920 lines
126 KiB
XML
920 lines
126 KiB
XML
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<mods:title id="7D4F168EBF8B35E4710E7DA766F85965">Shifting shell morphology in a Late Miocene-Pliocene land snail species lineage (Gastropoda: Stylommatophora: Spiraxidae), with the description of a new species</mods:title>
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<mods:namePart id="440AD5D68155FCB6327125D9281A89F2">Albesa, Joaquín</mods:namePart>
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<mods:affiliation id="FE06EBA66FF24EF9E60EA8283AC3B17B">Departamento de Botánica y Geología, Universitat de València, 46100 Burjassot (Spain) and Museu Valencià d’Historia Natural, P. O. Box 8460, 46018 Alginet (Spain) joaquin. albesa @ uv. es</mods:affiliation>
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<mods:namePart id="A563E41A20977CD87537483AFD4A319A">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="F1A9E546A3D3F69EE5A45149CC747B8B">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich (Germany) and Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands)</mods:affiliation>
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<mods:title id="C73610A982C7A0C310E0879AA12C95D5">Geodiversitas</mods:title>
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<mods:date id="4C6B50BA3CC8B691A21018BE6CA8887F">2023</mods:date>
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<taxonomicName id="4C194D67BB26FFEFFC71FDE98CC1FDAC" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[963,1239,571,597]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
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<emphasis id="B96DEAF6BB26FFEFFC71FDE98CC1FDAC" bold="true" box="[963,1239,571,597]" italics="true" pageId="7" pageNumber="648">Palaeoglandina turolensis</emphasis>
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<taxonomicNameLabel id="A25E578DBB26FFEFFB6CFDE88D0FFDAD" box="[1246,1305,570,596]" pageId="7" pageNumber="648" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="13222A61BB26FFEFFC45FD8B8C21FD8D" box="[1015,1079,601,628]" captionStart="FIG" captionStartId="8.[132,143,1369,1386]" captionTargetBox="[132,1455,215,1326]" captionTargetId="figure-332@8.[132,1455,215,1326]" captionTargetPageId="8" captionText="FIG. 4. — A, Palaeoglandina turolensis n. sp. from La Gloria 4 (Ruscinian, Teruel Basin): A1-A3, MAP-8405, holotype; B, MAP-8406, paratype; C, MAP-8407, paratype; D, MAP-8408, paratype; E, MAP-8409, paratype; F, MAP-8410, paratype; G, MAP-8411, paratype; H, MAP-8412, paratype; I, MAP-8413, paratype; J, MAP-8414, paratype; K, MAP-8415; L, MAP-8416; M, MAP-8417; N, MAP-8418; O, MAP-8419. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10144338" httpUri="https://zenodo.org/record/10144338/files/figure.png" pageId="7" pageNumber="648">Figs 4</figureCitation>
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A-O; 5A-O; 6G)
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||
</paragraph>
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||
<paragraph id="8BA636E4BB26FFEFFC9FFD4F8DB9FD57" blockId="7.[813,1455,669,686]" box="[813,1455,669,686]" pageId="7" pageNumber="648">
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<subSubSection id="C303656FBB26FFEFFC9FFD328BF2FBF7" pageId="7" pageNumber="648" type="reference_group">
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<paragraph id="8BA636E4BB26FFEFFC9FFD328B7BFCEB" blockId="7.[812,1456,735,786]" pageId="7" pageNumber="648">
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<treatmentCitationGroup id="AB0911CABB26FFEFFC9FFD328B7BFCEB" pageId="7" pageNumber="648">
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<taxonomicName id="4C194D67BB26FFEFFC9FFD328BC5FD0E" ID-CoL="3G57S" authority="Kraus" authorityName="Kraus" box="[813,979,736,760]" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="species" species="antiqua">
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||
<emphasis id="B96DEAF6BB26FFEFFC9FFD328BC5FD0E" box="[813,979,736,760]" italics="true" pageId="7" pageNumber="648">Glandina antiqua</emphasis>
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</taxonomicName>
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||
–
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<treatmentCitation id="0AB810F5BB26FFEFFC59FD328CADFD0E" author="VILANOVA I PIERA J." box="[1003,1211,735,760]" pageId="7" pageNumber="648" year="1863">
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<bibRefCitation id="EF884B15BB26FFEFFC59FD328CADFD0E" author="VILANOVA I PIERA J." box="[1003,1211,735,760]" pageId="7" pageNumber="648" refId="ref13616" refString="VILANOVA I PIERA J. 1863. - Ensayo de descripcion geognostica de la Provincia de Teruel, en sus relaciones con la agricultura de la misma. Junta General de Estadistica, Imprenta Nacional, 342 p." type="book" year="1863">Vilanova I Piera 1863</bibRefCitation>
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||
</treatmentCitation>
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||
: pl. 2, fig. 26 [non
|
||
<bibRefCitation id="EF884B15BB26FFEFFAC7FD328B74FCEB" author="KLEIN A." pageId="7" pageNumber="648" pagination="157 - 164" refId="ref11867" refString="KLEIN A. 1852. - Conchylien der Susswasserkalkformation Wurttembergs. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg 8: 157 - 164. https: // www. biodiversitylibrary. org / page / 7984477" type="journal article" year="1852">Klein, 1852</bibRefCitation>
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||
].
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BA636E4BB26FFEFFC9FFCF98DA6FCBA" blockId="7.[813,1456,811,835]" box="[813,1456,811,835]" pageId="7" pageNumber="648">
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<treatmentCitationGroup id="AB0911CABB26FFEFFC9FFCF98DA6FCBA" box="[813,1456,811,835]" pageId="7" pageNumber="648">
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<taxonomicName id="4C194D67BB26FFEFFC9FFCF98BA9FCBA" authorityName="Reuss" authorityYear="1849" box="[813,959,811,835]" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="species" species="inflata">
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<emphasis id="B96DEAF6BB26FFEFFC9FFCF98BA9FCBA" box="[813,959,811,835]" italics="true" pageId="7" pageNumber="648">Glandina inflata</emphasis>
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</taxonomicName>
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–
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||
<treatmentCitation id="0AB810F5BB26FFEFFC7DFCF98C97FCBA" author="DEREIMS M. A." box="[975,1153,811,835]" page="175" pageId="7" pageNumber="648" year="1898">
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<bibRefCitation id="EF884B15BB26FFEFFC7DFCF98C97FCBA" author="DEREIMS M. A." box="[975,1153,811,835]" pageId="7" pageNumber="648" refId="ref10350" refString="DEREIMS M. A. 1898. - Recherches geologiques dans le Sud de l'Aragon. Le Bigot Freres, Lille, 199 p." type="book" year="1898">Dereims 1898: 175</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 179 [non E.von Martens, 1891].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8BA636E4BB26FFEFFC9FFC8E8C70FC50" blockId="7.[811,1455,860,937]" pageId="7" pageNumber="648">
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<treatmentCitationGroup id="AB0911CABB26FFEFFC9FFC8E8C2EFC77" pageId="7" pageNumber="648">
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<taxonomicName id="4C194D67BB26FFEFFC9FFC8E8C5EFC8D" authorityName="Deperet" authorityYear="1890" box="[813,1096,860,884]" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="variety" species="aquensis" variety="obtusa">
|
||
<emphasis id="B96DEAF6BB26FFEFFC9FFC8E8BCDFC8A" box="[813,987,860,884]" italics="true" pageId="7" pageNumber="648">Glandina aquensis</emphasis>
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||
var.
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||
<emphasis id="B96DEAF6BB26FFEFFBBEFC8E8C5EFC8D" box="[1036,1096,860,884]" italics="true" pageId="7" pageNumber="648">obtusa</emphasis>
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</taxonomicName>
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–
|
||
<treatmentCitation id="0AB810F5BB26FFEFFBD0FC8E8D66FC8D" author="DEPERET C. & SAYN G." box="[1122,1392,860,884]" page="108" pageId="7" pageNumber="648" year="1900">
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<bibRefCitation id="EF884B15BB26FFEFFBD0FC8E8D66FC8D" author="DEPERET C. & SAYN G." box="[1122,1392,860,884]" pageId="7" pageNumber="648" pagination="107 - 126" refId="ref10302" refString="DEPERET C. & SAYN G. 1900. - Monographie de la faune fluvioterrestre du Miocene du Cucuron (Vaucluse). Annales de la Societe linneenne de Lyon 47: 107 - 126. https: // www. biodiversitylibrary. org / page / 43976795" type="journal article" year="1900">Depéret & Sayn 1900: 108</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 1, fig. 77 [non
|
||
<bibRefCitation id="EF884B15BB26FFEFFC12FCA48C3BFC77" author="DEPERET C." box="[928,1069,886,910]" pageId="7" pageNumber="648" refId="ref10275" refString="DEPERET C. 1890. - Les animaux pliocenes du Roussillon. Societe geologique de France, Paleontologie, Memoire no. 3, 194 p." type="book" year="1890">Depéret, 1890</bibRefCitation>
|
||
].
|
||
</treatmentCitationGroup>
|
||
—
|
||
<bibRefCitation id="EF884B15BB26FFEFFBE5FCA58D7CFC77" author="ROYO GOMEZ J." box="[1111,1386,886,911]" pageId="7" pageNumber="648" pagination="1 - 230" refId="ref13107" refString="ROYO GOMEZ J. 1922. - El Mioceno continental iberico y su fauna malacologica. Comision de Investigaciones Paleontologicas y Prehistoricas. Junta Ampliacion Estudios Investigaciones Cientificas, Memorias 30: 1 - 230." type="journal article" year="1922">Royo Gómez 1922: 136-138</bibRefCitation>
|
||
, pl. 12, fig. 17a, b [non
|
||
<bibRefCitation id="EF884B15BB26FFEFFC7BFC438C4AFC50" author="DEPERET C." box="[969,1116,913,937]" pageId="7" pageNumber="648" refId="ref10275" refString="DEPERET C. 1890. - Les animaux pliocenes du Roussillon. Societe geologique de France, Paleontologie, Memoire no. 3, 194 p." type="book" year="1890">Depéret, 1890</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB26FFEFFC9FFC138BF2FBF7" blockId="7.[811,1455,961,1038]" pageId="7" pageNumber="648">
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<treatmentCitationGroup id="AB0911CABB26FFEFFC9FFC138BE3FC0D" pageId="7" pageNumber="648">
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<taxonomicName id="4C194D67BB26FFEFFC9FFC138BCCFC20" authority="Matheron, 1843" authorityName="Matheron" authorityYear="1843" box="[813,986,961,985]" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB26FFEFFC9FFC138BCCFC20" box="[813,986,961,985]" italics="true" pageId="7" pageNumber="648">Glandina aquensis</emphasis>
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||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AB810F5BB26FFEFFC46FC108D02FC20" author="ROYO GOMEZ J." box="[1012,1300,961,986]" page="136-138" pageId="7" pageNumber="648" year="1922">
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<bibRefCitation id="EF884B15BB26FFEFFC46FC108D06FC20" author="ROYO GOMEZ J." box="[1012,1296,961,986]" pageId="7" pageNumber="648" pagination="1 - 230" refId="ref13107" refString="ROYO GOMEZ J. 1922. - El Mioceno continental iberico y su fauna malacologica. Comision de Investigaciones Paleontologicas y Prehistoricas. Junta Ampliacion Estudios Investigaciones Cientificas, Memorias 30: 1 - 230." type="journal article" year="1922">Royo Gómez 1922: 136-138</bibRefCitation>
|
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,
|
||
</treatmentCitation>
|
||
pl. 12, fig. 16a, b [non
|
||
<bibRefCitation id="EF884B15BB26FFEFFCC0FC0E8BFDFC0D" author="KLEIN A." box="[882,1003,988,1012]" pageId="7" pageNumber="648" pagination="157 - 164" refId="ref11867" refString="KLEIN A. 1852. - Conchylien der Susswasserkalkformation Wurttembergs. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg 8: 157 - 164. https: // www. biodiversitylibrary. org / page / 7984477" type="journal article" year="1852">Klein, 1852</bibRefCitation>
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].
|
||
</treatmentCitationGroup>
|
||
—
|
||
<bibRefCitation id="EF884B15BB26FFEFFBAAFC0E8D17FC0D" author="GOMEZ LLUECA F." box="[1048,1281,988,1012]" pageId="7" pageNumber="648" refId="ref11315" refString="GOMEZ LLUECA F. 1931. - Mapa geologico de Espana a escala 1: 50.000. Explicacion de la Hoja nº 567, Teruel. IGME, Madrid, 94 p." type="book" year="1931">Gómez Llueca 1931: 57</bibRefCitation>
|
||
, pl. 51, figs 14-17 [non
|
||
<bibRefCitation id="EF884B15BB26FFEFFCD3FC248BCDFBF7" author="KLEIN A." box="[865,987,1014,1038]" pageId="7" pageNumber="648" pagination="157 - 164" refId="ref11867" refString="KLEIN A. 1852. - Conchylien der Susswasserkalkformation Wurttembergs. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg 8: 157 - 164. https: // www. biodiversitylibrary. org / page / 7984477" type="journal article" year="1852">Klein, 1852</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB26FFEFFC9FFBFA8C4DFAB1" pageId="7" pageNumber="648" type="diagnosis">
|
||
<paragraph id="8BA636E4BB26FFEFFC9FFBFA8C4DFAB1" blockId="7.[811,1457,1063,1352]" pageId="7" pageNumber="648">
|
||
<smallCapsWord id="8D40A038BB26FFEFFC9FFBFA8B8DFBC7" baselines="1081,1082" box="[813,923,1064,1087]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="7" pageNumber="648">DIAGNOSIS</smallCapsWord>
|
||
. — Large oval to fusiform shell, with SH/SW ratio ranging between 1.63-2.05, and with up to 3.9 smoothly to moderately convex whorls separated by shallow sutures. Short spire constituting 10-15% of the total shell height. Protoconch up to 1.9 whorls, ornamented by deep spiral groove flanked by thick, short, prosocyrt riblets towards the beginning of the shell. Large last whorl constituting about 85-90% of total shell height. Large, pyriform and often flattened aperture on palatal margin, corresponding to almost 60% of total shell height; outer lip thin, sharp, non-reflexed, mostly subvertical, angular at its adapical limit, strongly curved at base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB26FFEFFC9FFAB38D4AF859" pageId="7" pageNumber="648" type="materials_examined">
|
||
<paragraph id="8BA636E4BB26FFEFFC9FFAB38BE5F9C8" blockId="7.[811,1457,1376,1586]" pageId="7" pageNumber="648">
|
||
<smallCapsWord id="8D40A038BB26FFEFFC9FFAB38B49FA8F" baselines="1394,1394" box="[813,863,1377,1400]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="648">
|
||
<collectionCode id="ED08AE21BB26FFEFFC9FFAB38B49FA8F" box="[813,863,1377,1400]" pageId="7" pageNumber="648">TYPE</collectionCode>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB26FFEFFCD5FAB48BD1FA8E" baselines="1395" box="[871,967,1382,1399]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="648">
|
||
<collectionCode id="ED08AE21BB26FFEFFCD5FAB48BD1FA8E" box="[871,967,1382,1399]" pageId="7" pageNumber="648">MATERIAL</collectionCode>
|
||
</smallCapsWord>
|
||
. —
|
||
<materialsCitation id="3B713CB9BB26FFEFFC4BFAB38B98FA6A" country="Spain" latitude="40.346806" location="La Gloria 4" longLatPrecision="1" longitude="-1.0671945" municipality="Teruel" pageId="7" pageNumber="648" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B96DEAF6BB26FFEFFC4BFAB38C4EFA80" bold="true" box="[1017,1112,1377,1401]" pageId="7" pageNumber="648">
|
||
<typeStatus id="54A28846BB26FFEFFC4BFAB38C4EFA80" box="[1017,1112,1377,1401]" pageId="7" pageNumber="648" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. MAP-8405: SH:
|
||
<quantity id="4CE19B01BB26FFEFFAAAFAB38D6EFA81" box="[1304,1400,1377,1401]" metricMagnitude="-2" metricUnit="m" metricValue="3.82" pageId="7" pageNumber="648" unit="mm" value="38.2">38.2 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC9CFAA98B9CFA6A" box="[814,906,1403,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="648" unit="mm" value="19.1">19.1 mm</quantity>
|
||
.
|
||
</materialsCitation>
|
||
—
|
||
<materialsCitation id="3B713CB9BB26FFEFFC1FFAA98D08FA6A" box="[941,1310,1403,1427]" pageId="7" pageNumber="648">
|
||
<emphasis id="B96DEAF6BB26FFEFFC1FFAA98C1AFA6A" bold="true" box="[941,1036,1403,1427]" pageId="7" pageNumber="648">
|
||
<typeStatus id="54A28846BB26FFEFFC1FFAA98C1AFA6A" box="[941,1036,1403,1427]" pageId="7" pageNumber="648" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. MAP-8406: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB0CFAA98D0FFA6A" box="[1214,1305,1403,1427]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="7" pageNumber="648" unit="mm" value="36.2">36.2 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFA90FAA98C21FA54" pageId="7" pageNumber="648">
|
||
SW: 18.0 mm. MAP-8407: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC64FA478C25FA54" box="[982,1075,1429,1453]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" pageId="7" pageNumber="648" unit="mm" value="39.5">39.5 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFB8CFA448D05FA3E" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFBDCFA448CDAFA54" box="[1134,1228,1430,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="7" pageNumber="648" unit="mm" value="20.9">20.9 mm</quantity>
|
||
. MAP-8408: SH:
|
||
<quantity id="4CE19B01BB26FFEFFA33FA448B4EFA3E" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="7" pageNumber="648" unit="mm" value="32.9">32.9 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC24FA628BE3FA3E" box="[918,1013,1456,1480]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="7" pageNumber="648" unit="mm" value="16.7">16.7 mm</quantity>
|
||
. MAP-8409: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB1DFA628D19FA3E" box="[1199,1295,1456,1480]" metricMagnitude="-2" metricUnit="m" metricValue="3.3899999999999997" pageId="7" pageNumber="648" unit="mm" value="33.9">33.9 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFAA9FA628C2DFA1B" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFAFEFA628DBAFA3E" box="[1356,1452,1456,1480]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="7" pageNumber="648" unit="mm" value="18.6">18.6 mm</quantity>
|
||
. MAP-8410: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC6BFA198C20FA1B" box="[985,1078,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="7" pageNumber="648" unit="mm" value="31.8">31.8 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFBF3FA198D05FA05" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFBC0FA188CC6FA1B" box="[1138,1232,1482,1506]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="648" unit="mm" value="15.5">15.5 mm</quantity>
|
||
. MAP-8411: SH: 31.0 mm; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC24FA378BE3FA05" box="[918,1013,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="7" pageNumber="648" unit="mm" value="15.4">15.4 mm</quantity>
|
||
. MAP-8412: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB1DFA378D19FA05" box="[1199,1295,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="7" pageNumber="648" unit="mm" value="34.5">34.5 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFAA9FA378C2DF9EE" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFAFEFA378DBAFA05" box="[1356,1452,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="7" pageNumber="648" unit="mm" value="17.2">17.2 mm</quantity>
|
||
. MAP-8413: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC6BFA2D8C20F9EE" box="[985,1078,1535,1559]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="7" pageNumber="648" unit="mm" value="32.2">32.2 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFBF3FA2D8B4DF9C8" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFBC0FA2D8CC6F9EE" box="[1138,1232,1535,1559]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="7" pageNumber="648" unit="mm" value="15.9">15.9 mm</quantity>
|
||
. MAP-8414: SH:
|
||
<quantity id="4CE19B01BB26FFEFFA36FA2D8B41F9C8" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="7" pageNumber="648" unit="mm" value="31.9">31.9 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFCD0F9C88BE5F9C8" box="[866,1011,1562,1586]" pageId="7" pageNumber="648">SW: 16.0 mm.</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB26FFEFFC9FF9998D4AF859" blockId="7.[811,1457,1610,1952]" pageId="7" pageNumber="648">
|
||
<smallCapsWord id="8D40A038BB26FFEFFC9FF9998BBDF998" baselines="1629,1629" box="[813,939,1611,1635]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Additional" pageId="7" pageNumber="648">ADDITIONAL</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB26FFEFFC03F9828C07F998" baselines="1629" box="[945,1041,1616,1633]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="648">
|
||
<collectionCode id="ED08AE21BB26FFEFFC03F9828C07F998" box="[945,1041,1616,1633]" pageId="7" pageNumber="648">MATERIAL</collectionCode>
|
||
</smallCapsWord>
|
||
. —
|
||
<materialsCitation id="3B713CB9BB26FFEFFB8FF9988C7BF984" pageId="7" pageNumber="648">
|
||
MAP-8415: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB5BF9998D5EF99B" box="[1257,1352,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="7" pageNumber="648" unit="mm" value="32.9">32.9 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFA30F9998B4EF984" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="7" pageNumber="648" unit="mm" value="16.4">16.4 mm</quantity>
|
||
. MAP-8416: SH: 33.0 mm;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFBC0F9B78B9CF96E" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFB10F9B78D16F984" box="[1186,1280,1637,1661]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="7" pageNumber="648" unit="mm" value="17.9">17.9 mm</quantity>
|
||
. MAP-8417: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC99F9528B90F96E" box="[811,902,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="7" pageNumber="648" unit="mm" value="33.8">33.8 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFC3DF9528BE1F935" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFC0EF9528C0EF96E" box="[956,1048,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="7" pageNumber="648" unit="mm" value="17.3">17.3 mm</quantity>
|
||
. MAP-8418: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB77F9528D36F96E" box="[1221,1312,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="7" pageNumber="648" unit="mm" value="31.2">31.2 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFAE4F9AD8DB8F96E" box="[1366,1454,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="7" pageNumber="648" unit="mm" value="16.2">16.2 mm</quantity>
|
||
MAP-8419: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC65F9488C22F948" box="[983,1076,1690,1714]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="7" pageNumber="648" unit="mm" value="34.9">34.9 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFBDFF9488CDAF948" box="[1133,1228,1690,1714]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="7" pageNumber="648" unit="mm" value="16.6">16.6 mm</quantity>
|
||
. MAP-8421: SH:
|
||
<quantity id="4CE19B01BB26FFEFFA32F9488B40F935" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="7" pageNumber="648" unit="mm" value="45.6">45.6 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC21F9678BE5F935" box="[915,1011,1717,1741]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="7" pageNumber="648" unit="mm" value="24.9">24.9 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFC4CF9678D07F935" box="[1022,1297,1716,1741]" pageId="7" pageNumber="648">
|
||
MAP-8422: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB1CF9668D1BF935" box="[1198,1293,1716,1740]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="7" pageNumber="648" unit="mm" value="42.5">42.5 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFAABF9678D5EF8F8" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFAF8F9678DBCF935" box="[1354,1450,1717,1741]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="648" unit="mm" value="21.3">21.3 mm</quantity>
|
||
. MAP-8423: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC6BF91D8C21F91F" box="[985,1079,1743,1767]" metricMagnitude="-2" metricUnit="m" metricValue="4.01" pageId="7" pageNumber="648" unit="mm" value="40.1">40.1 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFBC2F91D8CD9F91F" box="[1136,1231,1743,1767]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="7" pageNumber="648" unit="mm" value="21.4">21.4 mm</quantity>
|
||
. MAP-8424: SH:
|
||
<quantity id="4CE19B01BB26FFEFFA30F91D8B40F8F8" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="7" pageNumber="648" unit="mm" value="39.4">39.4 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC21F93B8BE5F8F8" box="[915,1011,1769,1793]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="7" pageNumber="648" unit="mm" value="20.8">20.8 mm</quantity>
|
||
; MNCNI-28900-A: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB4AF93B8D52F8F8" box="[1272,1348,1769,1793]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="648" unit="mm" value="40.0">40 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFAFDF93B8B4AF8E2" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFA32F93B8B41F8E2" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="7" pageNumber="648" unit="mm" value="21.2">21.2 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFCD3F8D68C37F8CF" pageId="7" pageNumber="648">
|
||
MNCNI-28900-D: SH:
|
||
<quantity id="4CE19B01BB26FFEFFBE7F8D68CA7F8E2" box="[1109,1201,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="7" pageNumber="648" unit="mm" value="33.4">33.4 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFB5BF8D68D50F8E2" box="[1257,1350,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="7" pageNumber="648" unit="mm" value="20.5">20.5 mm</quantity>
|
||
. MNCNI-28900-E: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC0DF8CC8C0BF8CF" box="[959,1053,1822,1846]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="7" pageNumber="648" unit="mm" value="44.4">44.4 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFB95F8CC8DB8F8A9" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFBE5F8CC8CA3F8CF" box="[1111,1205,1822,1846]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="7" pageNumber="648" unit="mm" value="24.9">24.9 mm</quantity>
|
||
; MNCNI-11300-E: SH:
|
||
<quantity id="4CE19B01BB26FFEFFC9EF8EB8B9DF8A9" box="[812,907,1849,1873]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="7" pageNumber="648" unit="mm" value="39.8">39.8 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC7AF8EB8C31F8A9" box="[968,1063,1849,1873]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="7" pageNumber="648" unit="mm" value="23.8">23.8 mm</quantity>
|
||
; MVHN-040822JA01: SH:
|
||
<quantity id="4CE19B01BB26FFEFFAF9F8EB8DBCF8A9" box="[1355,1450,1849,1873]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="7" pageNumber="648" unit="mm" value="37.4">37.4 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFC9FF8818D5FF892" box="[813,1353,1875,1899]" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFCEDF8818BA9F892" box="[863,959,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="7" pageNumber="648" unit="mm" value="20.4">20.4 mm</quantity>
|
||
; MVHN-040822JA02: SH:
|
||
<quantity id="4CE19B01BB26FFEFFB56F8818D52F892" box="[1252,1348,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="7" pageNumber="648" unit="mm" value="40.6">40.6 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFAE2F8818B4BF87C" pageId="7" pageNumber="648">
|
||
SW:
|
||
<quantity id="4CE19B01BB26FFEFFA30F8818B4FF87C" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="7" pageNumber="648" unit="mm" value="21.7">21.7 mm</quantity>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CB9BB26FFEFFCD0F8BC8D4AF859" pageId="7" pageNumber="648">
|
||
<collectionCode id="ED08AE21BB26FFEFFCD0F8BC8BBEF87F" box="[866,936,1902,1926]" pageId="7" pageNumber="648">MGM</collectionCode>
|
||
118M: SH:
|
||
<quantity id="4CE19B01BB26FFEFFBAFF8BC8C6DF87C" box="[1053,1147,1902,1926]" metricMagnitude="-2" metricUnit="m" metricValue="4.61" pageId="7" pageNumber="648" unit="mm" value="46.1">46.1 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFB06F8BC8D05F87C" box="[1204,1299,1902,1926]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="7" pageNumber="648" unit="mm" value="24.8">24.8 mm</quantity>
|
||
.
|
||
<collectionCode id="ED08AE21BB26FFEFFAAEF8BC8D76F87F" box="[1308,1376,1902,1926]" pageId="7" pageNumber="648">MGM</collectionCode>
|
||
3732M: SH:
|
||
<quantity id="4CE19B01BB26FFEFFCE9F85A8BACF859" box="[859,954,1928,1952]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" pageId="7" pageNumber="648" unit="mm" value="33.1">33.1 mm</quantity>
|
||
; SW:
|
||
<quantity id="4CE19B01BB26FFEFFC44F85A8C41F859" box="[1014,1111,1928,1952]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="648" unit="mm" value="19.1">19.1 mm</quantity>
|
||
. MAP-8420: protoconch.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB26FFEFFC9FF86B8BCCF812" pageId="7" pageNumber="648" type="etymology">
|
||
<paragraph id="8BA636E4BB26FFEFFC9FF86B8BCCF812" blockId="7.[812,1455,1977,2027]" pageId="7" pageNumber="648">
|
||
<smallCapsWord id="8D40A038BB26FFEFFC9FF86B8BB0F836" baselines="1994,1995" box="[813,934,1977,2000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="7" pageNumber="648">ETYMOLOGY</smallCapsWord>
|
||
. — In reference to the Teruel Basin, where the species is described from.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF66666CBB29FFE0FF36FA8B8C40FA6E" ID-DOI="http://doi.org/10.5281/zenodo.10144338" ID-Zenodo-Dep="10144338" httpUri="https://zenodo.org/record/10144338/files/figure.png" pageId="8" pageNumber="649" startId="8.[132,143,1369,1386]" targetBox="[132,1455,215,1326]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BA636E4BB29FFE0FF36FA8B8C40FA6E" blockId="8.[132,1457,1369,1431]" pageId="8" pageNumber="649">
|
||
<smallCapsWord id="8D40A038BB29FFE0FF36FA8B888BFA90" baselines="1382,1382" box="[132,157,1369,1386]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="649">FIG</smallCapsWord>
|
||
. 4. —
|
||
<emphasis id="B96DEAF6BB29FFE0FF65FA8B88F2FA93" bold="true" box="[215,228,1369,1386]" pageId="8" pageNumber="649">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C194D67BB29FFE0FF5DFA8B89D2FA93" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[239,452,1369,1386]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="649" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
|
||
<emphasis id="B96DEAF6BB29FFE0FF5DFA8B89D2FA93" box="[239,452,1369,1386]" italics="true" pageId="8" pageNumber="649">Palaeoglandina turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB29FFE0FE79FA8B89EAFA93" box="[459,508,1369,1386]" pageId="8" pageNumber="649" rank="species">n. sp.</taxonomicNameLabel>
|
||
from La Gloria 4 (Ruscinian, Teruel Basin):
|
||
<emphasis id="B96DEAF6BB29FFE0FCD9FA8B8BB7FA93" bold="true" box="[875,929,1369,1386]" pageId="8" pageNumber="649">A1-A3</emphasis>
|
||
, MAP-8405, holotype;
|
||
<emphasis id="B96DEAF6BB29FFE0FBD7FA8B8C64FA93" bold="true" box="[1125,1138,1369,1386]" pageId="8" pageNumber="649">B</emphasis>
|
||
, MAP-8406, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FA85FA8B8D53FA93" bold="true" box="[1335,1349,1369,1386]" pageId="8" pageNumber="649">C</emphasis>
|
||
, MAP-8407, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FF6BFAA288F1FA78" bold="true" box="[217,231,1392,1409]" pageId="8" pageNumber="649">D</emphasis>
|
||
, MAP-8408, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FE18FAA289A0FA78" bold="true" box="[426,438,1392,1409]" pageId="8" pageNumber="649">E</emphasis>
|
||
, MAP-8409, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FDC8FAA28A93FA78" bold="true" box="[634,645,1392,1409]" pageId="8" pageNumber="649">F</emphasis>
|
||
, MAP-8410, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FCFBFAA28B41FA78" bold="true" box="[841,855,1392,1409]" pageId="8" pageNumber="649">G</emphasis>
|
||
, MAP-8411, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FBA8FAA28C3EFA78" bold="true" box="[1050,1064,1392,1409]" pageId="8" pageNumber="649">H</emphasis>
|
||
, MAP-8412, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FB5EFAA28CE4FA78" bold="true" box="[1260,1266,1392,1409]" pageId="8" pageNumber="649">I</emphasis>
|
||
, MAP-8413, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FF36FA548898FA6E" bold="true" box="[132,142,1414,1431]" pageId="8" pageNumber="649">J</emphasis>
|
||
, MAP-8414, paratype;
|
||
<emphasis id="B96DEAF6BB29FFE0FEE3FA548948FA6E" bold="true" box="[337,350,1414,1431]" pageId="8" pageNumber="649">K</emphasis>
|
||
, MAP-8415;
|
||
<emphasis id="B96DEAF6BB29FFE0FE7EFA5489C1FA6E" bold="true" box="[460,471,1414,1431]" pageId="8" pageNumber="649">L</emphasis>
|
||
, MAP-8416;
|
||
<emphasis id="B96DEAF6BB29FFE0FDF7FA548A40FA6E" bold="true" box="[581,598,1414,1431]" pageId="8" pageNumber="649">M</emphasis>
|
||
, MAP-8417;
|
||
<emphasis id="B96DEAF6BB29FFE0FD76FA548AC4FA6E" bold="true" box="[708,722,1414,1431]" pageId="8" pageNumber="649">N</emphasis>
|
||
, MAP-8418;
|
||
<emphasis id="B96DEAF6BB29FFE0FCF2FA548B59FA6E" bold="true" box="[832,847,1414,1431]" pageId="8" pageNumber="649">O</emphasis>
|
||
, MAP-8419. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C303656FBB29FFE0FF36FA01890EF966" pageId="8" pageNumber="649" type="materials_examined">
|
||
<paragraph id="8BA636E4BB29FFE0FF36FA018A54F9AD" blockId="8.[131,776,1491,1621]" pageId="8" pageNumber="649">
|
||
<smallCapsWord id="8D40A038BB29FFE0FF36FA0188A3FA10" baselines="1508,1509" box="[132,181,1491,1514]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="649">
|
||
<typeStatus id="54A28846BB29FFE0FF36FA0188A3FA10" box="[132,181,1491,1514]" pageId="8" pageNumber="649">
|
||
<collectionCode id="ED08AE21BB29FFE0FF36FA0188A3FA10" box="[132,181,1491,1514]" pageId="8" pageNumber="649">TYPE</collectionCode>
|
||
</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB29FFE0FF0EFA0A8901FA10" baselines="1509" box="[188,279,1496,1513]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="8" pageNumber="649">
|
||
<collectionCode id="ED08AE21BB29FFE0FF0EFA0A8901FA10" box="[188,279,1496,1513]" pageId="8" pageNumber="649">HORIZON</collectionCode>
|
||
</smallCapsWord>
|
||
. — Marls of Lower Pliocene age (Zanclean, lower Ruscinian, Neogene mammal zone
|
||
<collectionCode id="ED08AE21BB29FFE0FE6FFA3F8A1CF9FC" box="[477,522,1517,1541]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="8" pageNumber="649">MN</collectionCode>
|
||
14) rich in organic matter, with variable shades between grey-brown and purple and with frequent whitish carbonate nodulations, as well as local accumulations of iron oxides/carbonates (
|
||
<bibRefCitation id="EF884B15BB29FFE0FE3AF9EF8A20F9AC" author="ALBESA J. & CALVO J. P. & ALCALA L. & ALONSO ZARZA A. M." box="[392,566,1597,1621]" pageId="8" pageNumber="649" pagination="239 - 264" refId="ref9371" refString="ALBESA J., CALVO J. P., ALCALA L. & ALONSO ZARZA A. M. 1997. - Interpretacion paleoambiental del yacimiento de la Gloria 4 (Plioceno, Fosa de Teruel) a partir del analisis de facies y de asociaciones de gasteropodos y de mamiferos. Cuadernos de Geologia Iberica 22: 239 - 264." type="journal article" year="1997">
|
||
Albesa
|
||
<emphasis id="B96DEAF6BB29FFE0FE7FF9EF89EAF9AC" box="[461,508,1597,1621]" italics="true" pageId="8" pageNumber="649">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB29FFE0FF36F9BF890EF966" blockId="8.[132,775,1644,1695]" pageId="8" pageNumber="649">
|
||
<smallCapsWord id="8D40A038BB29FFE0FF36F9BF88A0F97B" baselines="1662,1662" box="[132,182,1645,1668]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="649">
|
||
<typeStatus id="54A28846BB29FFE0FF36F9BF88A0F97B" box="[132,182,1645,1668]" pageId="8" pageNumber="649">
|
||
<collectionCode id="ED08AE21BB29FFE0FF36F9BF88A0F97B" box="[132,182,1645,1668]" pageId="8" pageNumber="649">TYPE</collectionCode>
|
||
</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB29FFE0FF0DF9A0890AF97A" baselines="1663" box="[191,284,1650,1667]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="8" pageNumber="649">
|
||
<collectionCode id="ED08AE21BB29FFE0FF0DF9A0890AF97A" box="[191,284,1650,1667]" pageId="8" pageNumber="649">LOCALITY</collectionCode>
|
||
</smallCapsWord>
|
||
. —
|
||
<location id="8EC6603FBB29FFE0FEFFF9BF89D5F97D" LSID="urn:lsid:plazi:treatment:03B087F2BB26FFE3FC71FDE989D4F812:8EC6603FBB29FFE0FEFFF9BF89D5F97D" box="[333,451,1644,1668]" country="Spain" latitude="40.346806" longLatPrecision="1" longitude="-1.0671945" municipality="Teruel" name="La Gloria 4" pageId="8" pageNumber="649">La Gloria 4</location>
|
||
(
|
||
<collectingMunicipality id="6BC2AC9EBB29FFE0FE60F9BE8A00F97D" box="[466,534,1644,1668]" pageId="8" pageNumber="649">Teruel</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F30E7674BB29FFE0FD90F9BE8A4DF97D" box="[546,603,1644,1668]" name="Spain" pageId="8" pageNumber="649">Spain</collectingCountry>
|
||
) (
|
||
<geoCoordinate id="EE2D5023BB29FFE0FDC3F9BE8B15F97D" box="[625,771,1644,1668]" degrees="40" direction="north" minutes="20" orientation="latitude" pageId="8" pageNumber="649" precision="1" seconds="48.5" value="40.346806">40°20’48.5”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2D5023BB29FFE0FF34F955891AF966" box="[134,268,1671,1695]" degrees="1" direction="west" minutes="04" orientation="longitude" pageId="8" pageNumber="649" precision="1" seconds="01.9" value="-1.0671945">1°04’01.9”W</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB29FFE0FF36F9648A85F8E7" pageId="8" pageNumber="649" type="distribution">
|
||
<paragraph id="8BA636E4BB29FFE0FF36F9648A85F8E7" blockId="8.[131,777,1718,1822]" pageId="8" pageNumber="649">
|
||
<smallCapsWord id="8D40A038BB29FFE0FF36F964890BF934" baselines="1736,1737" box="[132,285,1718,1742]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stratigraphic" pageId="8" pageNumber="649">STRATIGRAPHIC</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB29FFE0FE96F96E8958F934" baselines="1737" box="[292,334,1724,1741]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="8" pageNumber="649">AND</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB29FFE0FEE4F96E89C4F934" baselines="1737" box="[342,466,1724,1741]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="geographic" pageId="8" pageNumber="649">GEOGRAPHIC</smallCapsWord>
|
||
<smallCapsWord id="8D40A038BB29FFE0FE68F96E8A75F934" baselines="1737" box="[474,611,1724,1741]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="distribution" pageId="8" pageNumber="649">DISTRIBUTION</smallCapsWord>
|
||
. — This species is recorded from the late Tortonian (Turolian,
|
||
<collectionCode id="ED08AE21BB29FFE0FDEAF9038A90F910" box="[600,646,1745,1769]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="8" pageNumber="649">MN</collectionCode>
|
||
12, 7.87 Ma) and Zanclean (Ruscinian,
|
||
<collectionCode id="ED08AE21BB29FFE0FE33F93989B9F8FA" box="[385,431,1771,1795]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="8" pageNumber="649">MN</collectionCode>
|
||
14, 4.4-4.54 Ma) of the Teruel Basin (
|
||
<bibRefCitation id="EF884B15BB29FFE0FF3EF8D48950F8E7" author="ROYO GOMEZ J." box="[140,326,1798,1822]" pageId="8" pageNumber="649" pagination="1 - 230" refId="ref13107" refString="ROYO GOMEZ J. 1922. - El Mioceno continental iberico y su fauna malacologica. Comision de Investigaciones Paleontologicas y Prehistoricas. Junta Ampliacion Estudios Investigaciones Cientificas, Memorias 30: 1 - 230." type="journal article" year="1922">Royo Gómez 1922</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB29FFE0FEE0F8D48A0EF8E7" author="GOMEZ LLUECA F." box="[338,536,1798,1822]" pageId="8" pageNumber="649" refId="ref11315" refString="GOMEZ LLUECA F. 1931. - Mapa geologico de Espana a escala 1: 50.000. Explicacion de la Hoja nº 567, Teruel. IGME, Madrid, 94 p." type="book" year="1931">Gómez Llueca 1931</bibRefCitation>
|
||
; this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB29FFE1FF36F8808B1EFEA8" lastPageId="9" lastPageNumber="650" pageId="8" pageNumber="649" type="description">
|
||
<paragraph id="8BA636E4BB29FFE0FF36F8808903F893" blockId="8.[132,776,1874,2028]" box="[132,277,1874,1900]" pageId="8" pageNumber="649">
|
||
<smallCapsWord id="8D40A038BB29FFE0FF36F8808903F893" baselines="1893,1893" box="[132,277,1874,1900]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="8" pageNumber="649">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB29FFE1FF36F8A08B1EFEA8" blockId="8.[132,776,1874,2028]" lastBlockId="9.[131,776,215,337]" lastPageId="9" lastPageNumber="650" pageId="8" pageNumber="649">
|
||
Large oval to fusiform shell with up to 3.9 smooth to moderately convex whorls separated by shallow, often slightly indented sutures. Short spire consisting of up to 2.9 whorls and accounting for 10-15% of total shell height; obtuse apex. Shell shape, particularly spire height and SH/SW ratio, varies considerably among assemblages (
|
||
<tableCitation id="C69B035FBB29FFE0FB44FA218D5CF9F4" box="[1270,1354,1523,1549]" captionStart="TABLE" captionStartId="11.[133,142,219,236]" captionTargetPageId="11" captionText="TABLE 1. — Shell measurements of several of the studied specimens.Data for species other than P. turolensis n. sp. derive from Matheron (1843) and Montenat & Truc (1971)." httpUri="http://table.plazi.org/id/DF66666CBB2AFFE3FF37FF0988F1FEFA" pageId="8" pageNumber="649" tableUuid="DF66666CBB2AFFE3FF37FF0988F1FEFA">Table 1</tableCitation>
|
||
; see also
|
||
<figureCitation id="13222A61BB29FFE0FC9DF9C08B7DF9D4" box="[815,875,1554,1581]" captionStart="FIG" captionStartId="6.[132,143,1277,1294]" captionTargetBox="[159,1428,215,1235]" captionTargetId="graphics-261@6.[244,1428,215,1235]" captionTargetPageId="6" captionText="FIG. 3. — Results of the morphometric analyses: A, principal components analysis of the Procrustes shape coordinates; shown are the first three axes explaining 88.5% of the total variance; B, thin-plate spline deformation grid illustrating the extreme values for each principal component; deformations were magnified to a factor of 2 to highlight variation of distinct traits; C, reconstructed mean shapes for all material as well as across specimens for each locality." figureDoi="http://doi.org/10.5281/zenodo.10144336" httpUri="https://zenodo.org/record/10144336/files/figure.png" pageId="8" pageNumber="649">Fig. 3</figureCitation>
|
||
). Protoconch up to 1.9 whorls, with nucleus smooth and remaining portion ornamented by deep spiral groove located at approximately one third below upper suture and flanked by thick, short, prosocyrt riblets, creating altogether a pattern remotely resembling a fern pinna with pinnules. Passage into teleoconch marked by irregularly shaped scar and onset of growth lines. Teleoconch formed by 2 whorls. Teleoconch ornamentation consists of strong and numerous growth lines, weakly prosocline to almost orthocline, crossed by fine, almost imperceptible spiral grooves, which give rise to weak reticulation. Large last whorl, about 85-90% of total shell height. Large aperture constituting almost 60 % of total shell height; pyriform, acuminate at suture and rounded at base, often flattened at palatal edge and provided with wellmarked notch at boundary between columellar and basal edges; peristome simple with columellar edge arched and forming callosity which is obliquely truncated at base; outer lip thin, sharp, not reflexed, mostly subvertical, often adapically thickened and strongly curved at base. Not umbilicate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303656FBB28FFE3FF36FEA589D4F812" lastPageId="11" lastPageNumber="652" pageId="9" pageNumber="650" type="discussion">
|
||
<paragraph id="8BA636E4BB28FFE1FF36FEA588FFFE76" blockId="9.[130,777,375,2028]" box="[132,233,375,401]" pageId="9" pageNumber="650">
|
||
<smallCapsWord id="8D40A038BB28FFE1FF36FEA588FFFE76" baselines="394,394" box="[132,233,375,401]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="9" pageNumber="650">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB28FFE1FF36FE448C04FA77" blockId="9.[130,777,375,2028]" lastBlockId="9.[811,1457,215,2028]" pageId="9" pageNumber="650">
|
||
Of the 23 species of
|
||
<taxonomicName id="4C194D67BB28FFE1FEEAFE4489E1FE49" authorityName="Wenz" authorityYear="1914" box="[344,503,406,432]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96DEAF6BB28FFE1FEEAFE4489E1FE49" box="[344,503,406,432]" italics="true" pageId="9" pageNumber="650">Palaeoglandina</emphasis>
|
||
</taxonomicName>
|
||
currently recognized, the majority (18) come from Lower Eocene (Ypresian) to Middle Miocene (Langhian) deposits (e.g.
|
||
<bibRefCitation id="EF884B15BB28FFE1FDB0FE058A96FE09" author="WENZ W." box="[514,640,470,497]" pageId="9" pageNumber="650" pagination="737 - 1068" refId="ref13770" refString="WENZ W. 1923. - Fossilium Catalogus I: Animalia. Gastropoda extramarina tertiaria. Pars 20. W. Junk, Berlin: 737 - 1068. https: // www. biodiversitylibrary. org / page / 40596679" type="book chapter" year="1923">Wenz 1923</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FD3FFE058AE9FE09" author="TRUC G." box="[653,767,470,497]" pageId="9" pageNumber="650" pagination="79 - 129" refId="ref13525" refString="TRUC G. 1971. - Gasteropodes continentaux neogenes du Bassin rhodanien. Document du Laboratoire geologique de l'Universite de Lyon, hors serie 11: 79 - 129." type="journal article" year="1971">Truc 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FF36FE2589BEFDE8" author="LE RENARD J. & PACAUD J." box="[132,424,502,529]" pageId="9" pageNumber="650" pagination="65 - 132" refId="ref11904" refString="LE RENARD J. & PACAUD J. 1995. - Revision des mollusques Paleogenes du Bassin de Paris. II. Liste des references primaires des especes. Cossmanniana 3: 65 - 132." type="journal article" year="1995">Le Renard & Pacaud 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FE05FE248A56FDE9" author="BINDER H." box="[439,576,502,528]" pageId="9" pageNumber="650" pagination="189 - 229" refId="ref9922" refString="BINDER H. 2004. - Terrestrial, freshwater and brachyhaline Gastropoda from Lower Miocene deposits of Oberdorf (Styria, Austria). Annalen des Naturhistorischen Museum in Wien 105 A: 189 - 229. https: // www. jstor. org / stable / 41702058" type="journal article" year="2004">Binder 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FDFDFE2488DEFDC9" author="HARZHAUSER M. & NEUBAUER T. A. & GEORGOPOULOU E. & HARL J." pageId="9" pageNumber="650" pagination="819 - 908" refId="ref11456" refString="HARZHAUSER M., NEUBAUER T. A., GEORGOPOULOU E. & HARL J. 2014 a. - The Early Miocene (Burdigalian) mollusc fauna of the North Bohemian Lake (Most Basin). Bulletin of Geosciences 89 (4): 819 - 908. https: // doi. org / 10.3140 / bull. geosci. 1503" type="journal article" year="2014">
|
||
Harzhauser
|
||
<emphasis id="B96DEAF6BB28FFE1FD61FE258B11FDE9" box="[723,775,502,528]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
2014a
|
||
</bibRefCitation>
|
||
,b;
|
||
<bibRefCitation id="EF884B15BB28FFE1FF5BFDC489CFFDC9" author="HOLTKE O. & RASSER M. W." box="[233,473,534,561]" pageId="9" pageNumber="650" pagination="267 - 302" refId="ref11740" refString="HOLTKE O. & RASSER M. W. 2017. - Land snails from the Miocene Steinheim impact crater lake sediments (Baden-Wurttemberg, South Germany). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 285 (3): 267 - 302. https: // doi. org / 10.1127 / njgpa / 2017 / 0681" type="journal article" year="2017">Höltke & Rasser 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FE57FDC48AB3FDC9" author="HOLTKE O. & SALVADOR R. B. & RASSER M. W." box="[485,677,534,560]" pageId="9" pageNumber="650" pagination="17 - 44" refId="ref11800" refString="HOLTKE O., SALVADOR R. B. & RASSER M. W. 2018. - Miocene continental gastropods from the southern margin of the Swabian Alb (Baden-Wurttemberg, SW Germany). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 287 (1): 17 - 44. https: // doi. org / 10.1127 / njgpa / 2018 / 0704" type="journal article" year="2018">
|
||
Höltke
|
||
<emphasis id="B96DEAF6BB28FFE1FD84FDC58A7FFDC9" box="[566,617,534,560]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FD00FDC4891FFDA9" author="MOLLUSCABASE" pageId="9" pageNumber="650" refId="ref12299" refString="MOLLUSCABASE (eds) 2023. - MolluscaBase. Accessed at https: // www. molluscabase. org on 2023 - 07 - 25. https: // doi. org / 10.14284 / 448" type="book" year="2023">MolluscaBase 2023</bibRefCitation>
|
||
). Only five species have been recorded from the Late Miocene (Tortonian) to the Late Pliocene (Piacenzian)/ Early Pleistocene (Calabrian) (e.g.
|
||
<bibRefCitation id="EF884B15BB28FFE1FDBDFDA48A98FD69" author="WENZ W." box="[527,654,630,656]" pageId="9" pageNumber="650" pagination="737 - 1068" refId="ref13770" refString="WENZ W. 1923. - Fossilium Catalogus I: Animalia. Gastropoda extramarina tertiaria. Pars 20. W. Junk, Berlin: 737 - 1068. https: // www. biodiversitylibrary. org / page / 40596679" type="book chapter" year="1923">Wenz 1923</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FD2DFDA488AAFD49" author="ESU D. & GIROTTI O. & KOTSAKIS T." pageId="9" pageNumber="650" pagination="101 - 119" refId="ref10437" refString="ESU D., GIROTTI O. & KOTSAKIS T. 1993. - Palaeobiogeographical observations on Villafranchian molluscs of Italy. Scripta Geologica, special issue 2: 101 - 119." type="journal article" year="1993">
|
||
Esu
|
||
<emphasis id="B96DEAF6BB28FFE1FD62FDA58B11FD69" box="[720,775,630,656]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
1993
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FF7BFD44899AFD49" author="GLIOZZI E. & ABBAZZI L. & ARGENTI P. & AZZAROLI A. & CALOI L. & CAPASSO BARBATO L. & DI STEFANO G. & ESU D. & FICCARELLI G. & GIROTTI O. & KOTSAKI T. & MASINI F. & MAZZA P. & MEZZABOTTA C. & PAOMBO M. R. & PETRONIO C. & ROOK L. & SALA B. & SARDELLA R. & ZANALDA E. & TORRE D." box="[201,396,662,688]" pageId="9" pageNumber="650" pagination="369 - 388" refId="ref10838" refString="GLIOZZI E., ABBAZZI L., ARGENTI P., AZZAROLI A., CALOI L., CAPASSO BARBATO L., DI STEFANO G., ESU D., FICCARELLI G., GIROTTI O., KOTSAKI T., MASINI F., MAZZA P., MEZZABOTTA C., PAOMBO M. R., PETRONIO C., ROOK L., SALA B., SARDELLA R., ZANALDA E. & TORRE D. 1997. - Biochronology of selected mammals, molluscs and ostracods from the Middle Pliocene to the Late Pleistocene in Italy. The state of the art. Rivista Italiana di Paleontologia e Stratigrafia 103: 369 - 388. https: // doi. org / 10.13130 / 2039 - 4942 / 5299" type="journal article" year="1997">
|
||
Gliozzi
|
||
<emphasis id="B96DEAF6BB28FFE1FEAFFD448959FD49" box="[285,335,662,688]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FE2BFD448A80FD49" author="MONTENAT C. & TRUC G." box="[409,662,662,688]" pageId="9" pageNumber="650" pagination="52 - 58" refId="ref12335" refString="MONTENAT C. & TRUC G. 1971. - Pliocene superieur et Villafranchien dans le Levant espagnol (Provinces de Murcia et d'Alicante). Boletin Geologico y Minero 82 (1): 52 - 58." type="journal article" year="1971">Montenat & Truc 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FD11FD448917FD29" author="MONTOYA P. & ALBERDI M. T. & BLAZQUEZ A. M. & BARBADILLO L. J. & FUMANAL P. & VAN DER MADE J. & MARIN J. M. & MOLINA A. & MORALES J. & MURELAGA X. & PENALVER E. & ROBLES F. & RUIZ BUSTOS A. & SANCHEZ A. & SANCHIZ B. & SORIA D. & SZYNDLAR Z." pageId="9" pageNumber="650" pagination="127 - 161" refId="ref12374" refString="MONTOYA P., ALBERDI M. T., BLAZQUEZ A. M., BARBADILLO L. J., FUMANAL P., VAN DER MADE J., MARIN J. M., MOLINA A., MORALES J., MURELAGA X., PENALVER E., ROBLES F., RUIZ BUSTOS A., SANCHEZ A., SANCHIZ B., SORIA D. & SZYNDLAR Z. 1999 a. - La fauna del Pleistoceno inferior de la Sierra de Quibas (Abanilla, Murcia). Estudios Geologicos 55 (3 - 4): 127 - 161. https: // doi. org / 10.3989 / egeol. 99553 - 4171" type="journal article" year="1999">
|
||
Montoya
|
||
<emphasis id="B96DEAF6BB28FFE1FF36FD6488AEFD36" box="[132,184,693,719]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
1999a
|
||
</bibRefCitation>
|
||
, b, 2001;
|
||
<bibRefCitation id="EF884B15BB28FFE1FEC2FD678A55FD29" author="MOLLUSCABASE" box="[368,579,693,720]" pageId="9" pageNumber="650" refId="ref12299" refString="MOLLUSCABASE (eds) 2023. - MolluscaBase. Accessed at https: // www. molluscabase. org on 2023 - 07 - 25. https: // doi. org / 10.14284 / 448" type="book" year="2023">MolluscaBase 2023</bibRefCitation>
|
||
) and thus fall in a similar stratigraphic range as
|
||
<taxonomicName id="4C194D67BB28FFE1FE0CFD078AD0FD16" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[446,710,725,751]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
|
||
<emphasis id="B96DEAF6BB28FFE1FE0CFD078AD0FD16" box="[446,710,725,751]" italics="true" pageId="9" pageNumber="650">Palaeoglandina turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB28FFE1FD7FFD048B11FD09" box="[717,775,726,752]" pageId="9" pageNumber="650" rank="species">n. sp.</taxonomicNameLabel>
|
||
Two species have been located in supposedly Late Miocene sites, i.e.
|
||
<taxonomicName id="4C194D67BB28FFE1FF6DFCC78A8CFCD6" authority="(Matheron, 1843)" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[223,666,789,815]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FF6DFCC789C4FCD6" box="[223,466,789,815]" italics="true" pageId="9" pageNumber="650">Palaeoglandina aquensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FE52FCC78A84FCD6" author="MATHERON P." box="[480,658,789,815]" pageId="9" pageNumber="650" pagination="81 - 341" refId="ref12003" refString="MATHERON P. 1843. - Catalogue methodique et descriptif des corps organises fossiles du department des Bouches-du-Rhone et lieux circonvoisins. Repertoire des Travaux de la Societe de Statistique de Marseille 6: 81 - 341." type="journal article" year="1843">Matheron, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, originally described from the Burdigalian of Aix-en-Provence (Bouches-du-Rhône,
|
||
<collectingCountry id="F30E7674BB28FFE1FEB2FC84895AFC89" box="[256,332,854,880]" name="France" pageId="9" pageNumber="650">France</collectingCountry>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FEE6FC8489AAFC96" author="ESU D." box="[340,444,853,880]" pageId="9" pageNumber="650" pagination="328 - 354" refId="ref10470" refString="ESU D. 1999. - Contribution to the knowledge of Neogene climatic changes in western and central Europa by means of non-marine molluscs, in AGUSTI J., ROOK L. & ANDREWS P. (eds), The Evolution of Neogene Terrestrial Ecosystems in Europa. Hominoid Evolution and Climatic Change in Europa. Vol. I. Cambridge University Press: 328 - 354. https: // doi. org / 10.1017 / cbo 9780511542329.016" type="book chapter" year="1999">Esu 1999</bibRefCitation>
|
||
) but also cited from Tortonian strata (e.g.
|
||
<bibRefCitation id="EF884B15BB28FFE1FF4AFCA7898EFC76" author="SCHLOSSER M." box="[248,408,885,911]" pageId="9" pageNumber="650" pagination="1 - 42" refId="ref13341" refString="SCHLOSSER M. 1907. - Ueber Saugetiere und Susswassergastropoden aus Pliocanablagerungen Spaniens und uber die naturliche Grenze von Miocan und Pliocan. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie 2: 1 - 42. https: // www. biodiversitylibrary. org / page / 44553775" type="journal article" year="1907">Schlosser 1907</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FE15FCA78A64FC76" author="ROYO GOMEZ J." box="[423,626,885,911]" pageId="9" pageNumber="650" pagination="1 - 230" refId="ref13107" refString="ROYO GOMEZ J. 1922. - El Mioceno continental iberico y su fauna malacologica. Comision de Investigaciones Paleontologicas y Prehistoricas. Junta Ampliacion Estudios Investigaciones Cientificas, Memorias 30: 1 - 230." type="journal article" year="1922">Royo Gómez 1922</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="4C194D67BB28FFE1FD06FCA489D3FC56" authority="(Matheron, 1843)" baseAuthorityName="Matheron" baseAuthorityYear="1843" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="galloprovincialis">
|
||
<emphasis id="B96DEAF6BB28FFE1FD06FCA488EDFC56" italics="true" pageId="9" pageNumber="650">P. galloprovincialis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FEBAFC4789ABFC56" author="MATHERON P." box="[264,445,917,943]" pageId="9" pageNumber="650" pagination="81 - 341" refId="ref12003" refString="MATHERON P. 1843. - Catalogue methodique et descriptif des corps organises fossiles du department des Bouches-du-Rhone et lieux circonvoisins. Repertoire des Travaux de la Societe de Statistique de Marseille 6: 81 - 341." type="journal article" year="1843">Matheron, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, described from the Tortonian (?) of Peyrolles (Bouches-du-Rhône,
|
||
<collectingCountry id="F30E7674BB28FFE1FDBFFC678A4DFC36" box="[525,603,949,975]" name="France" pageId="9" pageNumber="650">France</collectingCountry>
|
||
). (Note that the age of the deposits at Peyrolles is uncertain; it was indicated as “Pontien” by
|
||
<bibRefCitation id="EF884B15BB28FFE1FE85FC2789D0FBF6" author="WENZ W." box="[311,454,1013,1039]" pageId="9" pageNumber="650" pagination="737 - 1068" refId="ref13770" refString="WENZ W. 1923. - Fossilium Catalogus I: Animalia. Gastropoda extramarina tertiaria. Pars 20. W. Junk, Berlin: 737 - 1068. https: // www. biodiversitylibrary. org / page / 40596679" type="book chapter" year="1923">Wenz (1923)</bibRefCitation>
|
||
, while species recorded from that site were attributed to both the Early Miocene (mammal stage MN4) and the Late Miocene (MN8) by
|
||
<bibRefCitation id="EF884B15BB28FFE1FDD1FBE78ADFFBB7" author="ESU D." box="[611,713,1076,1103]" pageId="9" pageNumber="650" pagination="328 - 354" refId="ref10470" refString="ESU D. 1999. - Contribution to the knowledge of Neogene climatic changes in western and central Europa by means of non-marine molluscs, in AGUSTI J., ROOK L. & ANDREWS P. (eds), The Evolution of Neogene Terrestrial Ecosystems in Europa. Hominoid Evolution and Climatic Change in Europa. Vol. I. Cambridge University Press: 328 - 354. https: // doi. org / 10.1017 / cbo 9780511542329.016" type="book chapter" year="1999">Esu 1999</bibRefCitation>
|
||
). The possible
|
||
<typeStatus id="54A28846BB28FFE1FF69FB87893AFB96" box="[219,300,1109,1135]" pageId="9" pageNumber="650" type="syntype">syntype</typeStatus>
|
||
series of both species have been located in the MHNM. Since the entire Matheron collection is deposited in the MHNM (C. Borrely pers. comm.), both the specimens of
|
||
<taxonomicName id="4C194D67BB28FFE1FF13FB678919FB37" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[161,271,1205,1230]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FF13FB678919FB37" box="[161,271,1205,1230]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
</taxonomicName>
|
||
from Aix and those of
|
||
<taxonomicName id="4C194D67BB28FFE1FDBBFB678ADCFB37" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[521,714,1204,1230]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="galloprovincialis">
|
||
<emphasis id="B96DEAF6BB28FFE1FDBBFB678ADCFB37" box="[521,714,1204,1230]" italics="true" pageId="9" pageNumber="650">P. galloprovincialis</emphasis>
|
||
</taxonomicName>
|
||
from Peyrolles could be the ones used by Matheron to describe the two species and thus belong to the type series. We examined
|
||
<specimenCount id="9D1FFD6DBB28FFE1FF31FAC6890AFAD4" box="[131,284,1299,1326]" count="2" pageId="9" pageNumber="650" type="generic">two specimens</specimenCount>
|
||
whose measurements are similar to those given by Matheron (
|
||
<taxonomicName id="4C194D67BB28FFE1FE95FAE68981FAB4" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[295,407,1332,1357]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FE95FAE68981FAB4" box="[295,407,1332,1357]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
</taxonomicName>
|
||
: MHNM 6034.9314.1,
|
||
<figureCitation id="13222A61BB28FFE1FD02FAE18880FA94" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">
|
||
Fig. 6A;
|
||
<taxonomicName id="4C194D67BB28FFE1FF36FA868880FA94" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[132,150,1364,1389]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="galloprovincialis">
|
||
<emphasis id="B96DEAF6BB28FFE1FF36FA868880FA94" box="[132,150,1364,1389]" italics="true" pageId="9" pageNumber="650">P</emphasis>
|
||
</taxonomicName>
|
||
</figureCitation>
|
||
.galloprovincialis: MHNM 6034.9317.1,
|
||
<figureCitation id="13222A61BB28FFE1FDEDFA818AA4FA94" box="[607,690,1363,1389]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6B</figureCitation>
|
||
). However, none of the specimens match the figures of
|
||
<bibRefCitation id="EF884B15BB28FFE1FD28FAA188D3FA54" author="MATHERON P." pageId="9" pageNumber="650" pagination="81 - 341" refId="ref12003" refString="MATHERON P. 1843. - Catalogue methodique et descriptif des corps organises fossiles du department des Bouches-du-Rhone et lieux circonvoisins. Repertoire des Travaux de la Societe de Statistique de Marseille 6: 81 - 341." type="journal article" year="1843">Matheron (1843</bibRefCitation>
|
||
: pl. 34, figs 8, 9; pl. 34, fig. 18), so we refrain from designating
|
||
<typeStatus id="54A28846BB28FFE1FEA3FA618994FA34" box="[273,386,1459,1485]" pageId="9" pageNumber="650" type="lectotype">lectotypes</typeStatus>
|
||
until the material can be studied in greater detail. Among the above-mentioned records of
|
||
<taxonomicName id="4C194D67BB28FFE1FF34FA2188E5F9F5" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[134,243,1523,1548]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FF34FA2188E5F9F5" box="[134,243,1523,1548]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
</taxonomicName>
|
||
in the Iberian Peninsula, only the specimens from the Teruel Basin have been located.
|
||
<taxonomicName id="4C194D67BB28FFE1FDBEF9C0888DF9B5" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
|
||
<emphasis id="B96DEAF6BB28FFE1FDBEF9C0888DF9B5" italics="true" pageId="9" pageNumber="650">Palaeoglandina turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB28FFE1FF16F9E188F4F9B4" box="[164,226,1587,1613]" pageId="9" pageNumber="650" rank="species">n. sp.</taxonomicNameLabel>
|
||
, besides being much younger, clearly differs from
|
||
<taxonomicName id="4C194D67BB28FFE1FF34F98188E3F995" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[134,245,1619,1644]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FF34F98188E3F995" box="[134,245,1619,1644]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
</taxonomicName>
|
||
in its shell morphology.
|
||
<taxonomicName id="4C194D67BB28FFE1FDBBF9808B11F995" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[521,775,1618,1644]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="aquensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FDBBF9808B11F995" box="[521,775,1618,1644]" italics="true" pageId="9" pageNumber="650">Palaeoglandina aquensis</emphasis>
|
||
</taxonomicName>
|
||
has, according to the description, figures and measurements provided by
|
||
<bibRefCitation id="EF884B15BB28FFE1FEBCF94089E9F955" author="MATHERON P." box="[270,511,1682,1709]" pageId="9" pageNumber="650" pagination="81 - 341" refId="ref12003" refString="MATHERON P. 1843. - Catalogue methodique et descriptif des corps organises fossiles du department des Bouches-du-Rhone et lieux circonvoisins. Repertoire des Travaux de la Societe de Statistique de Marseille 6: 81 - 341." type="journal article" year="1843">Matheron (1843: 279</bibRefCitation>
|
||
, pl. 34, figs 8, 9), a SH/ SW ratio greater than 2 (
|
||
<figureCitation id="13222A61BB28FFE1FE28F96089FBF935" box="[410,493,1714,1740]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6A</figureCitation>
|
||
;
|
||
<tableCitation id="C69B035FBB28FFE1FE4BF9608A5CF935" box="[505,586,1714,1740]" captionStart="TABLE" captionStartId="11.[133,142,219,236]" captionTargetPageId="11" captionText="TABLE 1. — Shell measurements of several of the studied specimens.Data for species other than P. turolensis n. sp. derive from Matheron (1843) and Montenat & Truc (1971)." httpUri="http://table.plazi.org/id/DF66666CBB2AFFE3FF37FF0988F1FEFA" pageId="9" pageNumber="650" tableUuid="DF66666CBB2AFFE3FF37FF0988F1FEFA">Table 1</tableCitation>
|
||
), as well as a less obtuse spire and a proportionally lower last whorl than the new species from Teruel, whose SH/SW ratio is on average less than 2 (
|
||
<tableCitation id="C69B035FBB28FFE1FEB8F8C0894DF8D5" box="[266,347,1810,1836]" captionStart="TABLE" captionStartId="11.[133,142,219,236]" captionTargetPageId="11" captionText="TABLE 1. — Shell measurements of several of the studied specimens.Data for species other than P. turolensis n. sp. derive from Matheron (1843) and Montenat & Truc (1971)." httpUri="http://table.plazi.org/id/DF66666CBB2AFFE3FF37FF0988F1FEFA" pageId="9" pageNumber="650" tableUuid="DF66666CBB2AFFE3FF37FF0988F1FEFA">Table 1</tableCitation>
|
||
) and has a more convex shell.
|
||
<bibRefCitation id="EF884B15BB28FFE1FD02F8C088D9F8B5" author="DEPERET C." pageId="9" pageNumber="650" refId="ref10275" refString="DEPERET C. 1890. - Les animaux pliocenes du Roussillon. Societe geologique de France, Paleontologie, Memoire no. 3, 194 p." type="book" year="1890">Depéret (1890)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C194D67BB28FFE1FEF7F8E08A3BF8B2" baseAuthorityName="Deperet" baseAuthorityYear="1890" box="[325,557,1841,1868]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="variety" species="aquensis" variety="obtusa">
|
||
<emphasis id="B96DEAF6BB28FFE1FEF7F8E089A2F8B2" box="[325,436,1842,1867]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
var.
|
||
<emphasis id="B96DEAF6BB28FFE1FE58F8E38A3BF8B2" box="[490,557,1841,1867]" italics="true" pageId="9" pageNumber="650">obtusa</emphasis>
|
||
</taxonomicName>
|
||
for specimens from the Pliocene of Roussillon with a slightly more obtuse spire and slightly more convex morphology. This author also indicated the possibility that the new variety is the same as
|
||
<taxonomicName id="4C194D67BB28FFE1FF37F8638A3EF835" authority="(Gobanz, 1854)" baseAuthorityName="Gobanz" baseAuthorityYear="1854" box="[133,552,1969,1996]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="porrecta">
|
||
<emphasis id="B96DEAF6BB28FFE1FF37F863896CF832" box="[133,378,1969,1995]" italics="true" pageId="9" pageNumber="650">Palaeoglandina porrecta</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FE37F8638A36F835" author="GOBANZ J." box="[389,544,1969,1996]" pageId="9" pageNumber="650" pagination="180 - 201" refId="ref10979" refString="GOBANZ J. 1854. - Die fossilen Land- und Susswasser-Mollusken des Beckens von Rein in Steiermark. Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 13: 180 - 201. https: // www. biodiversitylibrary. org / page / 6360108" type="journal article" year="1854">Gobanz, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Miocene of the Styrian Basin (
|
||
<collectingCountry id="F30E7674BB28FFE1FEFBF8038988F812" box="[329,414,2001,2027]" name="Austria" pageId="9" pageNumber="650">Austria</collectingCountry>
|
||
). However, although we have not been able to locate specimens from Roussillon and the work by
|
||
<bibRefCitation id="EF884B15BB28FFE1FCE2FF258BEDFEE8" author="DEPERET C." box="[848,1019,247,273]" pageId="9" pageNumber="650" refId="ref10275" refString="DEPERET C. 1890. - Les animaux pliocenes du Roussillon. Societe geologique de France, Paleontologie, Memoire no. 3, 194 p." type="book" year="1890">Depéret (1890)</bibRefCitation>
|
||
does not contain associated figures, we think that
|
||
<taxonomicName id="4C194D67BB28FFE1FC2CFECA8C94FEC8" baseAuthorityName="Deperet" baseAuthorityYear="1890" box="[926,1154,279,306]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="variety" species="aquensis" variety="obtusa">
|
||
<emphasis id="B96DEAF6BB28FFE1FC2CFECA8C1AFEC8" box="[926,1036,280,305]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
var.
|
||
<emphasis id="B96DEAF6BB28FFE1FB8DFEC58C94FEC8" box="[1087,1154,279,305]" italics="true" pageId="9" pageNumber="650">obtusa</emphasis>
|
||
</taxonomicName>
|
||
is different from
|
||
<taxonomicName id="4C194D67BB28FFE1FA8EFECA8DB1FEC8" baseAuthorityName="Gobanz" baseAuthorityYear="1854" box="[1340,1447,280,305]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="porrecta">
|
||
<emphasis id="B96DEAF6BB28FFE1FA8EFECA8DB1FEC8" box="[1340,1447,280,305]" italics="true" pageId="9" pageNumber="650">P. porrecta</emphasis>
|
||
</taxonomicName>
|
||
, a species found only in Middle Miocene sites (
|
||
<bibRefCitation id="EF884B15BB28FFE1FA81FEE58BA4FE88" author="HARZHAUSER M. & NEUBAUER T. A. & GROSS M. & BINDER H." pageId="9" pageNumber="650" pagination="1 - 71" refId="ref11521" refString="HARZHAUSER M., NEUBAUER T. A., GROSS M. & BINDER H. 2014 b. - The early Middle Miocene mollusc fauna of Lake Rein (Eastern Alps, Austria). Palaeontographica, Abteilung A: Palaeozoology - Stratigraphy 302 (1 - 6): 1 - 71. https: // doi. org / 10.1127 / pala / 302 / 2013 / 1" type="journal article" year="2014">
|
||
Harzhauser
|
||
<emphasis id="B96DEAF6BB28FFE1FC9FFE8A8B75FE88" box="[813,867,343,369]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
2014b
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FC70FE858CD6FE88" author="HOLTKE O. & RASSER M. W." box="[962,1216,343,369]" pageId="9" pageNumber="650" pagination="267 - 302" refId="ref11740" refString="HOLTKE O. & RASSER M. W. 2017. - Land snails from the Miocene Steinheim impact crater lake sediments (Baden-Wurttemberg, South Germany). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 285 (3): 267 - 302. https: // doi. org / 10.1127 / njgpa / 2017 / 0681" type="journal article" year="2017">Höltke & Rasser 2017</bibRefCitation>
|
||
). Besides the age difference, according to the description and figure of
|
||
<bibRefCitation id="EF884B15BB28FFE1FAE8FEA58BBBFE49" author="GOBANZ J." pageId="9" pageNumber="650" pagination="180 - 201" refId="ref10979" refString="GOBANZ J. 1854. - Die fossilen Land- und Susswasser-Mollusken des Beckens von Rein in Steiermark. Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 13: 180 - 201. https: // www. biodiversitylibrary. org / page / 6360108" type="journal article" year="1854">Gobanz (1854: 196</bibRefCitation>
|
||
, fig. 5),
|
||
<taxonomicName id="4C194D67BB28FFE1FBA3FE458C96FE49" baseAuthorityName="Gobanz" baseAuthorityYear="1854" box="[1041,1152,407,432]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="porrecta">
|
||
<emphasis id="B96DEAF6BB28FFE1FBA3FE458C96FE49" box="[1041,1152,407,432]" italics="true" pageId="9" pageNumber="650">P. porrecta</emphasis>
|
||
</taxonomicName>
|
||
is characterised by a sharp ovate-elongate aperture that attains 70% of the total shell height according to the measurements indicated by
|
||
<bibRefCitation id="EF884B15BB28FFE1FAEBFE048B61FDE8" author="GOBANZ J." pageId="9" pageNumber="650" pagination="180 - 201" refId="ref10979" refString="GOBANZ J. 1854. - Die fossilen Land- und Susswasser-Mollusken des Beckens von Rein in Steiermark. Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 13: 180 - 201. https: // www. biodiversitylibrary. org / page / 6360108" type="journal article" year="1854">Gobanz (1854)</bibRefCitation>
|
||
, as well as an elongate, regularly convex last whorl. Note, however, that
|
||
<taxonomicName id="4C194D67BB28FFE1FBB2FDC58C7FFDC9" baseAuthorityName="Gobanz" baseAuthorityYear="1854" box="[1024,1129,535,560]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="porrecta">
|
||
<emphasis id="B96DEAF6BB28FFE1FBB2FDC58C7FFDC9" box="[1024,1129,535,560]" italics="true" pageId="9" pageNumber="650">P. porrecta</emphasis>
|
||
</taxonomicName>
|
||
is currently considered a taxon inquirendum, since the type series is lost, and the limited available material is inconclusive (
|
||
<bibRefCitation id="EF884B15BB28FFE1FB2EFD848D89FD89" author="HARZHAUSER M. & NEUBAUER T. A. & GROSS M. & BINDER H." box="[1180,1439,598,624]" pageId="9" pageNumber="650" pagination="1 - 71" refId="ref11521" refString="HARZHAUSER M., NEUBAUER T. A., GROSS M. & BINDER H. 2014 b. - The early Middle Miocene mollusc fauna of Lake Rein (Eastern Alps, Austria). Palaeontographica, Abteilung A: Palaeozoology - Stratigraphy 302 (1 - 6): 1 - 71. https: // doi. org / 10.1127 / pala / 302 / 2013 / 1" type="journal article" year="2014">
|
||
Harzhauser
|
||
<emphasis id="B96DEAF6BB28FFE1FAADFD858D44FD89" box="[1311,1362,598,624]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
2014b
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EF884B15BB28FFE1FC9FFDA48C2CFD69" author="DEPERET C. & SAYN G." box="[813,1082,630,656]" pageId="9" pageNumber="650" pagination="107 - 126" refId="ref10302" refString="DEPERET C. & SAYN G. 1900. - Monographie de la faune fluvioterrestre du Miocene du Cucuron (Vaucluse). Annales de la Societe linneenne de Lyon 47: 107 - 126. https: // www. biodiversitylibrary. org / page / 43976795" type="journal article" year="1900">Depéret & Sayn (1900)</bibRefCitation>
|
||
listed the
|
||
<taxonomicName id="4C194D67BB28FFE1FB06FDA48DB9FD69" authority="Ratavoux" authorityName="Ratavoux" box="[1204,1455,630,656]" pageId="9" pageNumber="650" rank="variety" variety="from">variety from Ratavoux</taxonomicName>
|
||
(Cucuron, Vaucluse,
|
||
<collectingCountry id="F30E7674BB28FFE1FBBCFD448C4CFD49" box="[1038,1114,662,688]" name="France" pageId="9" pageNumber="650">France</collectingCountry>
|
||
), dated, like Los Aljezares, to the Middle Turolian (MN12) (
|
||
<bibRefCitation id="EF884B15BB28FFE1FBE8FD678D38FD29" author="BALLESIO R. & CARBONNEL G. & MEIN P. & TRUC G." box="[1114,1326,693,720]" pageId="9" pageNumber="650" pagination="467 - 471" refId="ref9761" refString="BALLESIO R., CARBONNEL G., MEIN P. & TRUC G. 1979. - Sur un nouveau gisement fossilifere du Miocene superieur (Tortonien- Turolien moyen) de Cucuron (Vaucluse). Geobios 12 (3): 467 - 471. https: // doi. org / 10.1016 / s 0016 - 6995 (79) 80126 - 6" type="journal article" year="1979">
|
||
Ballesio
|
||
<emphasis id="B96DEAF6BB28FFE1FB05FD648CFBFD36" box="[1207,1261,693,719]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
1979
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FA8CFD648DB1FD29" author="ESU D." box="[1342,1447,694,720]" pageId="9" pageNumber="650" pagination="328 - 354" refId="ref10470" refString="ESU D. 1999. - Contribution to the knowledge of Neogene climatic changes in western and central Europa by means of non-marine molluscs, in AGUSTI J., ROOK L. & ANDREWS P. (eds), The Evolution of Neogene Terrestrial Ecosystems in Europa. Hominoid Evolution and Climatic Change in Europa. Vol. I. Cambridge University Press: 328 - 354. https: // doi. org / 10.1017 / cbo 9780511542329.016" type="book chapter" year="1999">Esu 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FC9FFD048BC3FD09" author="MENOURET B." box="[813,981,726,752]" pageId="9" pageNumber="650" pagination="56 - 74" refId="ref12041" refString="MENOURET B. 2014. - Gisements paleontologiques a mammiferes ou empreintes de pas de mammiferes du Parc naturel regional du Luberon. Courrier scientifique du Parc naturel regional du Luberon et de la Reserve de biosphere Luberon-Lure 12: 56 - 74." type="journal article" year="2014">Ménouret 2014</bibRefCitation>
|
||
). Our new species bears a strong resemblance to those from Ratavoux figured by
|
||
<bibRefCitation id="EF884B15BB28FFE1FB1FFD278DBEFCF6" author="DEPERET C. & SAYN G." box="[1197,1448,757,783]" pageId="9" pageNumber="650" pagination="107 - 126" refId="ref10302" refString="DEPERET C. & SAYN G. 1900. - Monographie de la faune fluvioterrestre du Miocene du Cucuron (Vaucluse). Annales de la Societe linneenne de Lyon 47: 107 - 126. https: // www. biodiversitylibrary. org / page / 43976795" type="journal article" year="1900">Depéret & Sayn (1900</bibRefCitation>
|
||
: pl. 1, fig. 77) and those deposited at CERESE (UCBL-FSL 148027,
|
||
<figureCitation id="13222A61BB28FFE1FC23FCE78BFEFCA9" box="[913,1000,821,848]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6D</figureCitation>
|
||
<figureCitation id="13222A61BB28FFE1FC5AFCE78C03FCA9" box="[1000,1045,821,848]" captionStart="FIG" captionStartId="4.[132,143,1045,1062]" captionTargetId="figure-532@4.[698,1265,211,742]" captionTargetPageId="4" captionText="FIG. 1. — Geographical and geological overview of the studied sites in the Teruel Basin: A, schematic geological map of the Teruel Basin region indicating the areas in which the studied sites are located (modified after Ezquerro et al. 2022a); B, C, detailed locations of the Los Aljezares (AL), La Gloria 4 (LG4) and Orrios 1 (OR1) sites. Underlying images © Google Maps 2023, CNES/Airbus, Maxar Technologies." figureDoi="http://doi.org/10.5281/zenodo.10201351" httpUri="https://zenodo.org/record/10201351/files/figure.png" pageId="9" pageNumber="650">1-D</figureCitation>
|
||
<figureCitation id="13222A61BB28FFE1FBA7FCE78C35FCB6" box="[1045,1059,821,847]" captionStart="FIG" captionStartId="5.[132,143,723,740]" captionTargetBox="[341,1251,215,683]" captionTargetId="figure-664@5.[336,977,212,685]" captionTargetPageId="5" captionText="FIG. 2. — A, Shell measurements,following Solem & Climo (1985), and arrangement of landmarks (large circles) and semi-landmarks (small dots). The numbers in parentheses indicate the number of points for each of the two semi-landmark curves. See text for abbreviations. B, Protoconch measurements, following Verduin (1977). Abbreviations: N, nucleus; P/T boundary, protoconch/teleoconch boundary. Scale bar: A, 5 mm; B, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10144334" httpUri="https://zenodo.org/record/10144334/files/figure.png" pageId="9" pageNumber="650">2</figureCitation>
|
||
), all having a characteristic, convex shape. However, the degree of preservation of the French material makes a precise comparison difficult.
|
||
<taxonomicName id="4C194D67BB28FFE1FAA2FCA78B99FC56" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
|
||
<emphasis id="B96DEAF6BB28FFE1FAA2FCA78B99FC56" italics="true" pageId="9" pageNumber="650">Palaeoglandina turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB28FFE1FC2AFC478BC0FC56" box="[920,982,917,943]" pageId="9" pageNumber="650" rank="species">n. sp.</taxonomicNameLabel>
|
||
has a characteristic ornamentation on the protoconch not observed in any of the other European species we investigated, i.e.
|
||
<taxonomicName id="4C194D67BB28FFE1FB95FC068DB9FC16" authority="(Zieten, 1832)" baseAuthorityName="Zieten" baseAuthorityYear="1832" box="[1063,1455,980,1007]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis id="B96DEAF6BB28FFE1FB95FC068D18FC17" box="[1063,1294,980,1006]" italics="true" pageId="9" pageNumber="650">Palaeoglandina gracilis</emphasis>
|
||
(Zieten, 1832)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13222A61BB28FFE1FC84FC268B9BFBF6" box="[822,909,1012,1039]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6H</figureCitation>
|
||
),
|
||
<taxonomicName id="4C194D67BB28FFE1FC10FC278CA7FBF6" authority="Binder, 2004" authorityName="Binder" authorityYear="2004" box="[930,1201,1012,1039]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="dactylina">
|
||
<emphasis id="B96DEAF6BB28FFE1FC10FC278C0DFBF7" box="[930,1051,1012,1038]" italics="true" pageId="9" pageNumber="650">P. dactylina</emphasis>
|
||
<bibRefCitation id="EF884B15BB28FFE1FB90FC268CA7FBF6" author="BINDER H." box="[1058,1201,1012,1039]" pageId="9" pageNumber="650" pagination="189 - 229" refId="ref9922" refString="BINDER H. 2004. - Terrestrial, freshwater and brachyhaline Gastropoda from Lower Miocene deposits of Oberdorf (Styria, Austria). Annalen des Naturhistorischen Museum in Wien 105 A: 189 - 229. https: // www. jstor. org / stable / 41702058" type="journal article" year="2004">Binder, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FB73FC268D5DFBF6" author="BINDER H." box="[1217,1355,1012,1039]" pageId="9" pageNumber="650" pagination="189 - 229" refId="ref9922" refString="BINDER H. 2004. - Terrestrial, freshwater and brachyhaline Gastropoda from Lower Miocene deposits of Oberdorf (Styria, Austria). Annalen des Naturhistorischen Museum in Wien 105 A: 189 - 229. https: // www. jstor. org / stable / 41702058" type="journal article" year="2004">Binder 2004</bibRefCitation>
|
||
,
|
||
<figureCitation id="13222A61BB28FFE1FAEBFC268DB1FBF7" box="[1369,1447,1012,1038]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6b</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C194D67BB28FFE1FCEFFBC78C91FBD7" authority="(Sacco, 1886)" baseAuthorityName="Sacco" baseAuthorityYear="1886" box="[861,1159,1044,1070]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="taurinensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FCEFFBC78BFEFBD7" box="[861,1000,1045,1070]" italics="true" pageId="9" pageNumber="650">P. taurinensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FC4AFBC68C96FBD7" author="SACCO F." box="[1016,1152,1044,1070]" pageId="9" pageNumber="650" pagination="427 - 476" refId="ref13172" refString="SACCO F. 1886. - Nuove species terziarie di molluschi terrestri e d'aqua dolce e salmastra del Piemonte. Atti della Societa Italiana di Scienze Naturali 29: 427 - 476. https: // www. biodiversitylibrary. org / page / 16237730" type="journal article" year="1886">Sacco, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13222A61BB28FFE1FB2BFBC68CF2FBD7" box="[1177,1252,1044,1070]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6I</figureCitation>
|
||
). This ornamentation could be a diagnostic character, but the absence of an apex in the specimens of
|
||
<taxonomicName id="4C194D67BB28FFE1FB8CFB878D33FB97" baseAuthorityName="Deperet" baseAuthorityYear="1890" box="[1086,1317,1108,1135]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="variety" species="aquensis" variety="obtusa">
|
||
<emphasis id="B96DEAF6BB28FFE1FB8CFB878CBBFB97" box="[1086,1197,1109,1134]" italics="true" pageId="9" pageNumber="650">P. aquensis</emphasis>
|
||
var.
|
||
<emphasis id="B96DEAF6BB28FFE1FB50FB868D33FB97" box="[1250,1317,1108,1134]" italics="true" pageId="9" pageNumber="650">obtusa</emphasis>
|
||
</taxonomicName>
|
||
deposited at CERESE prevents us from studying it and comparing it with that of the new species. The strong resemblance between the Ratavoux and Los Aljezares specimens, supported by similar age, suggest that they may be conspecific taxa, but the state of preservation of the French specimens does not allow a final conclusion. If both taxa turn out to be synonymous, the name
|
||
<taxonomicName id="4C194D67BB28FFE1FC24FAE18BE1FAB4" authorityName="Albesa & Neubauer" authorityYear="2023" box="[918,1015,1331,1357]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="turolensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FC24FAE18BE1FAB4" box="[918,1015,1331,1357]" italics="true" pageId="9" pageNumber="650">turolensis</emphasis>
|
||
</taxonomicName>
|
||
still has priority, since
|
||
<taxonomicName id="4C194D67BB28FFE1FB5EFAE18C29FA94" authority="Deperet, 1890" authorityName="Deperet" authorityYear="1890" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="variety" species="aquensis" variety="obtusa">
|
||
<emphasis id="B96DEAF6BB28FFE1FB5EFAE18DB9FAB4" box="[1260,1455,1331,1357]" italics="true" pageId="9" pageNumber="650">Glandina aquensis</emphasis>
|
||
var.
|
||
<emphasis id="B96DEAF6BB28FFE1FCEBFA818B8AFA94" box="[857,924,1363,1389]" italics="true" pageId="9" pageNumber="650">obtusa</emphasis>
|
||
<bibRefCitation id="EF884B15BB28FFE1FC10FA818C29FA94" author="DEPERET C." box="[930,1087,1363,1389]" pageId="9" pageNumber="650" refId="ref10275" refString="DEPERET C. 1890. - Les animaux pliocenes du Roussillon. Societe geologique de France, Paleontologie, Memoire no. 3, 194 p." type="book" year="1890">Depéret, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
is a junior homonym of
|
||
<taxonomicName id="4C194D67BB28FFE1FAFBFA818C18FA77" authority="Pfeiffer, 1845" authorityName="Pfeiffer" authorityYear="1845" class="Gastropoda" family="Oleacinidae" genus="Glandina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="obtusa">
|
||
<emphasis id="B96DEAF6BB28FFE1FAFBFA818B66FA74" italics="true" pageId="9" pageNumber="650">Glandina obtusa</emphasis>
|
||
Pfeiffer, 1845
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB28FFE1FCF1FA418C15F9D4" blockId="9.[811,1457,215,2028]" pageId="9" pageNumber="650">
|
||
Regarding
|
||
<taxonomicName id="4C194D67BB28FFE1FC76FA468C80FA54" baseAuthorityName="Matheron" baseAuthorityYear="1843" box="[964,1174,1427,1453]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="galloprovincialis">
|
||
<emphasis id="B96DEAF6BB28FFE1FC76FA468C80FA54" box="[964,1174,1427,1453]" italics="true" pageId="9" pageNumber="650">P. galloprovincialis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13222A61BB28FFE1FB18FA418D12FA54" box="[1194,1284,1427,1453]" captionStart="FIG" captionStartId="12.[132,143,1674,1691]" captionTargetBox="[132,1455,215,1632]" captionTargetId="figure-140@12.[132,1455,215,1632]" captionTargetPageId="12" captionText="FIG. 6. — A, Palaeoglandina aquensis (Matheron,1843) from Aix (Burdigalian,Aix Basin),MHNM 6034.9314.1;B, P. galloprovincialis (Matheron, 1843) from Peyrolles (Tortonian, Durance Basin), MHNM 6034.9317.1; C1, C2, P. montenati Truc, 1971 from Quibas (Calabrian, Quibas site), MVHN-180222GM05;D1, D2, P. aquensis var. obtusa (Depéret, 1890), UCBL-FSL 148027; E1, E2, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040; F1, F2, P. taurinensis (Sacco, 1886), BS.093.02.001; G, P. turolensis n. sp., MAP-8420, detail of protoconch; H, P. gracilis (Zieten, 1832), NHMW 2013/0572/0040, detail of protoconch; I, P. taurinensis (Sacco, 1886), BS.093.02.001,detail of protoconch;J, P.montenati Truc,1971 from Quibas,MVHN-180222GM05,detail of protoconch.Scale bars:A-F,10 mm;G,1 mm;H-J,2 mm." figureDoi="http://doi.org/10.5281/zenodo.10144342" httpUri="https://zenodo.org/record/10144342/files/figure.png" pageId="9" pageNumber="650">Fig. 6B</figureCitation>
|
||
), according to Matheron’s description and the measurements he provided (
|
||
<bibRefCitation id="EF884B15BB28FFE1FC84FA018C0BFA14" author="MATHERON P." box="[822,1053,1491,1517]" pageId="9" pageNumber="650" pagination="81 - 341" refId="ref12003" refString="MATHERON P. 1843. - Catalogue methodique et descriptif des corps organises fossiles du department des Bouches-du-Rhone et lieux circonvoisins. Repertoire des Travaux de la Societe de Statistique de Marseille 6: 81 - 341." type="journal article" year="1843">Matheron 1843: 279</bibRefCitation>
|
||
, pl. 34, fig. 10), the species has, with a much higher SH/SW ratio than the new species, a more slender appearance.
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB28FFE1FCF1F9E08C94F8B5" blockId="9.[811,1457,215,2028]" pageId="9" pageNumber="650">
|
||
Three species of the genus
|
||
<taxonomicName id="4C194D67BB28FFE1FBC4F9E08D0FF9B5" authorityName="Wenz" authorityYear="1914" box="[1142,1305,1586,1612]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96DEAF6BB28FFE1FBC4F9E08D0FF9B5" box="[1142,1305,1586,1612]" italics="true" pageId="9" pageNumber="650">Palaeoglandina</emphasis>
|
||
</taxonomicName>
|
||
are currently recognised in the European Pliocene:
|
||
<taxonomicName id="4C194D67BB28FFE1FB0BF9818B65F975" authority="(Michaud, 1877)" baseAuthorityName="Michaud" baseAuthorityYear="1877" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="paladilhei">
|
||
<emphasis id="B96DEAF6BB28FFE1FB0BF9818D20F995" box="[1209,1334,1618,1644]" italics="true" pageId="9" pageNumber="650">P. paladilhei</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B15BB28FFE1FAF7F9808B7CF975" author="MICHAUD G." pageId="9" pageNumber="650" refId="ref12085" refString="MICHAUD G. 1877. - Description des coquilles fossiles decouvertes dans les environs de Hauterives (Drome). Savy edit., Paris, 28 p." type="book" year="1877">Michaud, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, from the Early Pliocene (MN14) of Hauterives (
|
||
<collectingCountry id="F30E7674BB28FFE1FC86F9408B95F955" box="[820,899,1682,1708]" name="France" pageId="9" pageNumber="650">France</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C194D67BB28FFE1FC27F9418C16F955" baseAuthorityName="d'Ancona" baseAuthorityYear="1867" box="[917,1024,1682,1708]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="lunensis">
|
||
<emphasis id="B96DEAF6BB28FFE1FC27F9418C16F955" box="[917,1024,1682,1708]" italics="true" pageId="9" pageNumber="650">P. lunensis</emphasis>
|
||
</taxonomicName>
|
||
(D’Ancona in
|
||
<bibRefCitation id="EF884B15BB28FFE1FB17F9408D2BF955" author="COCCHI I." box="[1189,1341,1682,1708]" pageId="9" pageNumber="650" pagination="1 - 80" refId="ref10096" refString="COCCHI I. 1867. - L'uomo fossile nell'Italia centrale, studi paleontologici. Memorie della Societa Italiana di Scienze Naturali 2: 1 - 80." type="journal article" year="1867">Cocchi, 1867</bibRefCitation>
|
||
), from the Italian Late Pliocene to Early Pleistocene (MN16-MN17) (
|
||
<bibRefCitation id="EF884B15BB28FFE1FC84F9008B8AF915" author="ESU D." box="[822,924,1746,1772]" pageId="9" pageNumber="650" pagination="328 - 354" refId="ref10470" refString="ESU D. 1999. - Contribution to the knowledge of Neogene climatic changes in western and central Europa by means of non-marine molluscs, in AGUSTI J., ROOK L. & ANDREWS P. (eds), The Evolution of Neogene Terrestrial Ecosystems in Europa. Hominoid Evolution and Climatic Change in Europa. Vol. I. Cambridge University Press: 328 - 354. https: // doi. org / 10.1017 / cbo 9780511542329.016" type="book chapter" year="1999">Esu 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FC19F9008CA6F915" author="CIANGHEROTTI A. & ESU D. & MARTINETTO E. & GIUNTELLI P." box="[939,1200,1746,1772]" pageId="9" pageNumber="650" pagination="573 - 587" refId="ref10031" refString="CIANGHEROTTI A., ESU D., MARTINETTO E. & GIUNTELLI P. 2007. - The remarkable Middle Pliocene non-marine mollusc record from Ceresole d'Alba, Piedmont, north-west Italy: Biochronology, palaeobiogeography and palaeoecology supported by fossil plants. Geobios 40: 573 - 587. https: // doi. org / 10.1016 / j. geobios. 2006.10.005" type="journal article" year="2007">
|
||
Ciangherotti
|
||
<emphasis id="B96DEAF6BB28FFE1FB8EF9018C66F915" box="[1084,1136,1746,1772]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C194D67BB28FFE1FB5FF9018B75F8F5" authority="Truc, 1971" authorityName="Truc" authorityYear="1971" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="montenati">
|
||
<emphasis id="B96DEAF6BB28FFE1FB5FF9018D66F915" box="[1261,1392,1747,1772]" italics="true" pageId="9" pageNumber="650">P. montenati</emphasis>
|
||
<bibRefCitation id="EF884B15BB28FFE1FAC4F9008B75F8F5" author="TRUC G." pageId="9" pageNumber="650" pagination="79 - 129" refId="ref13525" refString="TRUC G. 1971. - Gasteropodes continentaux neogenes du Bassin rhodanien. Document du Laboratoire geologique de l'Universite de Lyon, hors serie 11: 79 - 129." type="journal article" year="1971">Truc, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Late Pliocene (MN16) and Early Pleistocene (MN17) of Alicante and Murcia (
|
||
<bibRefCitation id="EF884B15BB28FFE1FB11F8C08DBEF8D5" author="MONTENAT C. & TRUC G." box="[1187,1448,1810,1836]" pageId="9" pageNumber="650" pagination="52 - 58" refId="ref12335" refString="MONTENAT C. & TRUC G. 1971. - Pliocene superieur et Villafranchien dans le Levant espagnol (Provinces de Murcia et d'Alicante). Boletin Geologico y Minero 82 (1): 52 - 58." type="journal article" year="1971">Montenat & Truc 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF884B15BB28FFE1FC9FF8E08C00F8B5" author="MONTOYA P. & ALBERDI M. T. & BLAZQUEZ A. M. & BARBADILLO L. J. & FUMANAL P. & VAN DER MADE J. & MARIN J. M. & MOLINA A. & MORALES J. & MURELAGA X. & PENALVER E. & ROBLES F. & RUIZ BUSTOS A. & SANCHEZ A. & SANCHIZ B. & SORIA D. & SZYNDLAR Z." box="[813,1046,1841,1868]" pageId="9" pageNumber="650" pagination="127 - 161" refId="ref12374" refString="MONTOYA P., ALBERDI M. T., BLAZQUEZ A. M., BARBADILLO L. J., FUMANAL P., VAN DER MADE J., MARIN J. M., MOLINA A., MORALES J., MURELAGA X., PENALVER E., ROBLES F., RUIZ BUSTOS A., SANCHEZ A., SANCHIZ B., SORIA D. & SZYNDLAR Z. 1999 a. - La fauna del Pleistoceno inferior de la Sierra de Quibas (Abanilla, Murcia). Estudios Geologicos 55 (3 - 4): 127 - 161. https: // doi. org / 10.3989 / egeol. 99553 - 4171" type="journal article" year="1999">
|
||
Montoya
|
||
<emphasis id="B96DEAF6BB28FFE1FC25F8E08BDAF8B2" box="[919,972,1841,1867]" italics="true" pageId="9" pageNumber="650">et al.</emphasis>
|
||
1999a
|
||
</bibRefCitation>
|
||
,b, 2001).
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB28FFE2FCF1F8838965F812" blockId="9.[811,1457,215,2028]" lastBlockId="10.[131,777,1650,2027]" lastPageId="10" lastPageNumber="651" pageId="9" pageNumber="650">
|
||
<taxonomicName id="4C194D67BB28FFE1FCF1F8838C5AF892" ID-CoL="7SL33" baseAuthorityName="Michaud" baseAuthorityYear="1877" box="[835,1100,1873,1899]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="650" phylum="Mollusca" rank="species" species="paladilhei">
|
||
<emphasis id="B96DEAF6BB28FFE1FCF1F8838C5AF892" box="[835,1100,1873,1899]" italics="true" pageId="9" pageNumber="650">Palaeoglandina paladilhei</emphasis>
|
||
</taxonomicName>
|
||
is a problematic species described from incomplete fragments of several specimens “three fragments, two of which show the last whorl and leave the opening fairly complete and the third comprising the apex” (
|
||
<bibRefCitation id="EF884B15BB28FFE1FC84F8038BEAF812" author="MICHAUD G." box="[822,1020,2001,2028]" pageId="9" pageNumber="650" refId="ref12085" refString="MICHAUD G. 1877. - Description des coquilles fossiles decouvertes dans les environs de Hauterives (Drome). Savy edit., Paris, 28 p." type="book" year="1877">Michaud 1877: 7</bibRefCitation>
|
||
), whose reconstruction was considered “unsatisfactory” by
|
||
<bibRefCitation id="EF884B15BB2BFFE2FEE7F9A18A83F974" author="MONTENAT C. & TRUC G." box="[341,661,1650,1677]" pageId="10" pageNumber="651" pagination="52 - 58" refId="ref12335" refString="MONTENAT C. & TRUC G. 1971. - Pliocene superieur et Villafranchien dans le Levant espagnol (Provinces de Murcia et d'Alicante). Boletin Geologico y Minero 82 (1): 52 - 58." type="journal article" year="1971">Montenat & Truc (1971: 57)</bibRefCitation>
|
||
. Although the reconstruction is indeed problematic and, in our opinion, shows one whorl too many, the fragments of the apical part of the shell and of the last whorl with the opening are drawn in great detail and are easily distinguishable from the inferred intermediate part. According to this information and the original description,
|
||
<taxonomicName id="4C194D67BB2BFFE2FDB1F8E08A9EF8B2" ID-CoL="7SL33" baseAuthorityName="Michaud" baseAuthorityYear="1877" box="[515,648,1841,1867]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="paladilhei">
|
||
<emphasis id="B96DEAF6BB2BFFE2FDB1F8E08A9EF8B2" box="[515,648,1841,1867]" italics="true" pageId="10" pageNumber="651">P. paladilhei</emphasis>
|
||
</taxonomicName>
|
||
is very different from
|
||
<taxonomicName id="4C194D67BB2BFFE2FEBCF880899DF892" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[270,395,1873,1899]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2BFFE2FEBCF880899DF892" box="[270,395,1873,1899]" italics="true" pageId="10" pageNumber="651">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2BFFE2FE20F88089C4F895" box="[402,466,1874,1900]" pageId="10" pageNumber="651" rank="species">n. sp.</taxonomicNameLabel>
|
||
, easily distinguished by the shape of the first whorls, flat in the Hauterives species and rounded in the new species, as well as by the shape of the last whorl and the aperture, narrower and more elongated in the French species.
|
||
</paragraph>
|
||
<caption id="DF66666CBB2BFFE2FF36FA238D52F9D6" ID-DOI="http://doi.org/10.5281/zenodo.10144340" ID-Zenodo-Dep="10144340" httpUri="https://zenodo.org/record/10144340/files/figure.png" pageId="10" pageNumber="651" startId="10.[132,143,1521,1538]" targetBox="[132,1455,215,1477]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BA636E4BB2BFFE2FF36FA238D52F9D6" blockId="10.[132,1457,1521,1583]" pageId="10" pageNumber="651">
|
||
<smallCapsWord id="8D40A038BB2BFFE2FF36FA23888BF9F8" baselines="1534,1534" box="[132,157,1521,1538]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="651">FIG</smallCapsWord>
|
||
. 5. —
|
||
<emphasis id="B96DEAF6BB2BFFE2FF6AFA2388ECF9FB" bold="true" box="[216,250,1521,1538]" pageId="10" pageNumber="651">A -D</emphasis>
|
||
,
|
||
<emphasis id="B96DEAF6BB2BFFE2FEB5FA238900F9FB" bold="true" box="[263,278,1521,1538]" pageId="10" pageNumber="651">O</emphasis>
|
||
,
|
||
<taxonomicName id="4C194D67BB2BFFE2FE90FA2389EAF9FB" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[290,508,1521,1538]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="turolensis" status="sp. nov.">
|
||
<emphasis id="B96DEAF6BB2BFFE2FE90FA2389EAF9FB" box="[290,508,1521,1538]" italics="true" pageId="10" pageNumber="651">Palaeoglandina turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2BFFE2FDB1FA238A23F9FB" box="[515,565,1521,1538]" pageId="10" pageNumber="651" rank="species">n. sp.</taxonomicNameLabel>
|
||
from Orrios 1 (Ruscinian, Teruel Basin):
|
||
<emphasis id="B96DEAF6BB2BFFE2FC25FA238BB2F9FB" bold="true" box="[919,932,1521,1538]" pageId="10" pageNumber="651">A</emphasis>
|
||
,
|
||
<emphasis id="B96DEAF6BB2BFFE2FC02FA238BA9F9FB" bold="true" box="[944,959,1521,1538]" pageId="10" pageNumber="651">O</emphasis>
|
||
, MAP-8421;
|
||
<emphasis id="B96DEAF6BB2BFFE2FB80FA238C29F9FB" bold="true" box="[1074,1087,1521,1538]" pageId="10" pageNumber="651">B</emphasis>
|
||
, MAP-8422;
|
||
<emphasis id="B96DEAF6BB2BFFE2FB01FA238CD7F9FB" bold="true" box="[1203,1217,1521,1538]" pageId="10" pageNumber="651">C</emphasis>
|
||
, MAP-8423;
|
||
<emphasis id="B96DEAF6BB2BFFE2FA87FA238D55F9FB" bold="true" box="[1333,1347,1521,1538]" pageId="10" pageNumber="651">D</emphasis>
|
||
, MAP-8424;
|
||
<emphasis id="B96DEAF6BB2BFFE2FF36F9D588B5F9E1" bold="true" box="[132,163,1543,1560]" pageId="10" pageNumber="651">E -L</emphasis>
|
||
,
|
||
<emphasis id="B96DEAF6BB2BFFE2FF1EF9D588ACF9E1" bold="true" box="[172,186,1543,1560]" pageId="10" pageNumber="651">N</emphasis>
|
||
,
|
||
<taxonomicName id="4C194D67BB2BFFE2FF71F9D5893EF9E1" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[195,296,1543,1560]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2BFFE2FF71F9D5893EF9E1" box="[195,296,1543,1560]" italics="true" pageId="10" pageNumber="651">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2BFFE2FE9EF9D5894DF9E1" box="[300,347,1543,1560]" pageId="10" pageNumber="651" rank="species">n. sp.</taxonomicNameLabel>
|
||
from Los Aljezares (Turolian,Teruel Basin):
|
||
<emphasis id="B96DEAF6BB2BFFE2FD0CF9D58ADCF9E1" bold="true" box="[702,714,1543,1560]" pageId="10" pageNumber="651">E</emphasis>
|
||
, 040822JA01;
|
||
<emphasis id="B96DEAF6BB2BFFE2FCF7F9D58B46F9E1" bold="true" box="[837,848,1543,1560]" pageId="10" pageNumber="651">F</emphasis>
|
||
,
|
||
<emphasis id="B96DEAF6BB2BFFE2FCEBF9D58B71F9E1" bold="true" box="[857,871,1543,1560]" pageId="10" pageNumber="651">N</emphasis>
|
||
, 040822JA02;
|
||
<emphasis id="B96DEAF6BB2BFFE2FC50F9D58BE6F9E1" bold="true" box="[994,1008,1543,1560]" pageId="10" pageNumber="651">G</emphasis>
|
||
, MGM-3723;
|
||
<emphasis id="B96DEAF6BB2BFFE2FBD0F9D58C66F9E1" bold="true" box="[1122,1136,1543,1560]" pageId="10" pageNumber="651">H</emphasis>
|
||
, MNCNI-28900-D;
|
||
<emphasis id="B96DEAF6BB2BFFE2FABDF9D58D03F9E1" bold="true" box="[1295,1301,1543,1560]" pageId="10" pageNumber="651">I</emphasis>
|
||
, MNCNI-28900-A;
|
||
<emphasis id="B96DEAF6BB2BFFE2FF36F9CC8898F9D6" bold="true" box="[132,142,1566,1583]" pageId="10" pageNumber="651">J</emphasis>
|
||
, MGM-118M;
|
||
<emphasis id="B96DEAF6BB2BFFE2FEBBF9CC8900F9D6" bold="true" box="[265,278,1566,1583]" pageId="10" pageNumber="651">K</emphasis>
|
||
, MNCNI-28900-E;
|
||
<emphasis id="B96DEAF6BB2BFFE2FE04F9CC89D7F9D6" bold="true" box="[438,449,1566,1583]" pageId="10" pageNumber="651">L</emphasis>
|
||
, MNCNI-11300-E;
|
||
<emphasis id="B96DEAF6BB2BFFE2FDD0F9CC8A65F9D6" bold="true" box="[610,627,1566,1583]" pageId="10" pageNumber="651">M</emphasis>
|
||
,
|
||
<taxonomicName id="4C194D67BB2BFFE2FDCFF9CC8AF5F9D6" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[637,739,1566,1583]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2BFFE2FDCFF9CC8AF5F9D6" box="[637,739,1566,1583]" italics="true" pageId="10" pageNumber="651">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2BFFE2FD5AF9CC8B0EF9D6" box="[744,792,1566,1583]" pageId="10" pageNumber="651" rank="species">n. sp.</taxonomicNameLabel>
|
||
from La Gloria 4, MAP-8405. Scale bars: A-J, 10 mm; M-O, 6 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636E4BB2BFFE2FCF1F9A08D07F8B2" blockId="10.[810,1457,1650,2028]" pageId="10" pageNumber="651">
|
||
<taxonomicName id="4C194D67BB2BFFE2FCF1F9A08C3AF975" ID-CoL="4C2FM" baseAuthorityName="d'Ancona" baseAuthorityYear="1867" box="[835,1068,1650,1676]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="lunensis">
|
||
<emphasis id="B96DEAF6BB2BFFE2FCF1F9A08C3AF975" box="[835,1068,1650,1676]" italics="true" pageId="10" pageNumber="651">Palaeoglandina lunensis</emphasis>
|
||
</taxonomicName>
|
||
, which has been figured and described in detail by
|
||
<bibRefCitation id="EF884B15BB2BFFE2FC19F9418C7BF955" author="DE STEFANI C." box="[939,1133,1682,1709]" pageId="10" pageNumber="651" pagination="9 - 108" refId="ref13469" refString="DE STEFANI C. 1880. - Molluschi continentali fino ad ora notati in ltalia nei terreni pliocenici, ed ordinamento di questi ultimi (continuazione). Atti della Societa Toscana di Scienze Naturali Residente in Pisa 5: 9 - 108. https: // www. biodiversitylibrary. org / page / 36825211" type="journal article" year="1880">De Stefani (1880)</bibRefCitation>
|
||
, has a regularly fusiform shell with low-convex whorls and, with one more whorl, a slower growth. Moreover, the Italian species appears to have a very marked reticulate ornamentation of teleoconch (
|
||
<bibRefCitation id="EF884B15BB2BFFE2FA8FF9208B72F8D5" author="DE STEFANI C." pageId="10" pageNumber="651" pagination="9 - 108" refId="ref13469" refString="DE STEFANI C. 1880. - Molluschi continentali fino ad ora notati in ltalia nei terreni pliocenici, ed ordinamento di questi ultimi (continuazione). Atti della Societa Toscana di Scienze Naturali Residente in Pisa 5: 9 - 108. https: // www. biodiversitylibrary. org / page / 36825211" type="journal article" year="1880">De Stefani 1880</bibRefCitation>
|
||
: pl. 2, fig. 13), which is not seen in
|
||
<taxonomicName id="4C194D67BB2BFFE2FB41F8C08D7BF8D5" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[1267,1389,1810,1836]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2BFFE2FB41F8C08D7BF8D5" box="[1267,1389,1810,1836]" italics="true" pageId="10" pageNumber="651">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2BFFE2FAC6F8C08DB9F8D5" box="[1396,1455,1810,1836]" pageId="10" pageNumber="651" rank="species">n. sp.</taxonomicNameLabel>
|
||
where the reticulation observed is very faint.
|
||
</paragraph>
|
||
<paragraph id="8BA636E4BB2BFFE3FCF1F88389D4F812" blockId="10.[810,1457,1650,2028]" lastBlockId="11.[131,777,1554,2027]" lastPageId="11" lastPageNumber="652" pageId="10" pageNumber="651">
|
||
<taxonomicName id="4C194D67BB2BFFE2FCF1F8838C58F892" ID-CoL="7SL32" authorityName="Truc" authorityYear="1971" box="[835,1102,1873,1899]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="651" phylum="Mollusca" rank="species" species="montenati">
|
||
<emphasis id="B96DEAF6BB2BFFE2FCF1F8838C58F892" box="[835,1102,1873,1899]" italics="true" pageId="10" pageNumber="651">Palaeoglandina montenati</emphasis>
|
||
</taxonomicName>
|
||
, a large species with a total height of
|
||
<emphasis id="B96DEAF6BB2BFFE2FCFEF8A08B4BF872" box="[844,861,1906,1931]" italics="true" pageId="10" pageNumber="651">c.</emphasis>
|
||
<quantity id="4CE19B01BB2BFFE2FCD4F8A38BCBF875" box="[870,989,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="10" pageNumber="651" unit="mm" value="65.0" valueMax="70.0" valueMin="60.0">60-70 mm</quantity>
|
||
(
|
||
<typeStatus id="54A28846BB2BFFE2FC5CF8A38C45F872" box="[1006,1107,1905,1931]" pageId="10" pageNumber="651" type="holotype">holotype</typeStatus>
|
||
, UCBL-FSL 98969:
|
||
<quantity id="4CE19B01BB2BFFE2FAF3F8A38DBDF875" box="[1345,1451,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="10" pageNumber="651" unit="mm" value="59.5">59.5 mm</quantity>
|
||
;
|
||
<typeStatus id="54A28846BB2BFFE2FC9CF8408B87F855" box="[814,913,1938,1964]" pageId="10" pageNumber="651" type="paratype">paratype</typeStatus>
|
||
, UCBL-FSL 98971:
|
||
<quantity id="4CE19B01BB2BFFE2FB30F8438CCEF855" box="[1154,1240,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="651" unit="mm" value="69.0">69 mm</quantity>
|
||
,
|
||
<bibRefCitation id="EF884B15BB2BFFE2FB54F8408B85F835" author="MONTENAT C. & TRUC G." pageId="10" pageNumber="651" pagination="52 - 58" refId="ref12335" refString="MONTENAT C. & TRUC G. 1971. - Pliocene superieur et Villafranchien dans le Levant espagnol (Provinces de Murcia et d'Alicante). Boletin Geologico y Minero 82 (1): 52 - 58." type="journal article" year="1971">Montenat & Truc 1971: 57</bibRefCitation>
|
||
), has a more fusiform morphology (
|
||
<bibRefCitation id="EF884B15BB2BFFE2FA91F8608BB6F812" author="MONTENAT C. & TRUC G." pageId="10" pageNumber="651" pagination="52 - 58" refId="ref12335" refString="MONTENAT C. & TRUC G. 1971. - Pliocene superieur et Villafranchien dans le Levant espagnol (Provinces de Murcia et d'Alicante). Boletin Geologico y Minero 82 (1): 52 - 58." type="journal article" year="1971">Montenat & Truc 1971</bibRefCitation>
|
||
: fig. 4a-c), with a SH/SW ratio of more than 2, a spire that attains about 10% of the total shell height and thus is proportionally shorter than that of the new species and a AH/SH ratio greater than in
|
||
<taxonomicName id="4C194D67BB2AFFE3FDF7F9818AD4F995" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[581,706,1618,1644]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="652" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2AFFE3FDF7F9818AD4F995" box="[581,706,1618,1644]" italics="true" pageId="11" pageNumber="652">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2AFFE3FD78F9818B11F994" box="[714,775,1619,1645]" pageId="11" pageNumber="652" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<tableCitation id="C69B035FBB2AFFE3FF3CF9A088CAF975" box="[142,220,1650,1676]" captionStart="TABLE" captionStartId="11.[133,142,219,236]" captionTargetPageId="11" captionText="TABLE 1. — Shell measurements of several of the studied specimens.Data for species other than P. turolensis n. sp. derive from Matheron (1843) and Montenat & Truc (1971)." httpUri="http://table.plazi.org/id/DF66666CBB2AFFE3FF37FF0988F1FEFA" pageId="11" pageNumber="652" tableUuid="DF66666CBB2AFFE3FF37FF0988F1FEFA">Table 1</tableCitation>
|
||
). The type specimens are moulds, but the specimens from Quibas are well preserved, showing a proportionally shorter spire with a flatter whorl profile, as well as a last whorl that is also proportionally wider and convex abapically. All this gives the shell a more fusiform morphology than the specimens from the Teruel Basin. On the other hand, the ornamentation of the protoconch is similar to that of
|
||
<taxonomicName id="4C194D67BB2AFFE3FF61F8808958F892" authority="Albesa & Neubauer, 2023" authorityName="Albesa & Neubauer" authorityYear="2023" box="[211,334,1873,1899]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="652" phylum="Mollusca" rank="species" species="turolensis" status="n. sp.">
|
||
<emphasis id="B96DEAF6BB2AFFE3FF61F8808958F892" box="[211,334,1873,1899]" italics="true" pageId="11" pageNumber="652">P. turolensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E578DBB2AFFE3FEE7F8808985F895" box="[341,403,1874,1900]" pageId="11" pageNumber="652" rank="species">n. sp.</taxonomicNameLabel>
|
||
, a circumstance that may indicate the possibility that they belong to the same evolutionary lineage. However, confirming this interpretation will require a more detailed assessment of well preserved specimens of other
|
||
<taxonomicName id="4C194D67BB2AFFE3FF74F80389D4F812" box="[198,450,2001,2027]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="652" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B96DEAF6BB2AFFE3FF74F803897EF812" box="[198,360,2001,2027]" italics="true" pageId="11" pageNumber="652">Palaeoglandina</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |