179 lines
29 KiB
XML
179 lines
29 KiB
XML
<document id="99A86325ADC916A7DFF3313221CEFDF0" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy & Toda, Masanori J." docDate="2021" docId="03D75C2D454E2E1BFF52631BB719B4A9" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia sabahana Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda 2021" docType="treatment" docVersion="7" lastPageNumber="37" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="37" updateTime="1699472124804" updateUser="ExternalLinkService">
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<mods:title id="CE1472C7D4D6AE7248B9BC070C5E3900">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="EAA9B242CBD5A1ED923F369C18F5918B">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="2F276617FED3CADADB3353A8189D86A3">Gao, Jian-Jun</mods:namePart>
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<mods:namePart id="F9932273C05D48D7CD77227B49A48AC7">Hu, Yao-Guang</mods:namePart>
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<mods:namePart id="716CC41B5DB972AA9DC363EBB9979569">Li, Nan-Nan</mods:namePart>
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<mods:namePart id="077122071977B26B85867B8D16F257C2">Yafuso, Masako</mods:namePart>
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<mods:namePart id="FE6293F8D703BEADD5CA85EF43D19E91">Suwito, Awit</mods:namePart>
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<mods:namePart id="7CBECE1AF738B0AC231413D429A360EE">Repin, Rimi</mods:namePart>
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<mods:namePart id="A7899194E5F377730A490A6A4DDCFC0F">Pungga, Runi Anak Sylvester</mods:namePart>
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<mods:namePart id="460DB22CD8A128050E8E2887A7A61C14">Meleng, Paulus Ak</mods:namePart>
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<mods:namePart id="6E9DF0D73DD6AA8D1028B0E33694D1D1">Kaliang, Clement Het</mods:namePart>
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<mods:namePart id="BC552B5D5DDA4481DC62B951DA555597">Chong, Lucy</mods:namePart>
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<mods:namePart id="8DDC60EAAC3F7FD1566D9EF9EB64F6FC">Toda, Masanori J.</mods:namePart>
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<mods:title id="CD270BAE4A71EF0FB9411F9414B9B9AC">Zootaxa</mods:title>
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<mods:date id="7A8BCE20A4F20A026A8FA82A8F88C9CE">2021</mods:date>
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<mods:number id="C03D4F763B494E98728197A1EE801F2B">2021-12-08</mods:number>
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<mods:number id="79B9A560F42A1CB98E599EBFEA639F3B">5079</mods:number>
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<mods:identifier id="55E1A26D14A518CE02025440336E2092" type="DOI">10.11646/zootaxa.5079.1.1</mods:identifier>
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<treatment id="03D75C2D454E2E1BFF52631BB719B4A9" ID-DOI="http://doi.org/10.5281/zenodo.5786179" ID-GBIF-Taxon="191651674" ID-Zenodo-Dep="5786179" LSID="urn:lsid:plazi:treatment:03D75C2D454E2E1BFF52631BB719B4A9" httpUri="http://treatment.plazi.org/id/03D75C2D454E2E1BFF52631BB719B4A9" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection id="C364BEB0454E2E1BFF52631BB04CB005" box="[151,768,798,825]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BC1ED3B454E2E1BFF52631BB04CB005" blockId="36.[151,768,798,825]" box="[151,768,798,825]" pageId="36" pageNumber="37">
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<emphasis id="B90A3129454E2E1BFF52631BB04CB005" bold="true" box="[151,768,798,825]" pageId="36" pageNumber="37">
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2.3
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<taxonomicName id="4C7E96B8454E2E1BFF06631BB2BDB005" ID-CoL="9HY7W" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[195,497,798,825]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sabahana">
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<emphasis id="B90A3129454E2E1BFF06631BB2BDB005" bold="true" box="[195,497,798,825]" italics="true" pageId="36" pageNumber="37">Colocasiomyia sabahana</emphasis>
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species complex, new
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</emphasis>
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</paragraph>
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<subSubSection id="C364BEB0454E2E1BFF526363B29AB44D" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="8BC1ED3B454E2E1BFF526363B73DB099" blockId="36.[151,1437,870,1941]" pageId="36" pageNumber="37">
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<emphasis id="B90A3129454E2E1BFF526363B246B0BC" bold="true" box="[151,266,870,896]" pageId="36" pageNumber="37">Diagnosis</emphasis>
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. Cibarial, posterior sensilla absent (ch.6-1;
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<figureCitation id="1345F1BE454E2E1BFCC66362B012B0BD" box="[771,862,871,897]" captionStart="FIGURE 4" captionStartId="10.[151,250,798,822]" captionTargetBox="[151,1435,181,773]" captionTargetId="figure-22@10.[151,1435,181,774]" captionTargetPageId="10" captionText="FIGURE 4. Characters 5 and 6 on the cibarium (dorsal view) of Colocasiomyia xenalocasiae (Okada) (A), C. cristata de Meijere (B) and C. sarawakana sp. nov. (C). Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensillum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766410" httpUri="https://zenodo.org/record/5766410/files/figure.png" pageId="36" pageNumber="37">Fig. 4C</figureCitation>
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). Phallal sheath gently curved dorsad and slightly projected triangularly at dorsobasal corner in lateral view (ch.40-2, 41-1;
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<figureCitation id="1345F1BE454E2E1BFC0B638EB763B099" box="[974,1071,907,933]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="36" pageNumber="37">Figs 11F</figureCitation>
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,
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<figureCitation id="1345F1BE454E2E1BFBFE638EB729B099" box="[1083,1125,907,933]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">24I</figureCitation>
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).
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</paragraph>
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<paragraph id="8BC1ED3B454E2E1BFF0263ABB29AB44D" blockId="36.[151,1437,870,1941]" pageId="36" pageNumber="37">
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<emphasis id="B90A3129454E2E1BFF0263ABB2EBB0F5" bold="true" box="[199,423,942,969]" pageId="36" pageNumber="37">Shared characters.</emphasis>
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Supracervical setae approximately 4 per side, distally more or less curved, longer than inner occipital setae. Anterior reclinate orbital seta small, situated behind proclinate orbital seta; additional, interfrontal setulae on fronto-orbital plate present (ch.2-0). Eye with stout, dense interfacetal setulae (ch.3-1). Distance between antennal sockets wider than half of socket width (ch.4-0). Cibarial, medial sensilla 3–4 per side, in parallel rows narrower than sensilla campaniformia (ch.5-2;
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<figureCitation id="1345F1BE454E2E1BFD61643AB1B0B765" box="[676,764,1087,1113]" captionStart="FIGURE 4" captionStartId="10.[151,250,798,822]" captionTargetBox="[151,1435,181,773]" captionTargetId="figure-22@10.[151,1435,181,774]" captionTargetPageId="10" captionText="FIGURE 4. Characters 5 and 6 on the cibarium (dorsal view) of Colocasiomyia xenalocasiae (Okada) (A), C. cristata de Meijere (B) and C. sarawakana sp. nov. (C). Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensillum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766410" httpUri="https://zenodo.org/record/5766410/files/figure.png" pageId="36" pageNumber="37">Fig. 4C</figureCitation>
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). Anterior dorsocentral setae behind transverse suture (ch.7- 0). Prescutellar acrostichal setae absent (ch.8-1); acrostichal setulae in 4 rows (ch.9-0). Katepisternal setae small, shorter than postpronotal setae (ch.10-1). Apical scutellar setae nearer to each other than to basal scutellar seta. Costal setae in middle row all weak, trichoid (ch.12-2). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch.13-1). Male abdominal sternites III–V wider than long (ch.16,17,18-0); V medially concaved on posterior margin. Female abdominal sternite III as long as wide (ch.19-0); V longer than wide (ch.21-1). Male abdominal sternite VI not pubescent at least partly, with a pair of divergent processes longer than
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<quantity id="4C8640DE454E2E1BFF17653EB261B669" box="[210,301,1339,1365]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="mm" value="0.1">0.1 mm</quantity>
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, widely separated from each other, bearing prominent seta on apex and small setae on submedial to distal portion (ch.23,25-0, 24,26,27-1;
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<figureCitation id="1345F1BE454E2E1BFDB5655AB1A4B645" box="[624,744,1375,1401]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="36" pageNumber="37">Fig. 9L,M</figureCitation>
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). Epandrium pubescent except for anterior to ventral margin and anteroventral elongation; setae on posteroventral portion as thick as cercal setae (ch.32-0); phragma small (narrower than epandrial anteroventral corner) expansion on antero-subapical margin (ch.33-2). Surstylus vestigial, very narrow process, apically with 1 or 2 tiny, trichoid seta(e) (ch.34,35-1). Cercus pubescent only on dorsal half; ventral/apical portion more or less differentiated from upper portion, with many small setae (ch.38-0). Phallal sheath not pubescent, as thick as wide, apically not bilobed (ch.42-0, 43,44-1;
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<figureCitation id="1345F1BE454E2E1BFBE56616B7ECB511" box="[1056,1184,1555,1581]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="36" pageNumber="37">Figs 11F,G</figureCitation>
|
||
,
|
||
<figureCitation id="1345F1BE454E2E1BFB6B6616B7A4B511" box="[1198,1256,1555,1581]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">24I,J</figureCitation>
|
||
); phallapodeme sclerotized plate directed nearly along or slightly oblique to phallal axis (ch.45-1, 46-0;
|
||
<figureCitation id="1345F1BE454E2E1BFBA36632B78BB56D" box="[1126,1223,1591,1617]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="36" pageNumber="37">Figs 11F</figureCitation>
|
||
,
|
||
<figureCitation id="1345F1BE454E2E1BFB166632B7B0B56D" box="[1235,1276,1591,1617]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">24I</figureCitation>
|
||
). Gonocoxites long, narrow slips (
|
||
<figureCitation id="1345F1BE454E2E1BFEA8665EB283B549" box="[365,463,1627,1653]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="36" pageNumber="37">Figs 11F</figureCitation>
|
||
,
|
||
<figureCitation id="1345F1BE454E2E1BFE1E665EB148B549" box="[475,516,1627,1653]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">24I</figureCitation>
|
||
). Epiproct and hypoproct with pubescence (ch.48-0). Oviscapt shorter than phallus (apodeme + sheath), with ovisensilla only on distal portion occupying less than 1/2 of total length (ch.49-0, 51-1). Puparium (3rd instar larva): segments with stout spicules on ventral surface; anterior spiracle with stalk ending in a whorl of 19–25 long branches; stalk as long as posterior spiracle and longer than longest branch; caudal segments not so elongate, with many small spicules, ending in a V-shaped pair of long posterior spiracles (
|
||
<figureCitation id="1345F1BE454E2E1BFAC166EEB6DCB439" box="[1284,1424,1771,1797]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">Fig. 24M,N</figureCitation>
|
||
). Mouth hook moderately, triangularly expanded medioventrally in lateral view; distal blade as long as basal portion, apically pointed, gently curved downward, with two rows of small, acute teeth on submedial to subapical portion of ventral margin (
|
||
<figureCitation id="1345F1BE454E2E1BFE8F6752B285B44D" box="[330,457,1879,1905]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda & Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="36" pageNumber="37">Fig. 24O,P</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0454E2E1BFF02677EB719B4A9" box="[199,1109,1914,1941]" pageId="36" pageNumber="37" type="reference_group">
|
||
<paragraph id="8BC1ED3B454E2E1BFF02677EB719B4A9" blockId="36.[151,1437,870,1941]" box="[199,1109,1914,1941]" pageId="36" pageNumber="37">
|
||
<emphasis id="B90A3129454E2E1BFF02677EB2DEB4A8" bold="true" box="[199,402,1914,1941]" pageId="36" pageNumber="37">Included species.</emphasis>
|
||
<taxonomicName id="4C7E96B8454E2E1BFE5D677EB1F8B4A9" ID-CoL="9HY7W" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[408,692,1915,1941]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sabahana" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFE5D677EB1F8B4A9" box="[408,692,1915,1941]" italics="true" pageId="36" pageNumber="37">Colocasiomyia sabahana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFD7E677EB059B4A9" bold="true" box="[699,789,1915,1941]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFD7E677EB059B4A9" box="[699,789,1915,1941]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7E96B8454E2E1BFC8E677EB0B9B4A9" ID-CoL="9HS2H" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[843,1013,1915,1941]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sarawakana" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFC8E677EB012B4A9" box="[843,862,1915,1941]" italics="true" pageId="36" pageNumber="37">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129454E2E1BFCA9677EB0B9B4A9" box="[876,1013,1915,1941]" italics="true" pageId="36" pageNumber="37">sarawakana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFC39677EB719B4A9" bold="true" box="[1020,1109,1915,1941]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFC39677EB719B4A9" box="[1020,1109,1915,1941]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |