212 lines
24 KiB
XML
212 lines
24 KiB
XML
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<mods:title id="C4933C5A5FE36CDEE1FC0872D81B5F00">A review of the Diapheridae (Gastropoda: Eupulmonata: Streptaxoidea), with special emphasis on India and Myanmar</mods:title>
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<mods:namePart id="72EB9F7C1A055BD80A3A19942C967D50">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="188D86D71739271E8311CDAF67FF6A17">Hunyadi, András</mods:namePart>
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<mods:namePart id="9250C12643F5E733C46C3C7C51BEB65C">Grego, Jozef</mods:namePart>
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<mods:namePart id="160270C5483AC93124EE641AD58D0E61">Sajan, Sheikh</mods:namePart>
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<mods:namePart id="57A712B7A6DCE75829BCD449B53BF8A3">Chen, Zhe-Yu</mods:namePart>
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<taxonomicName id="4C604D62FFAE4E06FF25FDB1FD593C6E" ID-CoL="7T5KV" authority="Pall-Gergely & Hunyadi" authorityName="Pall-Gergely & Hunyadi" authorityYear="2020" box="[152,699,540,565]" class="Gastropoda" family="Diapheridae" genus="Sinoennea" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="montawana" status="sp. nov.">
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<emphasis id="B914EAF3FFAE4E06FF25FDB1FE753C6E" bold="true" box="[152,407,541,565]" italics="true" pageId="27" pageNumber="709">Sinoennea montawana</emphasis>
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Páll-Gergely & Hunyadi
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,
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<taxonomicNameLabel id="A2275788FFAE4E06FD74FDB1FE133C0E" pageId="27" pageNumber="709" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="135B2A64FFAE4E06FEC7FDF1FE373C2E" box="[378,469,605,629]" captionStart="Fig" captionStartId="28.[115,150,1606,1628]" captionTargetBox="[113,1455,189,1591]" captionTargetId="figure-224@28.[113,1455,189,1591]" captionTargetPageId="28" captionText="Fig. 18. Paratype (A), holotype (B–K), and juvenile shell (L) of Sinoennea montawana Páll-Gergely & Hunyadi, new species." figureDoi="http://doi.org/10.5281/zenodo.5349831" httpUri="https://zenodo.org/record/5349831/files/figure.png" pageId="27" pageNumber="709">Figs. 18</figureCitation>
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,
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<figureCitation id="135B2A64FFAE4E06FE5FFDF1FDF43C2E" box="[482,534,605,629]" captionStart="Fig" captionStartId="33.[136,171,1851,1873]" captionTargetBox="[136,1474,230,1836]" captionTargetId="figure-149@33.[136,1474,229,1836]" captionTargetPageId="33" captionText="Fig. 22. Apertures of Streptaxidae (A) and Diapheridae (B–P). A, Gulella (Huttonella) bicolor (Hutton, 1834); B, Rowsonia beddomei (W.T. Blanford, 1881); C, Platylennea subcostulata (W.T. Blanford, 1881); D, Pupennea planguncula (Benson, 1863); E, Diaphera polita, Páll-Gergely new species; F, Diaphera cylindrelloidea (Stoliczka, 1871); G, Bruggennea cf. bongi (Dance, 1970), H, Parasinoennea ovulum (Bavay & Dautzenberg, 1912); I, Sinoennea otostoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely, Reischütz, Maassen, Grego & Hunyadi, 2020; J, Sinoennea infantilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; K, Sinoennea variabilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; L, Sinoennea sutchariti Páll-Gergely & Hunyadi in Páll-Gergely et al., 2020; M, Sinoennea montawana Páll-Gergely & Hunyadi, new species; N, Sinoennea angustistoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely et al., 2020, O, Sinoennea woodthorpei (Peile, 1929); P, Sinoennea fartoidea (Theobald, 1870). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5349843" httpUri="https://zenodo.org/record/5349843/files/figure.png" pageId="27" pageNumber="709">22M</figureCitation>
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)
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<subSubSection id="C37A656AFFAE4E06FF35FD31FE9B3D8D" pageId="27" pageNumber="709" type="materials_examined">
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<emphasis id="B914EAF3FFAE4E06FF35FD31FED23CEE" bold="true" box="[136,304,669,693]" pageId="27" pageNumber="709">Type material.</emphasis>
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(
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:
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,
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<collectionCode id="ED71AE24FFAE4E06FD97FD32FDA33CEE" box="[554,577,670,693]" pageId="27" pageNumber="709">D</collectionCode>
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:
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<quantity id="4C989B04FFAE4E06FDF6FD31FD7C3CEE" box="[587,670,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="709" unit="mm" value="1.3">1.3 mm</quantity>
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) (
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<specimenCode id="DBC69E9AFFAE4E06FD09FD31FF3E3C8E" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="27" pageNumber="709">HNHM 104856</specimenCode>
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) (
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<figureCitation id="135B2A64FFAE4E06FF4BFD11FE623C8E" box="[246,384,701,725]" captionStart="Fig" captionStartId="28.[115,150,1606,1628]" captionTargetBox="[113,1455,189,1591]" captionTargetId="figure-224@28.[113,1455,189,1591]" captionTargetPageId="28" captionText="Fig. 18. Paratype (A), holotype (B–K), and juvenile shell (L) of Sinoennea montawana Páll-Gergely & Hunyadi, new species." figureDoi="http://doi.org/10.5281/zenodo.5349831" httpUri="https://zenodo.org/record/5349831/files/figure.png" pageId="27" pageNumber="709">Fig. 18B–K</figureCitation>
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),
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<collectingCountry id="F3777671FFAE4E06FE2AFD12FDE63C8D" box="[407,516,702,726]" name="Myanmar" pageId="27" pageNumber="709">Myanmar</collectingCountry>
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,
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<collectingRegion id="49A4F803FFAE4E06FDAEFD11FD6D3C8E" box="[531,655,701,725]" country="Myanmar" name="Shan" pageId="27" pageNumber="709">Shan State</collectingRegion>
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,
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<collectingMunicipality id="6BBBAC9BFFAE4E06FD23FD11FCE53C8E" box="[670,775,701,725]" pageId="27" pageNumber="709">Taunggyi</collectingMunicipality>
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centre WSW,
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<location id="8EBF603AFFAE4E06FE96FD72FE3A3CAE" LSID="urn:lsid:plazi:treatment:03C987F7FFAE4E06FF25FDB1FA4C3CEE:8EBF603AFFAE4E06FE96FD72FE3A3CAE" box="[299,472,733,757]" country="Myanmar" latitude="20.7547" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi" name="Montawa Cave" pageId="27" pageNumber="709" stateProvince="Shan">Montawa Cave</location>
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,
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,
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<geoCoordinate id="EE545026FFAE4E06FD38FD70FCE03CAE" box="[645,770,732,757]" degrees="97" direction="east" minutes="1.057" orientation="longitude" pageId="27" pageNumber="709" precision="1" value="97.01762">97°1.057′E</geoCoordinate>
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,
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<quantity id="4C989B04FFAE4E06FF35FD51FF073D4E" box="[136,229,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="27" pageNumber="709" unit="m" value="1260.0">1,260 m</quantity>
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a.s.l.
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</elevation>
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(locality codes: 2018/34 and 20181005C), coll.
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<collectorName id="26955337FFAE4E06FF03FCB2FECB3D6E" box="[190,297,797,821]" pageId="27" pageNumber="709">K. Okubo</collectorName>
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, J.U. Otani &
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<collectorName id="26955337FFAE4E06FE7EFCB2FDA13D6E" box="[451,579,797,822]" pageId="27" pageNumber="709">A. Hunyadi</collectorName>
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,
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<date id="FFDE1021FFAE4E06FDF0FCB1FCE63D6E" box="[589,772,797,821]" pageId="27" pageNumber="709" value="2018-10-05">
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<collectingDate id="EF9AE9C9FFAE4E06FDF0FCB1FCE63D6E" box="[589,772,797,821]" pageId="27" pageNumber="709" value="2018-10-05">05 October 2018</collectingDate>
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</date>
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.
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<materialsCitation id="3B083CBCFFAE4E06FF35FC92FE553D2E" collectingDate="2018-10-05" collectionCode="H, D, HA" collectorName="K. Okubo & Okubo, J. U & A. Hunyadi" country="Myanmar" elevation="1260" latitude="20.7547" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi" pageId="27" pageNumber="709" specimenCount="1" stateProvince="Shan" typeStatus="paratype">
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<typeStatus id="54DB8843FFAE4E06FF35FC92FF113D0D" box="[136,243,830,854]" pageId="27" pageNumber="709" type="paratype">Paratypes</typeStatus>
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. 1 shell (
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<collectionCode id="ED71AE24FFAE4E06FEECFC91FE8A3D0E" box="[337,360,829,853]" pageId="27" pageNumber="709">H</collectionCode>
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:
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<quantity id="4C989B04FFAE4E06FECFFC91FE273D0E" box="[370,453,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="709" unit="mm" value="2.5">2.5 mm</quantity>
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,
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<collectionCode id="ED71AE24FFAE4E06FE6DFC92FE053D0E" box="[464,487,830,853]" pageId="27" pageNumber="709">D</collectionCode>
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:
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<quantity id="4C989B04FFAE4E06FE4DFC91FDA13D0E" box="[496,579,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="709" unit="mm" value="1.3">1.3 mm</quantity>
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) (Coll.
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<collectionCode id="ED71AE24FFAE4E06FD29FC91FD5C3D0E" box="[660,702,829,853]" pageId="27" pageNumber="709">HA</collectionCode>
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), same data as holotype (
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<figureCitation id="135B2A64FFAE4E06FEF7FCF1FE4D3D2E" box="[330,431,861,885]" captionStart="Fig" captionStartId="28.[115,150,1606,1628]" captionTargetBox="[113,1455,189,1591]" captionTargetId="figure-224@28.[113,1455,189,1591]" captionTargetPageId="28" captionText="Fig. 18. Paratype (A), holotype (B–K), and juvenile shell (L) of Sinoennea montawana Páll-Gergely & Hunyadi, new species." figureDoi="http://doi.org/10.5281/zenodo.5349831" httpUri="https://zenodo.org/record/5349831/files/figure.png" pageId="27" pageNumber="709">Fig. 18A</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BDF36E1FFAE4E06FF35FC31FE9B3D8D" blockId="27.[136,773,925,982]" pageId="27" pageNumber="709">
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<materialsCitation id="3B083CBCFFAE4E06FF35FC31FE973D8D" collectingDate="2018-10-05" collectionCode="H, D" collectorName="K. Okubo & Okubo, J. U & A. Hunyadi" country="Myanmar" elevation="1260" latitude="20.7547" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi" pageId="27" pageNumber="709" specimenCount="3" specimenCount-juvenile="3" stateProvince="Shan">
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<emphasis id="B914EAF3FFAE4E06FF35FC31FE8D3DEE" bold="true" box="[136,367,925,949]" pageId="27" pageNumber="709">Additional material.</emphasis>
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<specimenCount id="9D66FD68FFAE4E06FECEFC31FE3E3DEE" box="[371,476,925,949]" pageId="27" pageNumber="709" type="juvenile">3 juvenile</specimenCount>
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shells (
|
||
<figureCitation id="135B2A64FFAE4E06FD91FC31FD693DEE" box="[556,651,925,949]" captionStart="Fig" captionStartId="28.[115,150,1606,1628]" captionTargetBox="[113,1455,189,1591]" captionTargetId="figure-224@28.[113,1455,189,1591]" captionTargetPageId="28" captionText="Fig. 18. Paratype (A), holotype (B–K), and juvenile shell (L) of Sinoennea montawana Páll-Gergely & Hunyadi, new species." figureDoi="http://doi.org/10.5281/zenodo.5349831" httpUri="https://zenodo.org/record/5349831/files/figure.png" pageId="27" pageNumber="709">Fig. 18L</figureCitation>
|
||
), same data as holotype, coll. HA
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A656AFFAE4E06FF35FC51FF5E3A2E" pageId="27" pageNumber="709" type="diagnosis">
|
||
<paragraph id="8BDF36E1FFAE4E06FF35FC51FF5E3A2E" blockId="27.[136,775,1021,1141]" pageId="27" pageNumber="709">
|
||
<emphasis id="B914EAF3FFAE4E06FF35FC51FF1F3A4E" bold="true" box="[136,253,1021,1045]" pageId="27" pageNumber="709">Diagnosis.</emphasis>
|
||
A small, ovoid
|
||
<taxonomicName id="4C604D62FFAE4E06FE16FC52FD8C3A4E" box="[427,622,1021,1045]" class="Gastropoda" family="Diapheridae" genus="Sinoennea" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B914EAF3FFAE4E06FE16FC52FDF83A4E" box="[427,538,1022,1045]" italics="true" pageId="27" pageNumber="709">Sinoennea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with elongate oval aperture, four normally developed apertural barriers, narrow, rounded umbilicus, and a moderate periumbilical keel.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A656AFFAE4E06FF35FB31FB413F8E" pageId="27" pageNumber="709" type="description">
|
||
<paragraph id="8BDF36E1FFAE4E06FF35FB31FB753F2E" blockId="27.[136,776,1181,2101]" lastBlockId="27.[835,1474,189,373]" pageId="27" pageNumber="709">
|
||
<emphasis id="B914EAF3FFAE4E06FF35FB31FEF13AEE" bold="true" box="[136,275,1181,1205]" pageId="27" pageNumber="709">Description.</emphasis>
|
||
Shell shape ovoid to cylindrical, apex domed, the last three whorls the widest in standard apertural view. Shell colourless, semi-transparent, consisting of 5.5–5.75 whorls. Ribs low, fine, rather regularly arranged. Protoconch consisting of 1.5–1.75 whorls, glossy, almost smooth, very finely granular. Teleoconch sculpture glossier than that of protoconch, ribs rather regularly spaced, although the space between them increases on the last whorl towards aperture. Individual ribs look like low waves “pushed” backwards. Suture deep, whorls bulging, although side of body whorl straight with an outer angle of variable strength. Aperture elongate oval or almost rectangular, elongate. Parietal callus distinct, not “smeared” onto preceding whorl. Apertural dentition 4-fold. Parietal lamella moderately developed, nearly straight. Its outer part is slightly bent towards the parietal denticle, its inner part slightly bent towards the columella. Sinulus oval, opens slightly laterally, relatively small, and not conspicuously isolated due to the relatively weak parietal and palatal teeth. Palatal wall with two blunt, but well-developed denticles that are situated close to each other. Upper one situated on the peristome edge, lower one slightly lower in position and deeper. The inner one is slightly more elevated than the outer one. Columellar lamella situated relatively deep, but clearly visible from apertural view, rather low, elongated ridge-like, oblique to shell axis. Peristome strongly thickened and expanded, not reflected, relatively weak around the sinulus. Basal swelling very slightly indicated, nearly absent, situated relatively close to peristome. Depression anterior to basal swelling on the outer side very shallow (corresponding with the inner palatal tooth). Inner (umbilical) side with a slight depression corresponding with the columellar lamella. Umbilicus open, narrow, rounded. Periumbilical keel moderate, but relatively clearly visible from umbilical view, especially due to the slight furrow on the outer side.
|
||
</paragraph>
|
||
<paragraph id="8BDF36E1FFAE4E06FCFEFE31FB413F8E" blockId="27.[835,1475,413,469]" pageId="27" pageNumber="709">
|
||
Juveniles with a strong parietal and columellar lamella, and a lower palatal tooth (
|
||
<figureCitation id="135B2A64FFAE4E06FB89FE11FB743F8E" box="[1076,1174,445,469]" captionStart="Fig" captionStartId="28.[115,150,1606,1628]" captionTargetBox="[113,1455,189,1591]" captionTargetId="figure-224@28.[113,1455,189,1591]" captionTargetPageId="28" captionText="Fig. 18. Paratype (A), holotype (B–K), and juvenile shell (L) of Sinoennea montawana Páll-Gergely & Hunyadi, new species." figureDoi="http://doi.org/10.5281/zenodo.5349831" httpUri="https://zenodo.org/record/5349831/files/figure.png" pageId="27" pageNumber="709">Fig. 18L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A656AFFAE4E06FCFEFE51FA4C3CEE" pageId="27" pageNumber="709" type="diagnosis">
|
||
<paragraph id="8BDF36E1FFAE4E06FCFEFE51FA4C3CEE" blockId="27.[835,1475,509,693]" pageId="27" pageNumber="709">
|
||
<emphasis id="B914EAF3FFAE4E06FCFEFE51FBA83C4E" bold="true" box="[835,1098,509,533]" pageId="27" pageNumber="709">Differential diagnosis.</emphasis>
|
||
Only two
|
||
<taxonomicName id="4C604D62FFAE4E06FB77FE52FA773C4E" box="[1226,1429,509,533]" class="Gastropoda" family="Diapheridae" genus="Sinoennea" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B914EAF3FFAE4E06FB77FE52FADE3C4E" box="[1226,1340,510,533]" italics="true" pageId="27" pageNumber="709">Sinoennea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are known from
|
||
<collectingCountry id="F3777671FFAE4E06FC6AFDB2FBA73C6D" box="[983,1093,542,566]" name="Myanmar" pageId="27" pageNumber="709">Myanmar</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C604D62FFAE4E06FBEEFDB2FB343C6E" box="[1107,1238,541,565]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="fartoidea">
|
||
<emphasis id="B914EAF3FFAE4E06FBEEFDB2FB343C6E" box="[1107,1238,541,565]" italics="true" pageId="27" pageNumber="709">S. fartoidea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C604D62FFAE4E06FAAFFDB2FA593C6E" baseAuthorityName="Peile" baseAuthorityYear="1929" box="[1298,1467,541,565]" class="Gastropoda" family="Diapheridae" genus="Sinoennea" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="woodthorpei">
|
||
<emphasis id="B914EAF3FFAE4E06FAAFFDB2FA593C6E" box="[1298,1467,541,565]" italics="true" pageId="27" pageNumber="709">S. woodthorpei</emphasis>
|
||
</taxonomicName>
|
||
. The former is more ovoid, has a closed umbilicus, lacks the columellar lamella, and has a very weak lower palatal tooth.
|
||
<taxonomicName id="4C604D62FFAE4E06FCFEFDD2FBA03CCE" baseAuthorityName="Peile" baseAuthorityYear="1929" box="[835,1090,637,661]" class="Gastropoda" family="Diapheridae" genus="Sinoennea" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="709" phylum="Mollusca" rank="species" species="woodthorpei">
|
||
<emphasis id="B914EAF3FFAE4E06FCFEFDD2FBA03CCE" box="[835,1090,637,661]" italics="true" pageId="27" pageNumber="709">Sinoennea woodthorpei</emphasis>
|
||
</taxonomicName>
|
||
is larger with stronger ribbing and its parietal callus is smeared onto the penultimate whorl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |