treatments-xml/data/03/B0/87/03B087E55B08FFFEFF79F2BAFE1FF991.xml
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<document id="E9F7F2AAE3072D19C0B9709FE85DC13C" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B08FFFEFF79F2BAFE1FF991" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena pacificica Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="110" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="110" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A0CC2ED2CF3A48D8FC133496FD96EBA9">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="A37B308F6CC676338F4A87442F134483">Perkins, Philip D.</mods:namePart>
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<treatment id="03B087E55B08FFFEFF79F2BAFE1FF991" ID-DOI="http://doi.org/10.5281/zenodo.5291728" ID-Zenodo-Dep="5291728" LSID="urn:lsid:plazi:treatment:03B087E55B08FFFEFF79F2BAFE1FF991" httpUri="http://treatment.plazi.org/id/03B087E55B08FFFEFF79F2BAFE1FF991" lastPageNumber="110" pageId="109" pageNumber="110">
<subSubSection id="C30365785B08FFFEFF79F2BAFD9CFF27" box="[151,554,151,177]" pageId="109" pageNumber="110" type="nomenclature">
<paragraph id="8BA636F35B08FFFEFF79F2BAFD9CFF27" blockId="109.[151,554,151,210]" box="[151,554,151,177]" pageId="109" pageNumber="110">
<heading id="D0EE819F5B08FFFEFF79F2BAFD9CFF27" bold="true" box="[151,554,151,177]" fontSize="11" level="1" pageId="109" pageNumber="110" reason="1">
<emphasis id="B96DEAE15B08FFFEFF79F2BAFD9CFF27" bold="true" box="[151,554,151,177]" pageId="109" pageNumber="110">
<taxonomicName id="4C194D705B08FFFEFF79F2BAFE3DFF27" ID-CoL="6MJC4" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,395,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="pacificica" status="sp. nov.">
<emphasis id="B96DEAE15B08FFFEFF79F2BAFE3DFF27" bold="true" box="[151,395,151,177]" italics="true" pageId="109" pageNumber="110">Hydraena pacificica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B08FFFEFE74F2BAFD9CFF27" box="[410,554,151,177]" pageId="109" pageNumber="110" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF79F297FE34FF44" box="[151,386,186,210]" pageId="109" pageNumber="110" type="description">
<paragraph id="8BA636F35B08FFFEFF79F297FE34FF44" blockId="109.[151,554,151,210]" box="[151,386,186,210]" pageId="109" pageNumber="110">
(
<figureCitation id="13222A765B08FFFEFF71F297FEBDFF44" box="[159,267,186,210]" captionStart="FIGURE 194" captionStartId="290.[151,250,1753,1776]" captionTargetBox="[220,1371,215,1718]" captionTargetId="figure-28@290.[198,1388,193,1732]" captionTargetPageId="290" captionText="FIGURE 194. Hydraena pacificica, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283786" httpUri="https://zenodo.org/record/5283786/files/figure.png" pageId="109" pageNumber="110">Figs. 194</figureCitation>
,
<figureCitation id="13222A765B08FFFEFEF8F297FEF4FF44" box="[278,322,186,210]" captionStart="FIGURES 196197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="109" pageNumber="110">196</figureCitation>
,
<figureCitation id="13222A765B08FFFEFEA0F297FECFFF44" box="[334,377,186,210]" captionStart="FIGURES 519522" captionStartId="405.[151,264,1650,1673]" captionTargetBox="[165,1421,253,1615]" captionTargetId="figure-20@405.[165,1421,193,1629]" captionTargetPageId="405" captionText="FIGURES 519522. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284131" httpUri="https://zenodo.org/record/5284131/files/figure.png" pageId="109" pageNumber="110">521</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF79F32EFD4FFEA8" pageId="109" pageNumber="110" type="materials_examined">
<paragraph id="8BA636F35B08FFFEFF79F32EFD4FFEA8" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
Type Material.
<materialsCitation id="3B713CAE5B08FFFEFEBDF32EFD4FFEA8" collectingDate="2006-03-31" collectionCode="ZSM" collectorName="Balke &amp; Sagata" country="Papua New Guinea" elevation="850" latitude="-6.5818834" location="Huon Peninsula" municipality="Kwapsanek" pageId="109" pageNumber="110" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
<typeStatus id="54A288515B08FFFEFEBDF32EFE76FE8D" box="[339,448,259,283]" pageId="109" pageNumber="110" type="holotype">
<emphasis id="B96DEAE15B08FFFEFEBDF32EFE76FE8D" bold="true" box="[339,448,259,283]" pageId="109" pageNumber="110">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B08FFFEFDC2F32EFCB0FE8D" bold="true" box="[556,774,258,283]" pageId="109" pageNumber="110">
<collectingRegion id="49DDF8115B08FFFEFDC2F32EFCB7FE8D" box="[556,769,258,283]" country="Papua New Guinea" name="Morobe" pageId="109" pageNumber="110">Morobe Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B08FFFEFCF7F32EFC66FE8C" LSID="urn:lsid:plazi:treatment:03B087E55B08FFFEFF79F2BAFE1FF991:8EC660285B08FFFEFCF7F32EFC66FE8C" box="[793,976,258,282]" country="Papua New Guinea" latitude="-6.5818834" municipality="Kwapsanek" name="Huon Peninsula" pageId="109" pageNumber="110" stateProvince="Morobe">Huon Peninsula</location>
,
<collectingMunicipality id="6BC2AC895B08FFFEFC33F32FFBD5FE8C" box="[989,1123,258,282]" pageId="109" pageNumber="110">Kwapsanek</collectingMunicipality>
,
<quantity id="4CE19B165B08FFFEFB9EF32FFB0DFE8C" box="[1136,1211,258,282]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="109" pageNumber="110" unit="m" value="850.0">
<elevation id="0034D1C05B08FFFEFB9EF32FFB0DFE8C" box="[1136,1211,258,282]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="109" pageNumber="110" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B08FFFEFB27F32FFAEEFE8C" box="[1225,1368,258,282]" degrees="6" direction="south" minutes="34.913" orientation="latitude" pageId="109" pageNumber="110" precision="1" value="-6.5818834">6° 34.913' S</geoCoordinate>
, 147°.526' E,
<date id="FFA710335B08FFFEFF1DF30BFED9FEA8" box="[243,367,294,318]" pageId="109" pageNumber="110" value="2006-03-31">
<collectingDate id="EFE3E9DB5B08FFFEFF1DF30BFED9FEA8" box="[243,367,294,318]" pageId="109" pageNumber="110" value="2006-03-31">31 iii 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B08FFFEFE94F30BFE0DFEA8" box="[378,443,294,318]" pageId="109" pageNumber="110">Balke</collectorName>
&amp;
<collectorName id="26EC53255B08FFFEFE31F30BFD9FFEA8" box="[479,553,294,318]" pageId="109" pageNumber="110">Sagata</collectorName>
(
<collectingCountry id="F30E76635B08FFFEFDD6F30BFDC4FEA8" box="[568,626,294,318]" name="Papua New Guinea" pageId="109" pageNumber="110">PNG</collectingCountry>
25) (
<collectionCode id="ED08AE365B08FFFEFD40F30BFD5BFEA8" box="[686,749,294,318]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="109" pageNumber="110" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF29F367FCDEFDAC" pageId="109" pageNumber="110" type="diagnosis">
<paragraph id="8BA636F35B08FFFEFF29F367FCDEFDAC" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
<emphasis id="B96DEAE15B08FFFEFF29F367FE60FEF5" bold="true" box="[199,470,330,355]" pageId="109" pageNumber="110">Differential Diagnosis.</emphasis>
Similar in body proportions and dorsal sculpture to
<taxonomicName id="4C194D705B08FFFEFBABF366FB78FEF4" authorityName="Perkins" authorityYear="2011" box="[1093,1230,331,354]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="apertista">
<emphasis id="B96DEAE15B08FFFEFBABF366FBEFFEF4" box="[1093,1113,331,354]" italics="true" pageId="109" pageNumber="110">H</emphasis>
.
<emphasis id="B96DEAE15B08FFFEFB85F366FB78FEF4" box="[1131,1230,331,354]" italics="true" pageId="109" pageNumber="110">apertista</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B08FFFEFB0CF367FAE7FEF4" box="[1250,1361,330,354]" captionStart="FIGURE 191" captionStartId="288.[151,250,1753,1776]" captionTargetBox="[232,1354,193,1732]" captionTargetId="figure-29@288.[232,1354,193,1732]" captionTargetPageId="288" captionText="FIGURE 191. Hydraena apertista, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283778" httpUri="https://zenodo.org/record/5283778/files/figure.png" pageId="109" pageNumber="110">Figs. 191</figureCitation>
,
<figureCitation id="13222A765B08FFFEFA8FF367FA39FEF4" box="[1377,1423,330,354]" captionStart="FIGURE 194" captionStartId="290.[151,250,1753,1776]" captionTargetBox="[220,1371,215,1718]" captionTargetId="figure-28@290.[198,1388,193,1732]" captionTargetPageId="290" captionText="FIGURE 194. Hydraena pacificica, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283786" httpUri="https://zenodo.org/record/5283786/files/figure.png" pageId="109" pageNumber="110">194</figureCitation>
). These two species have the mesoventral intercoxal process (P2) very short (length slightly less than the apical width), compared with the other species in the
<emphasis id="B96DEAE15B08FFFEFD44F3BEFD4FFE3D" box="[682,761,403,427]" italics="true" pageId="109" pageNumber="110">Impala</emphasis>
group.
<taxonomicName id="4C194D705B08FFFEFCBFF3BEFB81FE3D" authorityName="Perkins" authorityYear="2011" box="[849,1079,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="pacificica">
<emphasis id="B96DEAE15B08FFFEFCBFF3BEFB81FE3D" box="[849,1079,403,427]" italics="true" pageId="109" pageNumber="110">Hydraena pacificica</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C194D705B08FFFEFB20F3BEFAE3FE3C" authorityName="Perkins" authorityYear="2011" box="[1230,1365,403,426]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="apertista">
<emphasis id="B96DEAE15B08FFFEFB20F3BEFB54FE3C" box="[1230,1250,403,426]" italics="true" pageId="109" pageNumber="110">H</emphasis>
.
<emphasis id="B96DEAE15B08FFFEFB1FF3BEFAE3FE3C" box="[1265,1365,403,426]" italics="true" pageId="109" pageNumber="110">apertista</emphasis>
</taxonomicName>
in the slightly more strongly emarginate anterior margin of the pronotum, the different plaque ratios (ca. 4/1/4/5 vs. 6/1/5/ 7), and in general a less uniform dorsal coloration, with an ill-defined pronotal macula. The general plan of the male genitalia of the two species also suggests a relationship, but the genitalia distinctively differ in details, and should be examined for reliable determinations (
<figureCitation id="13222A765B08FFFEFD58F00FFC97FDAC" box="[694,801,546,570]" captionStart="FIGURES 192193" captionStartId="289.[151,264,1650,1673]" captionTargetBox="[305,1295,206,1619]" captionTargetId="figure-29@289.[264,1323,193,1629]" captionTargetPageId="289" captionText="FIGURES 192193. Aedeagi of holotypes. 192. H. exhalista. 193. H. apertista." figureDoi="http://doi.org/10.5281/zenodo.5283780" httpUri="https://zenodo.org/record/5283780/files/figure.png" pageId="109" pageNumber="110">Figs. 193</figureCitation>
,
<figureCitation id="13222A765B08FFFEFCC3F00FFCEDFDAC" box="[813,859,546,570]" captionStart="FIGURES 196197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="109" pageNumber="110">196</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF29F06BFB1DFAE0" pageId="109" pageNumber="110" type="description">
<paragraph id="8BA636F35B08FFFEFF29F06BFCBDFD14" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
<emphasis id="B96DEAE15B08FFFEFF29F06BFEEEFDC9" bold="true" box="[199,344,582,607]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B08FFFEFE42F06BFDB8FDC8" box="[428,526,582,606]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.31/0.58; head 0.22/0.30; pronotum 0.31/0.44, PA 0.34, PB 0.36; elytra 0.77/0.58.
</paragraph>
<paragraph id="8BA636F35B08FFFEFF79F0A2FD0BFD78" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">Dorsum of head with frons dark brown to dark reddish brown, clypeus lighter, brown, labrum testaceous; pronotum brown around ill-defined dark reddish brown macula on disc; elytra dark brown to dark reddish brown; legs reddish brown; maxillary palpi testaceous, tip not darker.</paragraph>
<paragraph id="8BA636F35B08FFFEFF29F0D6FEFDFC50" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">Frons punctures ca. 12xef, slightly larger near eyes than medially; interstices shining, 13xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 24xpd, punctures larger and denser at anterior and posterior; PF1 and PF2 very shallow, obsolete; PF3 moderately deep; PF4 absent or very shallow.</paragraph>
<paragraph id="8BA636F35B08FFFEFF29F1FFFF47FBEC" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins rather wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 23xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B08FFFEFF29F6ABFB1DFAE0" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 4/1/4/5. P1 ca. 1/2 P2; median carina bisinuate in profile. P2 very short, l/w ca. 3/4, sides slightly converging toward blunt apex. Plaques very narrow, carinate, converging very slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. 1.7x P2. All legs moderately long and slender. Profemur (male) with very small tubercle next to trochanter; protibia slightly arcuate, anterior margin very slightly widened subapically, medial margin flat except small sharp tubercle subapically on posteromedial margin. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) very deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B08FFFEFBD2F773FB29FAE0" box="[1084,1183,1374,1398]" captionStart="FIGURES 196197" captionStartId="292.[151,264,1650,1673]" captionTargetBox="[310,1275,212,1616]" captionTargetId="figure-32@292.[264,1323,193,1629]" captionTargetPageId="292" captionText="FIGURES 196197. Aedeagi of holotypes. 196. H. pacificica. 197. H. confluenta." figureDoi="http://doi.org/10.5281/zenodo.5283792" httpUri="https://zenodo.org/record/5283792/files/figure.png" pageId="109" pageNumber="110">Fig. 196</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF29F7AEFDF8FA28" pageId="109" pageNumber="110" type="etymology">
<paragraph id="8BA636F35B08FFFEFF29F7AEFDF8FA28" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
<emphasis id="B96DEAE15B08FFFEFF29F7AEFEFBFA0D" bold="true" box="[199,333,1411,1435]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Etymology.</emphasis>
Named in reference to the shape of the distal process of the aedeagus, which recalls the two-fingered universal peace (or victory) sign.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B08FFFEFF29F7E7FE1FF991" pageId="109" pageNumber="110" type="distribution">
<paragraph id="8BA636F35B08FFFEFF29F7E7FE1FF991" blockId="109.[151,1437,258,1543]" pageId="109" pageNumber="110">
<emphasis id="B96DEAE15B08FFFEFF29F7E7FED6FA75" bold="true" box="[199,352,1482,1507]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B08FFFEFD23F7E6FD4BFA75" box="[717,765,1483,1507]" pageId="109" pageNumber="110">type</typeStatus>
locality in the Huon Peninsula, in southeastern Area 5; elevation
<quantity id="4CE19B165B08FFFEFF0AF7C3FE9DF990" box="[228,299,1518,1542]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="109" pageNumber="110" unit="m" value="850.0">850 m</quantity>
(
<figureCitation id="13222A765B08FFFEFED4F7C3FE2FF990" box="[314,409,1518,1542]" captionStart="FIGURES 519522" captionStartId="405.[151,264,1650,1673]" captionTargetBox="[165,1421,253,1615]" captionTargetId="figure-20@405.[165,1421,193,1629]" captionTargetPageId="405" captionText="FIGURES 519522. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284131" httpUri="https://zenodo.org/record/5284131/files/figure.png" pageId="109" pageNumber="110">Fig. 521</figureCitation>
).
</paragraph>
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