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<document id="65F14E0CB244DF1E8F2DC3E7BF8BE012" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy &amp; Toda, Masanori J." docDate="2021" docId="03D75C2D45752E1EFF5262D3B1D2B259" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia kinabaluana Toda &amp; Takano 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="32" updateTime="1699472124804" updateUser="ExternalLinkService">
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6)
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<emphasis id="B90A312945752E20FF7262D3B145B1CD" bold="true" box="[183,521,726,753]" italics="true" pageId="31" pageNumber="32">Colocasiomyia kinabaluana</emphasis>
Toda &amp; Takano
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(
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,
<figureCitation id="1345F1BE45752E20FF3A62FFB266B029" box="[255,298,762,789]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada &amp; Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="31" pageNumber="32">15F</figureCitation>
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<figureCitation id="1345F1BE45752E20FEF362FEB21FB029" box="[310,339,763,789]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">21</figureCitation>
)
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sp.2
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aff.
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:
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Sultana
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Takano
<emphasis id="B90A312945752E20FBB66345B7D3B06B" box="[1139,1183,831,855]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
., 2011: 22
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<treatmentCitation id="0ADFCB2A45752E20FAD1633AB263B04A" author="Takano, T. K. &amp; Repin, R. &amp; Mohamed, D. M. B. &amp; Toda, M. J." page="559" pageId="31" pageNumber="32" year="2012">
<bibRefCitation id="EFEF90CA45752E20FAD1633AB263B04A" author="Takano, T. K. &amp; Repin, R. &amp; Mohamed, D. M. B. &amp; Toda, M. J." pageId="31" pageNumber="32" pagination="555 - 564" refId="ref33627" refString="Takano, T. K., Repin, R., Mohamed, D. M. B. &amp; Toda, M. J. (2012) Pollination mutualism between Alocasia macrorrhizos (Araceae) and two taxonomically undescribed Colocasiomyia species (Diptera: Drosophilidae), in Sabah, Borneo. Plant Biology, 14, 555 - 564. https: // doi. org / 10.1111 / j. 1438 - 8677.2011.00541. x" type="journal article" year="2012">
Takano
<emphasis id="B90A312945752E20FAA16345B6DCB06B" box="[1380,1424,831,855]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
., 2012: 559
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, Supplemental File 1;
<bibRefCitation id="EFEF90CA45752E20FDD6635BB19AB04A" author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[531,726,862,886]" pageId="31" pageNumber="32" pagination="763 - 782" refId="ref32284" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. &amp; Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
Fartyal
<emphasis id="B90A312945752E20FDA4635AB1C2B04A" box="[609,654,862,886]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
., 2013
</bibRefCitation>
,
<figureCitation id="1345F1BE45752E20FD25635BB052B04A" box="[736,798,862,886]" captionStart="FIGURE 6" captionStartId="11.[151,250,1951,1975]" captionTargetBox="[286,1299,181,1927]" captionTargetId="figure-18@11.[286,1300,181,1927]" captionTargetPageId="11" captionText="FIGURE 6. Character 12 of costal setae on the wing (anterodistal portion) of Colocasiomyia xenalocasiae (Okada) (A), C. colocasiae (Duda) (B) and C. cristata de Meijere (C). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766414" httpUri="https://zenodo.org/record/5766414/files/figure.png" pageId="31" pageNumber="32">Fig. 6</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C364BEB045752E20FF5263A1B259B772" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BC1ED3B45752E20FF5263A1B259B772" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
<emphasis id="B90A312945752E20FF5263A1B25EB082" bold="true" box="[151,274,932,958]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
A pair of processes on male abdominal sternite VI short (&lt;
<quantity id="4C8640DE45752E20FC0C63A1B77AB083" box="[969,1078,932,959]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="0.05">0.05 mm</quantity>
), slightly divergent and curved inward, apically round (
<figureCitation id="1345F1BE45752E20FE6C63CDB2B7B0DE" box="[425,507,968,994]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson &amp; Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada &amp; Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson &amp; Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda &amp; Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="31" pageNumber="32">Fig. 9J</figureCitation>
). Epandrium posteroventrally extended as ventral lobe roundish on apical margin (
<figureCitation id="1345F1BE45752E20FF6563E9B249B73A" box="[160,261,1004,1030]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21C</figureCitation>
). Phallal sheath apicoventrally pointed like beak in lateral view, apically looking like arrowhead in ventral view (
<figureCitation id="1345F1BE45752E20FF1B6415B22CB716" box="[222,352,1040,1066]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21D,E</figureCitation>
). Epiproct and hypoproct nearly flat (
<figureCitation id="1345F1BE45752E20FCC96415B03DB716" box="[780,881,1040,1066]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21F</figureCitation>
). Apical portion of oviscapt somewhat triangular (
<figureCitation id="1345F1BE45752E20FF656431B244B772" box="[160,264,1076,1102]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB045752E20FF02645DB078B47E" pageId="31" pageNumber="32" type="description">
<paragraph id="8BC1ED3B45752E20FF02645DB14AB786" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
<emphasis id="B90A312945752E20FF02645DB21DB74E" bold="true" box="[199,337,1112,1138]" pageId="31" pageNumber="32">Description</emphasis>
(
<specimenCount id="9D7826B245752E20FEA4645DB23AB74E" box="[353,374,1112,1138]" pageId="31" pageNumber="32" type="male">♂</specimenCount>
and
<specimenCount id="9D7826B245752E20FE68645CB28DB74F" box="[429,449,1113,1139]" pageId="31" pageNumber="32" type="female">♀</specimenCount>
). Head. Supracervical setae 6–9 per side, distally more or less curved, longer than inner occipital setae. Cibarial, medial sensilla approximately 3 per side; posterior sensillum 1 per side. Supralateral setae outside prementum 3–4 per side.
</paragraph>
<paragraph id="8BC1ED3B45752E20FF0264C1B0D1B7E3" blockId="31.[151,1437,932,2039]" box="[199,925,1220,1247]" pageId="31" pageNumber="32">Thorax. Anterior dorsocentral setae just beside transverse suture.</paragraph>
<paragraph id="8BC1ED3B45752E20FF0264EDB25FB61A" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
Abdomen. Male sternite III and IV longer than wide (
<figureCitation id="1345F1BE45752E20FCE964EDB0DAB63E" box="[812,918,1256,1282]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21A</figureCitation>
). Female sternites V and VI longer than wide (
<figureCitation id="1345F1BE45752E20FF656509B24AB61A" box="[160,262,1292,1318]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21B</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B45752E20FF026535B1EBB6E6" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
Male terminalia. Epandrium pubescent except for anterior margin, anteroventral elongation and ventral portion, with 1–2 setae on dorsal portion and 21–23 setae thicker than cercal setae on ventral lobe of each side (
<figureCitation id="1345F1BE45752E20FAED6551B6DCB652" box="[1320,1424,1364,1390]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21C</figureCitation>
). Cercus pubescent on dorsal 2/3 but not on anterior margin and ventral portion, with 38–39 setae, extended below, apically forming small lobe marginally fringed with small setae (
<figureCitation id="1345F1BE45752E20FCA66599B084B68A" box="[867,968,1436,1462]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21C</figureCitation>
). Phallapodeme less sclerotized thin plate, nearly perpendicular to phallal axis (
<figureCitation id="1345F1BE45752E20FDF765C5B1D6B6E6" box="[562,666,1472,1498]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21D</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B45752E20FF0265E1B725B51E" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
Female terminalia. Hypoproct not pubescent. Oviscapt longer than phallus (apodeme + sheath), with 21–22 ovisensilla only on distal portion occupying less than 1/2 length of oviscapt (
<figureCitation id="1345F1BE45752E20FC31660DB711B51E" box="[1012,1117,1544,1570]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21G</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B45752E20FF026629B600B5B3" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
Indices (range of
<specimenCount id="9D7826B245752E20FE4C6629B2F6B57A" box="[393,442,1580,1606]" pageId="31" pageNumber="32" type="male">10♂</specimenCount>
and
<specimenCount id="9D7826B245752E20FE2A6629B16EB57A" box="[495,546,1580,1606]" pageId="31" pageNumber="32" type="female">10♀</specimenCount>
): FW/HW = 0.57–0.67, ch/o = 0.37–0.53, prorb = 0.94–1.15, rcorb = 0.25–0.54, vb = 0.23–0.38, dcl = 0.58–0.86, sctl = 0.68–0.86, sterno = 0.83–1.02, orbito = 0.44–0.84, dcp = 1.02–1.33, sctlp = 1.07–1.36, C = 1.68–2.38, 4c = 1.02–1.37, 4v = 1.45–1.86, 5x = 0.72–1.16, ac = 2.18–3.01, M = 0.35–0.48.
</paragraph>
<paragraph id="8BC1ED3B45752E20FF02669DB078B47E" blockId="31.[151,1437,932,2039]" pageId="31" pageNumber="32">
Puparium (3rd instar larva). Segments with stout spicules on ventral surface; anterior spiracle sessile, with a bundle of approximately 4 short branches; caudal segments elongate, with many small spicules, ending in a Vshaped pair of posterior spiracles (
<figureCitation id="1345F1BE45752E20FDE366E5B1EDB5C6" box="[550,673,1760,1786]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21H,I</figureCitation>
). Mouth hook less expanded medioventrally in lateral view; distal blade as long as basal portion, apically pointed, weakly curved downward, with two rows of small, acute teeth on submedial to subapical portion of ventral margin (
<figureCitation id="1345F1BE45752E20FD0D672DB06AB47E" box="[712,806,1832,1858]" captionStart="FIGURE 21" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1435,273,1812]" captionTargetId="figure-60@32.[151,1435,273,1813]" captionTargetPageId="32" captionText="FIGURE 21. Colocasiomyia kinabaluana Toda &amp; Takano, sp. nov. (♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I." figureDoi="http://doi.org/10.5281/zenodo.5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="31" pageNumber="32">Fig. 21J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB045752E1FFF026749B3BDB3C5" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BC1ED3B45752E20FF026749B622B45A" blockId="31.[151,1437,932,2039]" box="[199,1390,1868,1895]" pageId="31" pageNumber="32">
<materialsCitation id="3B16E76645752E20FF026749B622B45A" ID-GBIF-Occurrence="3418896961" box="[199,1390,1868,1895]" collectingDate="1999-01-03" collectionCode="BORN" country="Malaysia" location="Kota Kinabalu" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B90A312945752E20FF026749B277B45A" bold="true" box="[199,315,1868,1894]" pageId="31" pageNumber="32">
<typeStatus id="54C5539945752E20FF026749B27AB45A" box="[199,310,1868,1894]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D7826B245752E20FE876748B21BB45B" box="[322,343,1869,1895]" pageId="31" pageNumber="32" type="male">♂</specimenCount>
(
<collectionCode id="ED6F75FE45752E20FEA36749B2F5B45A" box="[358,441,1868,1894]" country="Malaysia" httpUri="http://grbio.org/cool/y0pn-rrd0" name="Institute for Tropical Biology and Conservation, Borneensis" pageId="31" pageNumber="32">BORN</collectionCode>
), “
<location id="8EA1BBE045752E20FE136749B1CCB45A" LSID="urn:lsid:plazi:treatment:03D75C2D45752E1EFF5262D3B1D2B259:8EA1BBE045752E20FE136749B1CCB45A" box="[470,640,1868,1894]" country="Malaysia" name="Kota Kinabalu" pageId="31" pageNumber="32" stateProvince="Sabah">Kota Kinabalu</location>
,
<collectingRegion id="49BA23D945752E20FD4E6749B19EB45A" box="[651,722,1868,1894]" country="Malaysia" name="Sabah" pageId="31" pageNumber="32">Sabah</collectingRegion>
,
<collectingCountry id="F369ADAB45752E20FD186749B00AB45A" box="[733,838,1868,1894]" name="Malaysia" pageId="31" pageNumber="32">Malaysia</collectingCountry>
,
<date id="FFC0CBFB45752E20FC976749B0FCB45A" box="[850,944,1868,1894]" pageId="31" pageNumber="32" value="1999-01-03">
<collectingDate id="EF84321345752E20FC976749B0FCB45A" box="[850,944,1868,1894]" pageId="31" pageNumber="32" value="1999-01-03">3.i.1999</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B845752E20FC186749B62CB45A" authority=", M. J. Toda" authorityName="M. J. Toda" box="[989,1376,1868,1895]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A312945752E20FC186749B797B45A" box="[989,1243,1868,1894]" italics="true" pageId="31" pageNumber="32">Alocasia macrorrhizos</emphasis>
, M.J. Toda
</taxonomicName>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BC1ED3B45752E1FFF026775B3BDB3C5" blockId="31.[151,1437,932,2039]" lastBlockId="32.[151,1436,151,249]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<materialsCitation id="3B16E76645752E20FF026775B03BB4B6" ID-GBIF-Occurrence="3418896448" box="[199,887,1904,1931]" collectingDate="1999-01-03" collectionCode="BORN" country="Malaysia" location="Kota Kinabalu" pageId="31" pageNumber="32" specimenCount="33" specimenCount-female="10" specimenCount-male="23" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B90A312945752E20FF026775B20AB4B6" bold="true" box="[199,326,1904,1930]" pageId="31" pageNumber="32">
<typeStatus id="54C5539945752E20FF026775B20DB4B6" box="[199,321,1904,1930]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F369ADAB45752E20FE8A6775B2F6B4B6" box="[335,442,1904,1930]" name="Malaysia" pageId="31" pageNumber="32">Malaysia</collectingCountry>
:
<specimenCount id="9D7826B245752E20FE026775B2B6B4B6" box="[455,506,1904,1930]" pageId="31" pageNumber="32" type="male">23♂</specimenCount>
,
<specimenCount id="9D7826B245752E20FDCD6775B177B4B6" box="[520,571,1904,1930]" pageId="31" pageNumber="32" type="female">10♀</specimenCount>
, same data as the holotype
</materialsCitation>
;
<materialsCitation id="3B16E76645752E20FC416775B21CB492" ID-GBIF-Occurrence="3418896505" collectingDate="2004-08-12" collectionCode="T, BORN, SEHU" collectorName="Takano" country="Malaysia" location="Kota Kinabalu" pageId="31" pageNumber="32" specimenCount="6" specimenCount-male="6" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D7826B245752E20FC416775B0E5B4B6" box="[900,937,1904,1930]" pageId="31" pageNumber="32" type="male">6♂</specimenCount>
, ditto except
<date id="FFC0CBFB45752E20FB836775B79CB4B6" box="[1094,1232,1904,1930]" pageId="31" pageNumber="32" value="2004-08-12">
<collectingDate id="EF84321345752E20FB836775B79CB4B6" box="[1094,1232,1904,1930]" pageId="31" pageNumber="32" value="2004-08-12">12.viii.2004</collectingDate>
</date>
, K.
<collectionCode id="ED6F75FE45752E20FB3D6774B647B4B7" box="[1272,1291,1905,1931]" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T</collectionCode>
.
<collectorName id="268B88ED45752E20FADD6775B626B4B6" box="[1304,1386,1904,1930]" pageId="31" pageNumber="32">Takano</collectorName>
leg. (
<collectionCode id="ED6F75FE45752E20FF5A6791B3BDB492" box="[159,241,1940,1966]" country="Malaysia" httpUri="http://grbio.org/cool/y0pn-rrd0" name="Institute for Tropical Biology and Conservation, Borneensis" pageId="31" pageNumber="32">BORN</collectionCode>
,
<collectionCode id="ED6F75FE45752E20FF3F6791B204B492" box="[250,328,1940,1966]" pageId="31" pageNumber="32">SEHU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B16E76645752E20FE9D6791B1D1B4EE" ID-GBIF-Occurrence="3418896621" collectingDate="2012-11-30" country="Philippines" latitude="13.147778" location="Luzon" longLatPrecision="21" longitude="123.73444" municipality="Legaspi" pageId="31" pageNumber="32" specimenCount="13" specimenCount-female="6" specimenCount-male="7" typeStatus="paratype">
<collectingCountry id="F369ADAB45752E20FE9D6791B297B492" box="[344,475,1940,1966]" name="Philippines" pageId="31" pageNumber="32">Philippines</collectingCountry>
:
<specimenCount id="9D7826B245752E20FE266790B144B493" box="[483,520,1941,1967]" pageId="31" pageNumber="32" type="male">7♂</specimenCount>
,
<specimenCount id="9D7826B245752E20FDD46791B179B492" box="[529,565,1940,1966]" pageId="31" pageNumber="32" type="female">6♀</specimenCount>
,
<collectingMunicipality id="6BA5774145752E20FD856791B1D7B492" box="[576,667,1940,1966]" pageId="31" pageNumber="32">Legaspi</collectingMunicipality>
, S.
<location id="8EA1BBE045752E20FD056790B046B493" LSID="urn:lsid:plazi:treatment:03D75C2D45752E1EFF5262D3B1D2B259:8EA1BBE045752E20FD056790B046B493" box="[704,778,1941,1967]" country="Philippines" latitude="13.147778" longLatPrecision="21" longitude="123.73444" municipality="Legaspi" name="Luzon" pageId="31" pageNumber="32">Luzon</location>
,
<geoCoordinate id="EE4A8BFC45752E20FCD66791B0D0B493" box="[787,924,1940,1967]" degrees="13" direction="north" minutes="08" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="52" value="13.147778">13°08’52”N</geoCoordinate>
<geoCoordinate id="EE4A8BFC45752E20FC646791B77BB492" box="[929,1079,1940,1966]" degrees="123" direction="east" minutes="44" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="04" value="123.73444">123°44’04”E</geoCoordinate>
,
<date id="FFC0CBFB45752E20FB856791B7F7B492" box="[1088,1211,1940,1966]" pageId="31" pageNumber="32" value="2012-11-30">
<collectingDate id="EF84321345752E20FB856791B7F7B492" box="[1088,1211,1940,1966]" pageId="31" pageNumber="32" value="2012-11-30">30.xi.2012</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B845752E20FB266791B13CB4EE" authority=", P. J. Matthews, M. Medecilo &amp; J. R. Castillo" authorityName="P. J. Matthews, M. Medecilo &amp; J. R. Castillo" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A312945752E20FB266791B7B8B492" box="[1251,1268,1940,1966]" italics="true" pageId="31" pageNumber="32">A</emphasis>
.
<emphasis id="B90A312945752E20FAC56791B6D9B492" box="[1280,1429,1940,1966]" italics="true" pageId="31" pageNumber="32">macrorrhizos</emphasis>
, P.J. Matthews, M. Medecilo &amp; J.R. Castillo
</taxonomicName>
leg
</materialsCitation>
.;
<materialsCitation id="3B16E76645752E20FD6E67BDB021B4CA" ID-GBIF-Occurrence="3418896767" collectingDate="2012-12-02" country="Philippines" latitude="12.015555" location="Gandara" longLatPrecision="21" longitude="124.79528" pageId="31" pageNumber="32" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Samar" typeStatus="paratype">
<specimenCount id="9D7826B245752E20FD6E67BDB192B4EE" box="[683,734,1976,2002]" pageId="31" pageNumber="32" type="male">10♂</specimenCount>
,
<specimenCount id="9D7826B245752E20FD2D67BDB057B4EE" box="[744,795,1976,2002]" pageId="31" pageNumber="32" type="female">10♀</specimenCount>
,
<location id="8EA1BBE045752E20FCE067BDB0CBB4EE" LSID="urn:lsid:plazi:treatment:03D75C2D45752E1EFF5262D3B1D2B259:8EA1BBE045752E20FCE067BDB0CBB4EE" box="[805,903,1976,2002]" country="Philippines" latitude="12.015555" longLatPrecision="21" longitude="124.79528" name="Gandara" pageId="31" pageNumber="32" stateProvince="Samar">Gandara</location>
,
<collectingRegion id="49BA23D945752E20FC4A67BDB094B4EE" box="[911,984,1976,2002]" country="Philippines" name="Samar" pageId="31" pageNumber="32">Samar</collectingRegion>
,
<geoCoordinate id="EE4A8BFC45752E20FC2467BDB729B4EF" box="[993,1125,1976,2003]" degrees="12" direction="north" minutes="00" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="56" value="12.015555">12°00'56&quot;N</geoCoordinate>
<geoCoordinate id="EE4A8BFC45752E20FBAF67BDB7B7B4EE" box="[1130,1275,1976,2002]" degrees="124" direction="east" minutes="47" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="43" value="124.79528">124°47'43&quot;E</geoCoordinate>
,
<date id="FFC0CBFB45752E20FAC167BDB634B4EE" box="[1284,1400,1976,2002]" pageId="31" pageNumber="32" value="2012-12-02">
<collectingDate id="EF84321345752E20FAC167BDB634B4EE" box="[1284,1400,1976,2002]" pageId="31" pageNumber="32" value="2012-12-02">2.xii.2012</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B845752E20FF5267D9B073B4CA" authority=", P. J. Matthews, M. Medecilo &amp; J. R. Castillo" authorityName="P. J. Matthews, M. Medecilo &amp; J. R. Castillo" box="[151,831,2012,2039]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A312945752E20FF5267D9B3E4B4CA" box="[151,168,2012,2038]" italics="true" pageId="31" pageNumber="32">A</emphasis>
.
<emphasis id="B90A312945752E20FF7367D9B207B4CA" box="[182,331,2012,2038]" italics="true" pageId="31" pageNumber="32">macrorrhizos</emphasis>
, P.J. Matthews, M. Medecilo &amp; J.R. Castillo
</taxonomicName>
leg
</materialsCitation>
.;
<materialsCitation id="3B16E76645752E1FFCB867D9B775B38D" ID-GBIF-Occurrence="3418896577" collectingDate="2012-12-03" country="Philippines" lastPageId="32" lastPageNumber="33" latitude="11.1175" location="Tanuan" longLatPrecision="21" longitude="125.015274" pageId="31" pageNumber="32" specimenCount="2" specimenCount-male="2" stateProvince="Southern Leyte" typeStatus="paratype">
<specimenCount id="9D7826B245752E20FCB867D9B0EFB4CA" box="[893,931,2012,2038]" pageId="31" pageNumber="32" type="male">2♂</specimenCount>
,
<location id="8EA1BBE045752E20FC6B67D8B74FB4CB" LSID="urn:lsid:plazi:treatment:03D75C2D45752E1EFF5262D3B1D2B259:8EA1BBE045752E20FC6B67D8B74FB4CB" box="[942,1027,2013,2039]" country="Philippines" latitude="11.1175" longLatPrecision="21" longitude="125.015274" name="Tanuan" pageId="31" pageNumber="32" stateProvince="Southern Leyte">Tanuan</location>
, N.
<collectingRegion id="49BA23D945752E20FBEA67D8B73EB4CB" box="[1071,1138,2013,2039]" country="Philippines" name="Southern Leyte" pageId="31" pageNumber="32">Leyte</collectingRegion>
,
<geoCoordinate id="EE4A8BFC45752E20FBB867D9B64CB4CA" box="[1149,1280,2012,2038]" degrees="11" direction="north" minutes="07" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="03" value="11.1175">11°07'03&quot;N</geoCoordinate>
<geoCoordinate id="EE4A8BFC45752E20FAC367D9B6DBB4CB" box="[1286,1431,2012,2039]" degrees="125" direction="east" minutes="00" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="55" value="125.015274">125°00'55&quot;E</geoCoordinate>
,
<date id="FFC0CBFB454A2E1FFF526092B247B38D" box="[151,267,151,177]" pageId="32" pageNumber="33" value="2012-12-03">
<collectingDate id="EF843213454A2E1FFF526092B247B38D" box="[151,267,151,177]" pageId="32" pageNumber="33" value="2012-12-03">3.xii.2012</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B8454A2E1FFE856092B74AB38D" authority=", P. J. Matthews, M. Medecilo &amp; J. R. Castillo" authorityName="P. J. Matthews, M. Medecilo &amp; J. R. Castillo" box="[320,1030,151,177]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A3129454A2E1FFE856092B21DB38D" box="[320,337,151,177]" italics="true" pageId="32" pageNumber="33">A</emphasis>
.
<emphasis id="B90A3129454A2E1FFEA76092B2BBB38D" box="[354,503,151,177]" italics="true" pageId="32" pageNumber="33">macrorrhizos</emphasis>
, P.J. Matthews, M. Medecilo &amp; J.R. Castillo
</taxonomicName>
leg
</materialsCitation>
.;
<materialsCitation id="3B16E766454A2E1FFB886092B3A7B3C5" ID-GBIF-Occurrence="3418896890" collectingDate="2013-02-05" collectionCode="MPMP, SEHU" country="Philippines" latitude="10.190001" location="Sabang" longLatPrecision="21" longitude="118.9" pageId="32" pageNumber="33" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Palawan" typeStatus="paratype">
<specimenCount id="9D7826B2454A2E1FFB886092B7CCB38D" box="[1101,1152,151,177]" pageId="32" pageNumber="33" type="male">10♂</specimenCount>
,
<specimenCount id="9D7826B2454A2E1FFB556092B78EB38D" box="[1168,1218,151,177]" pageId="32" pageNumber="33" type="female">10♀</specimenCount>
,
<location id="8EA1BBE0454A2E1FFB176092B66BB38D" LSID="urn:lsid:plazi:treatment:03D75C2D45752E1EFF5262D3B1D2B259:8EA1BBE0454A2E1FFB176092B66BB38D" box="[1234,1319,151,177]" country="Philippines" latitude="10.190001" longLatPrecision="21" longitude="118.9" name="Sabang" pageId="32" pageNumber="33" stateProvince="Palawan">Sabang</location>
,
<collectingRegion id="49BA23D9454A2E1FFAF36092B6D4B38D" box="[1334,1432,151,177]" country="Philippines" name="Palawan" pageId="32" pageNumber="33">Palawan</collectingRegion>
,
<geoCoordinate id="EE4A8BFC454A2E1FFF5260BEB256B3E9" box="[151,282,187,213]" degrees="10" direction="north" minutes="11" orientation="latitude" pageId="32" pageNumber="33" precision="15" seconds="24" value="10.190001">10°11'24&quot;N</geoCoordinate>
<geoCoordinate id="EE4A8BFC454A2E1FFEE760BFB2FEB3E9" box="[290,434,186,213]" degrees="118" direction="east" minutes="54" orientation="longitude" pageId="32" pageNumber="33" precision="15" seconds="00" value="118.9">118°54'00&quot;E</geoCoordinate>
,
<date id="FFC0CBFB454A2E1FFE7A60BFB169B3E9" box="[447,549,186,213]" pageId="32" pageNumber="33" value="2013-02-05">
<collectingDate id="EF843213454A2E1FFE7A60BFB169B3E9" box="[447,549,186,213]" pageId="32" pageNumber="33" value="2013-02-05">5.ii.2013</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B8454A2E1FFD9160BEB649B3E9" authority=", P. J. Matthews, E. M. Agoo &amp; D. A. Madulid" authorityName="P. J. Matthews, E. M. Agoo &amp; D. A. Madulid" box="[596,1285,187,213]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A3129454A2E1FFD9160BEB129B3E9" box="[596,613,187,213]" italics="true" pageId="32" pageNumber="33">A</emphasis>
.
<emphasis id="B90A3129454A2E1FFDB060BEB046B3E9" box="[629,778,187,213]" italics="true" pageId="32" pageNumber="33">macrorrhizos</emphasis>
, P.J. Matthews, E.M. Agoo &amp; D.A. Madulid
</taxonomicName>
leg. (
<collectionCode id="ED6F75FE454A2E1FFA8060BEB6D4B3E9" box="[1349,1432,187,213]" country="Philippines" httpUri="http://grbio.org/cool/i3xh-eq16" name="National Museum of the Philippines" pageId="32" pageNumber="33">MPMP</collectionCode>
,
<collectionCode id="ED6F75FE454A2E1FFF5260DAB3ACB3C5" box="[151,224,223,249]" pageId="32" pageNumber="33">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF01BDB3454A2E1FFF526729B70FB4E8" ID-DOI="http://doi.org/10.5281/zenodo.5766448" ID-Zenodo-Dep="5766448" httpUri="https://zenodo.org/record/5766448/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1836,1860]" targetBox="[151,1435,273,1812]" targetPageId="32">
<paragraph id="8BC1ED3B454A2E1FFF526729B70FB4E8" blockId="32.[151,1437,1836,2004]" pageId="32" pageNumber="33">
<emphasis id="B90A3129454A2E1FFF526729B26EB478" bold="true" box="[151,290,1836,1860]" pageId="32" pageNumber="33">FIGURE 21.</emphasis>
<taxonomicName id="4C7E96B8454A2E1FFEEF6729B1B8B478" authority="Toda &amp; Takano" authorityName="Toda &amp; Takano" authorityYear="2021" box="[298,756,1836,1860]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kinabaluana" status="sp. nov.">
<emphasis id="B90A3129454A2E1FFEEF6729B105B478" box="[298,585,1836,1860]" italics="true" pageId="32" pageNumber="33">Colocasiomyia kinabaluana</emphasis>
Toda &amp; Takano
</taxonomicName>
,
<emphasis id="B90A3129454A2E1FFCC56729B01EB478" bold="true" box="[768,850,1836,1860]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="A2398C52454A2E1FFCC56729B01EB478" box="[768,850,1836,1860]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, oviscapt (lateral view); H, anterior spiracles of puparium (dorsal view); I, puparium (ventral habitus; inset: caudal-most portion); J, mouth hook; K, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–G and K, 1.0 mm in I.
</paragraph>
</caption>
<subSubSection id="C364BEB0454B2E1EFF026093B750B38E" box="[199,1052,150,178]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BC1ED3B454B2E1EFF026093B750B38E" blockId="33.[151,1437,150,357]" box="[199,1052,150,178]" pageId="33" pageNumber="34">
<emphasis id="B90A3129454B2E1EFF026093B22DB38C" bold="true" box="[199,353,150,176]" pageId="33" pageNumber="34">Distribution.</emphasis>
Borneo (Sabah),
<collectingCountry id="F369ADAB454B2E1EFDE36092B1E8B38D" box="[550,676,151,177]" name="Philippines" pageId="33" pageNumber="34">Philippines</collectingCountry>
(Luzon,
<collectingRegion id="49BA23D9454B2E1EFCCF6092B01FB38D" box="[778,851,151,177]" country="Philippines" name="Samar" pageId="33" pageNumber="34">Samar</collectingRegion>
,
<collectingRegion id="49BA23D9454B2E1EFC9B6092B0EDB38D" box="[862,929,151,177]" country="Philippines" name="Southern Leyte" pageId="33" pageNumber="34">Leyte</collectingRegion>
,
<collectingRegion id="49BA23D9454B2E1EFC696092B75CB38E" box="[940,1040,151,178]" country="Philippines" name="Palawan" pageId="33" pageNumber="34">Palawan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454B2E1EFF0260BEB25CB27D" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BC1ED3B454B2E1EFF0260BEB25CB27D" blockId="33.[151,1437,150,357]" pageId="33" pageNumber="34">
<emphasis id="B90A3129454B2E1EFF0260BEB277B3E9" bold="true" box="[199,315,187,213]" pageId="33" pageNumber="34">Remarks.</emphasis>
This species is coupled with
<taxonomicName id="4C7E96B8454B2E1EFD5860BEB007B3E9" box="[669,843,187,213]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sulawesiana">
<emphasis id="B90A3129454B2E1EFD5860BEB1FCB3E9" box="[669,688,187,213]" italics="true" pageId="33" pageNumber="34">C</emphasis>
.
<emphasis id="B90A3129454B2E1EFD0460BEB007B3E9" box="[705,843,187,213]" italics="true" pageId="33" pageNumber="34">sulawesiana</emphasis>
</taxonomicName>
in the molecular phylogenetic tree (BP = 87;
<figureCitation id="1345F1BE454B2E1EFAB560BEB3EAB3C5" captionStart="FIGURE 2" captionStartId="7.[151,250,1764,1788]" captionTargetBox="[151,1433,385,1733]" captionTargetId="figure-124@7.[151,1436,381,1740]" captionTargetPageId="7" captionText="FIGURE 2. Maximum-likelihood (ML) tree of the Colocasiomyia cristata species group (ingroup) and the C. gigantea species group (outgroup), based on DNA sequences of two mitochondrial and six nuclear loci. The tree is rooted by outgroup rooting, and bootstrap percentage (BP) is shown for each internal branch. Within the cristata group, two highly supported (BP = 100) clades (I and II) and three subclades (IIa–c) are recognized." figureDoi="http://doi.org/10.5281/zenodo.5766406" httpUri="https://zenodo.org/record/5766406/files/figure.png" pageId="33" pageNumber="34">Fig. 2</figureCitation>
) and the cladogram (BP = 100;
<figureCitation id="1345F1BE454B2E1EFDE760DAB13BB3C5" box="[546,631,223,249]" captionStart="FIGURE 13" captionStartId="18.[151,250,1328,1352]" captionTargetBox="[151,1433,181,1302]" captionTargetId="figure-22@18.[151,1436,181,1305]" captionTargetPageId="18" captionText="FIGURE 13. The strict consensus tree resulting from the parsimony analysis (PAUP* v4.0a168) of the data matrix of 25 spp. × 51 morphological characters (Appendix 6) for 20 species of the Colocasiomyia cristata group (as ingroup) and five species of the C. gigantea group (as outgroup). The tree is rooted by outgroup rooting, and synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support value (BP). Codes of clades (I and II) and subclades (IIa–c) are shown beside the corresponding branches (see the text for more explanation)." figureDoi="http://doi.org/10.5281/zenodo.5766428" httpUri="https://zenodo.org/record/5766428/files/figure.png" pageId="33" pageNumber="34">Fig. 13</figureCitation>
), and shares the oviscapt longer than the phallus (apodeme + sheath) as a synapomorphy (ch.49-1) with it. However, they can be distinguished from each other by their diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454B2E1EFF02614EB1D2B259" box="[199,670,331,357]" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BC1ED3B454B2E1EFF02614EB1D2B259" blockId="33.[151,1437,150,357]" box="[199,670,331,357]" pageId="33" pageNumber="34">
<emphasis id="B90A3129454B2E1EFF02614EB202B259" bold="true" box="[199,334,331,357]" pageId="33" pageNumber="34">Etymology.</emphasis>
Referring to the
<typeStatus id="54C55399454B2E1EFDC8614EB171B259" box="[525,573,331,357]" pageId="33" pageNumber="34">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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