pageNumber="396" type="diagnosis">
<paragraph id="8BFC0A141570A14D10888A1DFE0B453B" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888A1DFE9A466F" bold="true" box="[199,322,1376,1403]" pageId="2" pageNumber="396">Diagnosis.</emphasis>
<taxonomicName id="4C4371971570A14D11088A1CFD89466F" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[327,593,1377,1403]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D6061570A14D11088A1CFD89466F" box="[327,593,1377,1403]" italics="true" pageId="2" pageNumber="396">Stiptopodius peringueyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1570A14D12188A1CFD04466F" box="[599,732,1377,1403]" pageId="2" pageNumber="396" rank="species">new species</taxonomicNameLabel>
can be distinguished from the other two species in the
<taxonomicName id="4C4371971570A14D15778A1CFA44466F" authorityName="Harold" authorityYear="1871" box="[1336,1436,1377,1403]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B937D6061570A14D15778A1CFA44466F" box="[1336,1436,1377,1403]" italics="true" pageId="2" pageNumber="396">S. doriae</emphasis>
</taxonomicName>
species-group (
<taxonomicName id="4C4371971570A14D110D8AF8FE7F468B" authorityName="Harold" authorityYear="1871" box="[322,423,1413,1439]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B937D6061570A14D110D8AF8FE7F468B" box="[322,423,1413,1439]" italics="true" pageId="2" pageNumber="396">S. doriae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4371971570A14D11938AF8FDB4468B" authorityName="Branco" authorityYear="1991" box="[476,620,1413,1439]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="longipedis">
<emphasis id="B937D6061570A14D11938AF8FDB4468B" box="[476,620,1413,1439]" italics="true" pageId="2" pageNumber="396">S. longipedis</emphasis>
</taxonomicName>
) by having a distinctive brown metallic sheen on the pronotum.
<taxonomicName id="4C4371971570A14D15778AF8FA44468B" authorityName="Harold" authorityYear="1871" box="[1336,1436,1413,1439]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B937D6061570A14D15778AF8FA44468B" box="[1336,1436,1413,1439]" italics="true" pageId="2" pageNumber="396">S. doriae</emphasis>
</taxonomicName>
has a very elongate body, unlike the short and wide body of
<taxonomicName id="4C4371971570A14D130C8AD4FC0046D7" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[835,984,1449,1475]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D6061570A14D130C8AD4FC0046D7" box="[835,984,1449,1475]" italics="true" pageId="2" pageNumber="396">S. peringueyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1570A14D13AF8AD4FBBF46D7" box="[992,1127,1449,1475]" pageId="2" pageNumber="396" rank="species">new species</taxonomicNameLabel>
. The meso- and metatarsomeres I to IV of the new species are triangular and wider than long, while in
<taxonomicName id="4C4371971570A14D13908AB0FBB746F3" authorityName="Branco" authorityYear="1991" box="[991,1135,1485,1511]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="longipedis">
<emphasis id="B937D6061570A14D13908AB0FBB746F3" box="[991,1135,1485,1511]" italics="true" pageId="2" pageNumber="396">S. longipedis</emphasis>
</taxonomicName>
meso- and metatarsomeres I and II are much longer than wide. Furthermore, the latter species lacks meso- and metatarsomeres IV and V and meso- and metatarsal claws.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D10888944FF6445AB" pageId="2" pageNumber="396" type="discussion">
<paragraph id="8BFC0A141570A14D10888944FF6445AB" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D10888944FEEC4547" bold="true" box="[199,308,1593,1619]" pageId="2" pageNumber="396">Remarks</emphasis>
. In
<collectingCountry id="F3544A841570A14D112B8944FE2F4547" box="[356,503,1593,1619]" name="South Africa" pageId="2" pageNumber="396">South Africa</collectingCountry>
,
<taxonomicName id="4C4371971570A14D124A8944FD434547" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[517,667,1593,1619]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="peringueyi" status="sp. nov.">
<emphasis id="B937D6061570A14D124A8944FD434547" box="[517,667,1593,1619]" italics="true" pageId="2" pageNumber="396">S. peringueyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1570A14D12EC8944FCF44547" box="[675,812,1593,1619]" pageId="2" pageNumber="396" rank="species">new species</taxonomicNameLabel>
can be misidentified as
<taxonomicName id="4C4371971570A14D140A8944FB0D4547" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[1093,1237,1593,1619]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="singularis">
S.
<emphasis id="B937D6061570A14D142A8944FB0D4547" box="[1125,1237,1593,1619]" italics="true" pageId="2" pageNumber="396">singularis</emphasis>
</taxonomicName>
because both are roughly the same size. However, the two species belong to different species groups (see Introduction). Furthermore, the new species bears a carina on the head divided into two tubercles, which is lacking in
<taxonomicName id="4C4371971570A14D143889FCFADA458F" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[1143,1282,1665,1691]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B937D6061570A14D143889FCFADA458F" box="[1143,1282,1665,1691]" italics="true" pageId="2" pageNumber="396">S. singularis</emphasis>
</taxonomicName>
(see
<figureCitation id="137816911570A14D157389FCFF7745AB" captionStart="FIGURES 1" captionStartId="3.[151,264,1877,1902]" captionTargetBox="[163,1426,206,1839]" captionTargetId="figure@3.[151,1436,198,1849]" captionTargetPageId="3" captionText="FIGURES 1A–D. Habitus of Stiptopodius Harold, 1871 species, holotypes. A, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; B, Stiptopodius savuti Daniel &amp; Deschodt, new species; C, Stiptopodius muellerae Daniel &amp; Deschodt, new species; D, Stiptopodius singularis (Péringuey, 1901). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759529" httpUri="https://zenodo.org/record/3759529/files/figure.png" pageId="2" pageNumber="396">Figs.1A, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359599F1570A14D108889B5FCD14483" pageId="2" pageNumber="396" type="conservation">
<paragraph id="8BFC0A141570A14D108889B5FCD14483" blockId="2.[151,1437,152,1943]" pageId="2" pageNumber="396">
<emphasis id="B937D6061570A14D108889B5FD6145F7" bold="true" box="[199,697,1736,1763]" pageId="2" pageNumber="396">Conservation status and potential threats.</emphasis>
The species was collected on coastal dunes in a six-year-old stand comprising a monoculture of
<taxonomicName id="4C4371971570A14D11918990FB514413" authority="(P. P. Sw. ex Coates Palgr.) Kyal. &amp; Boatwr" authorityName="Kyal. &amp; Boatwr" baseAuthorityName="Coates Palgr." box="[478,1161,1773,1800]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="396" phylum="Tracheophyta" rank="species" species="kosiensis">
<emphasis id="B937D6061570A14D11918990FD694413" box="[478,689,1773,1799]" italics="true" pageId="2" pageNumber="396">Vachellia kosiensis</emphasis>
(P.P.Sw. ex Coates Palgr.) Kyal. &amp; Boatwr
</taxonomicName>
that replaced the cleared dune forest after dredge-mining. The trees were spaced about one meter apart and were about two to three meters tall with a 100% closed canopy (CMD, personal observation). It is unclear if this species would prefer an open forest habitat and or if its occurrence was affected by mining strategies. The extent of occurrence of this species is not known and it should thus be assessed as Data Deficient.
</paragraph>
</subSubSection>
</treatment>
</document>
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