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<document id="5EEC6854706FE3D25BC48E2CA4891FD9" ID-DOI="10.5852/ejt.2024.926.2469" ID-GBIF-Dataset="4511b4bd-f026-49d3-9d4d-a39875e75cc5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFF9FFFE6FE1AD58CFBBBFD44" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum neuberti Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner 2024, sp. nov." docType="treatment" docUuid="9BABF583-23ED-4ACF-92DE-1A52D8F7D0E0" docUuidSource="ZooBank" docVersion="10" lastPageNumber="39" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="38" updateTime="1722640713706" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7CBCAEEC83B72199B1586AFFE7CD34E9">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
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<mods:namePart id="FC1FDF96286D0E103772FEC5DBB30FE2">Jochum, Adrienne</mods:namePart>
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<mods:affiliation id="4DD243659108B0BABCB3E49AAC52C83C">Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, D- 60325 Frankfurt / M., Germany.</mods:affiliation>
<mods:nameIdentifier id="730B9FA6D350127A2354D736E78BC547" type="email">nne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="84B09E834E2B8F8C1FE9BA9A4F94DD1C">Michalik, Peter</mods:namePart>
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<mods:affiliation id="013474E664DFD82226ED30EC6C5FFC7C">Zoological Institute and Museum, University of Greifswald, Loitzer Strasse 26, D- 17489 Greifswald, Germany.</mods:affiliation>
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<mods:namePart id="A9BD39D0C00BF986B3C4554CD2E5A878">Inäbnit, Thomas</mods:namePart>
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<mods:affiliation id="2B95942B6AFF8BF153DD62A2B43BB131">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="BEA9CA6C12228F19C0A94F68425ACE80" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1A0809776E7E8720EEFC179A6E757721">Kneubühler, Jeannette</mods:namePart>
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<mods:affiliation id="815CADB9A4FF084E6C9FE982E20EE10D">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="782024026D0FDC064102DB59111F28B1" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
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<mods:namePart id="60E3D25ED08889D2419BFEAE67228B0D">Slapnik, Rajko</mods:namePart>
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<mods:affiliation id="24E648396DFD21BF8B7365CC134B05CD">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="54ADAFE77F9373D61D45FD5EA8162E95" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AD1B065880E88A24D45A9F6E86566748">Vrabec, Marko</mods:namePart>
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<mods:affiliation id="A61063ECC4256DF4727192D1F3A7F751">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="97967FBD35353EF5BADF6290D06F9EBF" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
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<mods:namePart id="4EC4232188ED57AF19F6B8BDACE7B438">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="6C4C6DEE562EEECC58D4AE928D03F8FC">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="42FB3BFD405898929829B6C0FEA0A18E">Ruthensteiner, Bernhard</mods:namePart>
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<mods:affiliation id="E5D4B077ABA44949754E75FA4DF32457">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="ABF782576656AB3BD72A01E94A2EA303" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
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<mods:title id="250B552143CB95D7979A78A8F2C8249B">European Journal of Taxonomy</mods:title>
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<mods:date id="66DCB4A968FD8F5D40362CA68AD4F163">2024</mods:date>
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<mods:number id="4E535A6729760DA6249289F45AAE7680">2024-03-20</mods:number>
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<mods:number id="E1DFFD499671B7B832A6E7A6AC4E4F57">926</mods:number>
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<mods:url id="AE0F502CD9E9F526807B443978C7F00E">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973</mods:url>
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<treatment id="03B0879DFF9FFFE6FE1AD58CFBBBFD44" ID-DOI="http://doi.org/10.5281/zenodo.10847844" ID-GBIF-Taxon="224067514" ID-Zenodo-Dep="10847844" LSID="urn:lsid:zoobank.org:act:9BABF583-23ED-4ACF-92DE-1A52D8F7D0E0" httpUri="http://treatment.plazi.org/id/03B0879DFF9FFFE6FE1AD58CFBBBFD44" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="C3036500FF9FFFE5FE1AD58CFB9BFD72" box="[485,1103,651,678]" pageId="37" pageNumber="38" type="nomenclature">
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<heading id="D0EE81E7FF9FFFE5FE1AD58CFB9BFD72" box="[485,1103,651,678]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C194D08FF9FFFE5FE1AD58CFC39FD72" ID-CoL="C8NTB" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[485,1005,651,678]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="neuberti" status="sp. nov.">
<emphasis id="B96DEA99FF9FFFE5FE1AD58CFD16FD72" bold="true" box="[485,706,651,678]" italics="true" pageId="37" pageNumber="38">Zospeum neuberti</emphasis>
Jochum &amp; Ruthensteiner
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<taxonomicNameLabel id="A25E57E2FF9FFFE5FC0AD58CFB9BFD72" box="[1013,1103,652,678]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3036500FF9FFFE5FE9CD5AFFC97FD38" pageId="37" pageNumber="38" type="description">
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<uri id="FF883A89FF9FFFE5FE9CD5AFFB04FD1E" box="[355,1232,687,714]" pageId="37" pageNumber="38">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FD0FD5D2FC97FD38" blockId="37.[355,1232,651,748]" box="[752,835,722,748]" pageId="37" pageNumber="38">
<figureCitation id="13222A0EFF9FFFE5FD0FD5D2FC97FD38" box="[752,835,722,748]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[186,1395,802,1819]" captionTargetId="figure-215@38.[186,1395,802,1819]" captionTargetPageId="38" captionText="Fig. 12. Zospeum neuberti Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 32006). A–B. Light microscopic images of paratypes showing aperture and dorsal views and collection labels. C. Light microscopic images of holotype showing aperture and dorsal views. D–I. 3D visualizations of Micro-CT data of holotype. D. Aperture view. E. Aperture facing right view showing smooth columella. F. Apical view.G. Dorsal view showing smooth columella. H. Aperture facing left view. I. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10847838" httpUri="https://zenodo.org/record/10847838/files/figure.png" pageId="37" pageNumber="38">Fig. 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9FFFE5FF42D413FC0EFC12" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BA6368BFF9FFFE5FF42D413FEE2FCF9" blockId="37.[189,310,787,813]" box="[189,310,787,813]" pageId="37" pageNumber="38">
<heading id="D0EE81E7FF9FFFE5FF42D413FEE2FCF9" bold="true" box="[189,310,787,813]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96DEA99FF9FFFE5FF42D413FEE2FCF9" bold="true" box="[189,310,787,813]" pageId="37" pageNumber="38">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF42D442FC0EFC12" blockId="37.[188,1399,833,966]" pageId="37" pageNumber="38">
Shell ca
<quantity id="4CE19B6EFF9FFFE5FED8D442FE52FC89" box="[295,390,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="37" pageNumber="38" unit="mm" value="1.3">1.3 mm</quantity>
, conical with 5¼ regularly coiled whorls, with blunt irregular growth lines; parietal shield thickly callused; columella centrally aligned, completely smooth; no lamella; ventral side swollen with well-defined umbilical depression; columellar side of peristome is positioned directly above umbilical depression with last coiling of whorl leaning to right of it.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9FFFE5FF42D4EDFAA5FBA8" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8BA6368BFF9FFFE5FF42D4EDFE90FBD3" blockId="37.[189,324,1005,1031]" box="[189,324,1005,1031]" pageId="37" pageNumber="38">
<heading id="D0EE81E7FF9FFFE5FF42D4EDFE90FBD3" bold="true" box="[189,324,1005,1031]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96DEA99FF9FFFE5FF42D4EDFE90FBD3" bold="true" box="[189,324,1005,1031]" pageId="37" pageNumber="38">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF42D31CFAA5FBA8" blockId="37.[189,1399,1052,1149]" pageId="37" pageNumber="38">
This species honors the German malacologist, mentor, and curator of molluscs, Dr Eike Neubert at the Naturhistorisches Museum der Burgergemeinde
<collectingRegion id="49DDF869FF9FFFE5FD08D340FCE4FB8E" box="[759,816,1088,1114]" country="Switzerland" name="Bern" pageId="37" pageNumber="38">Bern</collectingRegion>
(NMBE),
<collectingRegion id="49DDF869FF9FFFE5FC51D340FC3EFB8E" box="[942,1002,1088,1114]" country="Switzerland" name="Bern" pageId="37" pageNumber="38">Bern</collectingRegion>
,
<collectingCountry id="F30E761BFF9FFFE5FC09D33FFB56FB8D" box="[1014,1154,1087,1113]" name="Switzerland" pageId="37" pageNumber="38">Switzerland</collectingCountry>
in recognition of his support, dedicated mentorship, and contributions to malacology and specifically, to carychiid research.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9FFFE5FF42D3A4FB20FA6E" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BA6368BFF9FFFE5FF42D3A4FEB8FB69" blockId="37.[189,364,1187,1214]" box="[189,364,1187,1214]" pageId="37" pageNumber="38">
<heading id="D0EE81E7FF9FFFE5FF42D3A4FEB8FB69" bold="true" box="[189,364,1187,1214]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96DEA99FF9FFFE5FF42D3A4FEB8FB69" bold="true" box="[189,364,1187,1214]" pageId="37" pageNumber="38">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF15D3D2FE88FB38" blockId="37.[189,1398,1234,1331]" box="[234,348,1234,1260]" pageId="37" pageNumber="38">
<emphasis id="B96DEA99FF9FFFE5FF15D3D2FE88FB38" bold="true" box="[234,348,1234,1260]" pageId="37" pageNumber="38">
<typeStatus id="54A28829FF9FFFE5FF15D3D2FE88FB38" box="[234,348,1234,1260]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF42D3F6FB37FAE7" blockId="37.[189,1398,1234,1331]" pageId="37" pageNumber="38">
<materialsCitation id="3B713CD6FF9FFFE5FF42D3F6FB37FAE7" ID-GBIF-Occurrence="4598261314" collectionCode="NHMW Mol.Coll." collectorName="Edlauer" country="Croatia" county="Kuscer" latitude="42.7438" location="Grabovica cave" longLatPrecision="6" longitude="18.0578" municipality="Trebinje" pageId="37" pageNumber="38" specimenCode="NHMW Mol.Coll.Edlauer 32006" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F30E761BFF9FFFE5FF42D3F6FEE8FAC4" box="[189,316,1270,1296]" name="Croatia" pageId="37" pageNumber="38">CROATIA</collectingCountry>
• (
<figureCitation id="13222A0EFF9FFFE5FE9FD3F5FE32FAC4" box="[352,486,1269,1296]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[186,1395,802,1819]" captionTargetId="figure-215@38.[186,1395,802,1819]" captionTargetPageId="38" captionText="Fig. 12. Zospeum neuberti Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 32006). A–B. Light microscopic images of paratypes showing aperture and dorsal views and collection labels. C. Light microscopic images of holotype showing aperture and dorsal views. D–I. 3D visualizations of Micro-CT data of holotype. D. Aperture view. E. Aperture facing right view showing smooth columella. F. Apical view.G. Dorsal view showing smooth columella. H. Aperture facing left view. I. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10847838" httpUri="https://zenodo.org/record/10847838/files/figure.png" pageId="37" pageNumber="38">Fig. 12C–I</figureCitation>
); “Grabovica-Höhle bei Trebinje com. Kuščer” [
<location id="8EC66050FF9FFFE5FBD6D3F5FB31FAC4" LSID="urn:lsid:plazi:treatment:03B0879DFF9FFFE6FE1AD58CFBBBFD44:8EC66050FF9FFFE5FBD6D3F5FB31FAC4" box="[1065,1253,1269,1296]" country="Croatia" county="Kuscer" latitude="42.7438" longLatPrecision="6" longitude="18.0578" municipality="Trebinje" name="Grabovica cave" pageId="37" pageNumber="38">Grabovica cave</location>
by Trebinje communicated with Kuščer]; [
<geoCoordinate id="EE2D504CFF9FFFE5FDDFD219FD79FAE7" box="[544,685,1305,1331]" degrees="42.7438" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="42.7438">42.7438° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFF9FFFE5FD46D218FC95FAE7" box="[697,833,1304,1331]" degrees="18.0578" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="18.0578">18.0578° E</geoCoordinate>
];
<collectionCode id="ED08AE4EFF9FFFE5FCA9D219FBFFFAE7" box="[854,1067,1305,1331]" pageId="37" pageNumber="38">NHMW Mol.Coll.</collectionCode>
<collectorName id="26EC535DFF9FFFE5FBD5D219FB5EFAE7" box="[1066,1162,1305,1331]" pageId="37" pageNumber="38">Edlauer</collectorName>
32006.
</materialsCitation>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF15D25AFEB3FAA0" blockId="37.[189,1398,1370,1467]" box="[234,359,1370,1396]" pageId="37" pageNumber="38">
<emphasis id="B96DEA99FF9FFFE5FF15D25AFEB3FAA0" bold="true" box="[234,359,1370,1396]" pageId="37" pageNumber="38">
<typeStatus id="54A28829FF9FFFE5FF15D25AFEB3FAA0" box="[234,359,1370,1396]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF42D27DFB20FA6E" blockId="37.[189,1398,1370,1467]" pageId="37" pageNumber="38">
<materialsCitation id="3B713CD6FF9FFFE5FF42D27DFAA2FA43" ID-GBIF-Occurrence="4598261307" box="[189,1398,1405,1432]" collectionCode="NHMW Mol.Coll." collectorName="Edlauer" country="Croatia" county="Kuscer" latitude="42.7438" location="CROATIA" longLatPrecision="6" longitude="18.0578" municipality="Trebinje" pageId="37" pageNumber="38" specimenCode="NHMW Mol.Coll.Edlauer 32006" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F30E761BFF9FFFE5FF42D27DFEE8FA43" box="[189,316,1405,1431]" name="Croatia" pageId="37" pageNumber="38">CROATIA</collectingCountry>
•
<specimenCount id="9D1FFD02FF9FFFE5FEABD27DFE72FA4C" box="[340,422,1405,1432]" count="1" pageId="37" pageNumber="38" type="generic">1 spec.</specimenCount>
(
<figureCitation id="13222A0EFF9FFFE5FE49D27DFDF7FA43" box="[438,547,1405,1431]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[186,1395,802,1819]" captionTargetId="figure-215@38.[186,1395,802,1819]" captionTargetPageId="38" captionText="Fig. 12. Zospeum neuberti Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 32006). A–B. Light microscopic images of paratypes showing aperture and dorsal views and collection labels. C. Light microscopic images of holotype showing aperture and dorsal views. D–I. 3D visualizations of Micro-CT data of holotype. D. Aperture view. E. Aperture facing right view showing smooth columella. F. Apical view.G. Dorsal view showing smooth columella. H. Aperture facing left view. I. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10847838" httpUri="https://zenodo.org/record/10847838/files/figure.png" pageId="37" pageNumber="38">Fig. 12A</figureCitation>
); same collection data as for holotype;
<collectionCode id="ED08AE4EFF9FFFE5FC0ED27EFB12FA43" box="[1009,1222,1405,1432]" pageId="37" pageNumber="38">NHMW Mol.Coll.</collectionCode>
<collectorName id="26EC535DFF9FFFE5FB3AD27DFAF1FA43" box="[1221,1317,1405,1431]" pageId="37" pageNumber="38">Edlauer</collectorName>
32006
</materialsCitation>
•
<materialsCitation id="3B713CD6FF9FFFE5FF30D2A1FB24FA6E" ID-GBIF-Occurrence="4598261331" box="[207,1264,1440,1467]" collectionCode="NHMW Mol.Coll." collectorName="Edlauer" country="Croatia" county="Kuscer" latitude="42.7438" location="Grabovica cave" longLatPrecision="6" longitude="18.0578" municipality="Trebinje" pageId="37" pageNumber="38" specimenCode="NHMW Mol.Coll.Edlauer 32006" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D1FFD02FF9FFFE5FF30D2A1FEF4FA6F" box="[207,288,1441,1467]" count="1" pageId="37" pageNumber="38" type="generic">1 spec.</specimenCount>
(
<figureCitation id="13222A0EFF9FFFE5FECFD2A0FE4FFA6F" box="[304,411,1440,1467]" captionStart="Fig" captionStartId="38.[189,232,1857,1883]" captionTargetBox="[186,1395,802,1819]" captionTargetId="figure-215@38.[186,1395,802,1819]" captionTargetPageId="38" captionText="Fig. 12. Zospeum neuberti Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 32006). A–B. Light microscopic images of paratypes showing aperture and dorsal views and collection labels. C. Light microscopic images of holotype showing aperture and dorsal views. D–I. 3D visualizations of Micro-CT data of holotype. D. Aperture view. E. Aperture facing right view showing smooth columella. F. Apical view.G. Dorsal view showing smooth columella. H. Aperture facing left view. I. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10847838" httpUri="https://zenodo.org/record/10847838/files/figure.png" pageId="37" pageNumber="38">Fig. 12B</figureCitation>
); same collection data as for holotype;
<collectionCode id="ED08AE4EFF9FFFE5FC98D2A1FBE8FA6E" box="[871,1084,1440,1467]" pageId="37" pageNumber="38">NHMW Mol.Coll.</collectionCode>
<collectorName id="26EC535DFF9FFFE5FBC4D2A0FB4FFA6E" box="[1083,1179,1440,1466]" pageId="37" pageNumber="38">Edlauer</collectorName>
32006
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9FFFE6FF42D2E2FA93FE93" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="8BA6368BFF9FFFE5FF42D2E2FE9AFA28" blockId="37.[189,334,1506,1532]" box="[189,334,1506,1532]" pageId="37" pageNumber="38">
<heading id="D0EE81E7FF9FFFE5FF42D2E2FE9AFA28" bold="true" box="[189,334,1506,1532]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96DEA99FF9FFFE5FF42D2E2FE9AFA28" bold="true" box="[189,334,1506,1532]" pageId="37" pageNumber="38">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9FFFE5FF42D124FC73F950" blockId="37.[188,1398,1571,1669]" pageId="37" pageNumber="38">
<smallCapsWord id="8D40A057FF9FFFE5FF42D124FEBAF9E8" baselines="1591,1591" box="[189,366,1572,1598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="37" pageNumber="38">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="54A28829FF9FFFE5FE83D123FE3AF9E9" box="[380,494,1571,1597]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
: sh:
<quantity id="4CE19B6EFF9FFFE5FDDED123FD4EF9EA" box="[545,666,1571,1598]" metricMagnitude="-3" metricUnit="m" metricValue="1.308" pageId="37" pageNumber="38" unit="mm" value="1.308">1.308 mm</quantity>
; sw:
<quantity id="4CE19B6EFF9FFFE5FD2AD123FC9AF9EA" box="[725,846,1571,1598]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" pageId="37" pageNumber="38" unit="mm" value="0.915">0.915 mm</quantity>
; ah:
<quantity id="4CE19B6EFF9FFFE5FC7BD123FC29F9EA" box="[900,1021,1571,1598]" metricMagnitude="-4" metricUnit="m" metricValue="5.55" pageId="37" pageNumber="38" unit="mm" value="0.555">0.555 mm</quantity>
; aw:
<quantity id="4CE19B6EFF9FFFE5FBC6D123FB66F9EA" box="[1081,1202,1571,1598]" metricMagnitude="-4" metricUnit="m" metricValue="5.62" pageId="37" pageNumber="38" unit="mm" value="0.562">0.562 mm</quantity>
; hlw:
<quantity id="4CE19B6EFF9FFFE5FB07D123FAA6F9EA" box="[1272,1394,1571,1598]" metricMagnitude="-4" metricUnit="m" metricValue="7.94" pageId="37" pageNumber="38" unit="mm" value="0.794">0.794 mm</quantity>
; SA: 61.88 deg.
<typeStatus id="54A28829FF9FFFE5FE8FD147FE36F9B5" box="[368,482,1607,1633]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
(N = 2): sh:
<quantity id="4CE19B6EFF9FFFE5FD8BD146FCE9F9B5" box="[628,829,1606,1633]" metricMagnitude="-3" metricUnit="m" metricValue="1.255" metricValueMax="1.305" metricValueMin="1.205" pageId="37" pageNumber="38" unit="mm" value="1.255" valueMax="1.305" valueMin="1.205">
<geoCoordinate id="EE2D504CFF9FFFE5FD8BD146FD61F9B5" box="[628,693,1606,1633]" degrees="1.205" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.205">1.205</geoCoordinate>
<geoCoordinate id="EE2D504CFF9FFFE5FD4AD146FCD2F9B5" box="[693,774,1606,1633]" degrees="1.305" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-1.305">–1.305</geoCoordinate>
mm
</quantity>
; sw:
<quantity id="4CE19B6EFF9FFFE5FC88D146FB95F9B5" box="[887,1089,1606,1633]" metricMagnitude="-4" metricUnit="m" metricValue="8.93" metricValueMax="9.32" metricValueMin="8.54" pageId="37" pageNumber="38" unit="mm" value="0.893" valueMax="0.932" valueMin="0.854">
<geoCoordinate id="EE2D504CFF9FFFE5FC88D146FC6DF9B5" box="[887,953,1606,1633]" degrees="0.854" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.854">0.854</geoCoordinate>
<geoCoordinate id="EE2D504CFF9FFFE5FC46D146FBDEF9B5" box="[953,1034,1606,1633]" degrees="0.932" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.932">–0.932</geoCoordinate>
mm
</quantity>
; ah:
<quantity id="4CE19B6EFF9FFFE5FB89D146FA94F9B5" box="[1142,1344,1606,1633]" metricMagnitude="-4" metricUnit="m" metricValue="5.244999999999999" metricValueMax="5.32" metricValueMin="5.17" pageId="37" pageNumber="38" unit="mm" value="0.5245" valueMax="0.532" valueMin="0.517">
<geoCoordinate id="EE2D504CFF9FFFE5FB89D146FB63F9B5" box="[1142,1207,1606,1633]" degrees="0.517" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.517">0.517</geoCoordinate>
<geoCoordinate id="EE2D504CFF9FFFE5FB47D146FADDF9B5" box="[1208,1289,1606,1633]" degrees="0.532" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.532">–0.532</geoCoordinate>
mm
</quantity>
; aw:
<quantity id="4CE19B6EFF9FFFE5FF43D16AFE53F950" box="[188,391,1642,1669]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.51" metricValueMin="5.39" pageId="37" pageNumber="38" unit="mm" value="0.545" valueMax="0.551" valueMin="0.539">
<geoCoordinate id="EE2D504CFF9FFFE5FF43D16AFF2BF951" box="[188,255,1642,1669]" degrees="0.539" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.539">0.539</geoCoordinate>
<geoCoordinate id="EE2D504CFF9FFFE5FF00D16AFE9BF951" box="[255,335,1642,1669]" degrees="0.551" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.551">–0.551</geoCoordinate>
mm
</quantity>
; hlw:
<quantity id="4CE19B6EFF9FFFE5FE31D16AFD4DF950" box="[462,665,1642,1668]" metricMagnitude="-4" metricUnit="m" metricValue="7.615" metricValueMax="7.86" metricValueMin="7.37" pageId="37" pageNumber="38" unit="mm" value="0.7615" valueMax="0.786" valueMin="0.737">
<geoCoordinate id="EE2D504CFF9FFFE5FE31D16AFDC4F950" box="[462,528,1642,1668]" degrees="0.737" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.737">0.737</geoCoordinate>
<geoCoordinate id="EE2D504CFF9FFFE5FDEFD16AFDB5F950" box="[528,609,1642,1668]" degrees="0.786" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.786">–0.786</geoCoordinate>
mm
</quantity>
; SA: 66.77–67.49 deg.
</paragraph>
<paragraph id="8BA6368BFF9FFFE6FF42D1ABFA93FE93" blockId="37.[189,1401,1706,1980]" lastBlockId="38.[189,1399,265,327]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Shell ca
<quantity id="4CE19B6EFF9FFFE5FED9D1ABFE51F912" box="[294,389,1707,1734]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="37" pageNumber="38" unit="mm" value="1.3">1.3 mm</quantity>
, conical, with 5–5¼ regularly coiled convex whorls; transparent when fresh, opaque when older; aperture reniform; teleoconch sculpture with few irregular and occasional course growth lines; some weak axial ribbing present immediately behind palatal lip; palatal-basal lip narrowly reflected; aperture wider than high; peristome thickly callused on parietal shield and on basal columellar side, upper palatal side thin and slightly expanded, columellar side more straight than oblique, roundish not angular; parietal shield long and straight with narrow notch indentation at upper parietal and palatal junction; height of last whorl greater than half of shell height; upper rim of peristome recedes ca
<superScript id="7C6C9BC3FF9FFFE5FACDD07EFAEEF858" attach="right" box="[1330,1338,1918,1932]" fontSize="6" pageId="37" pageNumber="38">1</superScript>
/
<subScript id="179D34CEFF9FFFE5FABDD089FA9FF843" attach="left" box="[1346,1355,1929,1943]" fontSize="6" pageId="37" pageNumber="38">6</subScript>
the width of the penultimate whorl (aperture facing left); columella centrally aligned, moderately thick, smooth; umbilical depression callused; columellar side of peristome positioned directly above with last coiling of whorl leaning to right of umbilical depression, alignment of last ¼ whorl is not compact.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9CFFE6FF42D66DFAF7FE62" box="[189,1315,365,439]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BA6368BFF9CFFE6FF42D66DFE82FE53" blockId="38.[189,342,365,391]" box="[189,342,365,391]" pageId="38" pageNumber="39">
<heading id="D0EE81E7FF9CFFE6FF42D66DFE82FE53" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B96DEA99FF9CFFE6FF42D66DFE82FE53" bold="true" box="[189,342,365,391]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9CFFE6FF42D69CFAF7FE62" blockId="38.[189,1315,412,439]" box="[189,1315,412,439]" pageId="38" pageNumber="39">
This species is only known from its
<typeStatus id="54A28829FF9CFFE6FD9AD69DFD4CFE63" box="[613,664,413,439]" pageId="38" pageNumber="39">type</typeStatus>
locality cave, Grabovica jama in southwestern
<collectingCountry id="F30E761BFF9CFFE6FB3AD69CFACBFE62" box="[1221,1311,412,438]" name="Croatia" pageId="38" pageNumber="39">Croatia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9CFFE6FF42D6DEFBBBFD44" pageId="38" pageNumber="39" type="discussion">
<paragraph id="8BA6368BFF9CFFE6FF42D6DEFEFBFE2C" blockId="38.[189,303,478,504]" box="[189,303,478,504]" pageId="38" pageNumber="39">
<heading id="D0EE81E7FF9CFFE6FF42D6DEFEFBFE2C" bold="true" box="[189,303,478,504]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B96DEA99FF9CFFE6FF42D6DEFEFBFE2C" bold="true" box="[189,303,478,504]" pageId="38" pageNumber="39">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9CFFE6FF42D50DFBBBFD44" blockId="38.[189,1398,524,656]" pageId="38" pageNumber="39">
<taxonomicName id="4C194D08FF9CFFE6FF42D50DFD18FDF2" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[189,716,524,550]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="neuberti" status="sp. nov.">
<emphasis id="B96DEA99FF9CFFE6FF42D50DFE42FDF2" box="[189,406,524,550]" italics="true" pageId="38" pageNumber="39">Zospeum neuberti</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF9CFFE6FD28D50DFCE2FDF3" box="[727,822,525,551]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
is the only species to possess a totally smooth columella. Although the narrow shell spire and the straight parietal shield resembles that of
<taxonomicName id="4C194D08FF9CFFE6FB0AD530FABBFD9D" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[1269,1391,559,585]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF9CFFE6FB0AD530FABBFD9D" box="[1269,1391,559,585]" italics="true" pageId="38" pageNumber="39">Z. simplex</emphasis>
</taxonomicName>
, it differs from this species in that
<taxonomicName id="4C194D08FF9CFFE6FDB4D553FD1AFDB8" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[587,718,594,620]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="neuberti">
<emphasis id="B96DEA99FF9CFFE6FDB4D553FD1AFDB8" box="[587,718,594,620]" italics="true" pageId="38" pageNumber="39">Z. neuberti</emphasis>
</taxonomicName>
is smaller in shell height, aperture height, aperture width, height of the last whorl and it bears a much broader spire than that of
<taxonomicName id="4C194D08FF9CFFE6FC11D576FBB3FD44" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[1006,1127,630,656]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF9CFFE6FC11D576FBB3FD44" box="[1006,1127,630,656]" italics="true" pageId="38" pageNumber="39">Z. simplex</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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