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<document id="28F1F0C0FB426A7510A2BE43B9BA371C" ID-CLB-Dataset="7783" ID-DOI="10.5852/ejt.2021.738.1261" ID-GBIF-Dataset="02cf5931-b18e-4d58-a6ff-9c7b1e37e68d" ID-ISSN="2118-9773" ID-Zenodo-Dep="4594541" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615410255512" checkinUser="plazi" docAuthor="Turner, Ian M. &amp; Cheek, Martin" docDate="2021" docId="355087D8FFEFE47A1864FC28EA6EC181" docLanguage="en" docName="ejt-738_1-60.pdf" docOrigin="European Journal of Taxonomy 738" docSource="http://dx.doi.org/10.5852/ejt.2021.738.1261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chassalia involucrata T. Y. Yu 2021, sp. nov." docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="C969FFA0FFF9E4631A5DFFA8EF2BC42F" masterDocTitle="Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species" masterLastPageNumber="60" masterPageNumber="1" pageNumber="23" updateTime="1698932934732" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="DD7EAE5A36B541D5EFF91802113C2AC7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="ED3A88503BDFBBFFEF28CA6D329CE635">Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species</mods:title>
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<mods:namePart id="F90E79A772D71AB6B7CF79ACA3958462">Turner, Ian M.</mods:namePart>
<mods:nameIdentifier id="84C1E28D506A162A7930F507FD318956" type="email">i.turner@kew.org</mods:nameIdentifier>
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<mods:namePart id="0A5B024929D4BF6E2393EA98A6A99593">Cheek, Martin</mods:namePart>
<mods:affiliation id="42511E2F8941CE30E0C23A474D6B9BD1">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Queen Mary University of London, E 1 4 NS, UK. Singapore Botanical Liaison Officer, Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569, Singapore.</mods:affiliation>
<mods:nameIdentifier id="6EE17E17E03391DC243A87F9C2AE0293" type="email">m.cheek@kew.org</mods:nameIdentifier>
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<mods:title id="BDB3E41C6701C7B0E9234E9DE5A31723">European Journal of Taxonomy</mods:title>
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<mods:part id="4C7CFC98EC118CE6DB5C1B87CE6EEA67">
<mods:date id="A693350A9D5448BB68CAD3A1262451FF">2021</mods:date>
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<mods:number id="B8CEE92399B680F1C2877774D9B78F47">2021-03-09</mods:number>
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<treatment id="355087D8FFEFE47A1864FC28EA6EC181" ID-DOI="http://doi.org/10.5281/zenodo.4599397" ID-GBIF-Taxon="180327052" ID-Zenodo-Dep="4599397" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181" httpUri="http://treatment.plazi.org/id/355087D8FFEFE47A1864FC28EA6EC181" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection id="F5E36545FFEFE4751864FC28ECD0C7B4" box="[569,1019,896,923]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="BD4636CEFFEFE4751864FC28ECD0C7B4" blockId="22.[569,1019,896,995]" box="[569,1019,896,923]" pageId="22" pageNumber="23">
<heading id="E60E81A2FFEFE4751864FC28ECD0C7B4" box="[569,1019,896,923]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName id="7AF94D4DFFEFE4751864FC28ECB1C7B4" ID-CoL="9V52Y" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[569,922,896,923]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="involucrata" status="sp. nov.">
<emphasis id="8F8DEADCFFEFE4751864FC28EC69C7B4" bold="true" box="[569,834,896,923]" italics="true" pageId="22" pageNumber="23">Chassalia involucrata</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFEFE47519FCFC29ECD0C7B4" box="[929,1019,897,923]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFEFE475181AFC0CEC71C7CC" pageId="22" pageNumber="23" type="description">
<paragraph id="BD4636CEFFEFE475181AFC0CEC71C7CC" blockId="22.[569,1019,896,995]" pageId="22" pageNumber="23">
urn:lsid:ipni.org:names:77215406-1
<figureCitation id="25C22A4BFFEFE4751887FC61EC71C7CC" box="[730,858,969,995]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="23.[190,233,1858,1884]" captionStartId-1="24.[189,232,1999,2025]" captionTargetBox-0="[224,1317,329,1810]" captionTargetBox-1="[267,1321,265,1961]" captionTargetId-0="figure-12@23.[223,1364,262,1818]" captionTargetId-1="figure-13@24.[266,1321,265,1961]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="Fig. 12. Chassalia involucrata T.Y.Yu sp. nov. A. Flowering shoot. B. Inflorescence. C. Flower. D. Longitudinal view of corolla showing stamens, band of hairs, stigma, style and disc.E. Infructescence with stem node showing fresh fruit. F. Dry fruit. G. Pyrene ventral surface. A, C–D, F–G from Carlo Hansen 1608; B, E from photographs of living plants in Ulu Temburong National Park, Brunei, taken by Mr Y.W. Low. Drawn by T.Y. Yu." captionText-1="Fig. 13. Holotype of Chassalia involucrata T.Y.Yu sp. nov. (Carlo Hansen 1608)." figureDoi-0="http://doi.org/10.5281/zenodo.4594571" figureDoi-1="http://doi.org/10.5281/zenodo.4594574" httpUri-0="https://zenodo.org/record/4594571/files/figure.png" httpUri-1="https://zenodo.org/record/4594574/files/figure.png" pageId="22" pageNumber="23">Fig. 12–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFEFE4751AE0FBA3EE87C17D" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="BD4636CEFFEFE4751AE0FBA3EE1DC00A" blockId="22.[189,310,1035,1061]" box="[189,310,1035,1061]" pageId="22" pageNumber="23">
<heading id="E60E81A2FFEFE4751AE0FBA3EE1DC00A" bold="true" box="[189,310,1035,1061]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="8F8DEADCFFEFE4751AE0FBA3EE1DC00A" bold="true" box="[189,310,1035,1061]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFEFE4751AE0FB93EE87C17D" blockId="22.[189,1400,1082,1363]" pageId="22" pageNumber="23">
Differs from other species of
<taxonomicName id="7AF94D4DFFEFE4751841FB93EDBAC07A" box="[540,657,1083,1109]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="8F8DEADCFFEFE4751841FB93EDBAC07A" box="[540,657,1083,1109]" italics="true" pageId="22" pageNumber="23">Chassalia</emphasis>
</taxonomicName>
in Borneo in having large leaves up to
<quantity id="7A019B2BFFEFE4751E37FB93EB9FC07A" box="[1130,1204,1083,1109]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" pageId="22" pageNumber="23" unit="cm" value="48.0">48 cm</quantity>
long and
<quantity id="7A019B2BFFEFE4751F70FB92EA5CC07A" box="[1325,1399,1082,1109]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="cm" value="15.0">15 cm</quantity>
wide (they are much smaller in other species), in the capitulum having outer and inner bracts up to
<quantity id="7A019B2BFFEFE4751AE0FB2BEE24C0B1" box="[189,271,1155,1182]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="cm" value="1.5">1.5 cm</quantity>
wide (broader than in any other species), in the infructescences usually persisting for a whole season until new inflorescences are produced (not soon disappearing after fruit maturity). Most similar to, but differing from,
<taxonomicName id="7AF94D4DFFEFE4751B8CFB64ED72C0CA" authority="Turner &amp; Cheek, 2021" authorityName="Turner &amp; Cheek" authorityYear="2021" box="[465,601,1227,1254]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="calamus" status="sp. nov.">
<emphasis id="8F8DEADCFFEFE4751B8CFB64ED72C0CA" box="[465,601,1227,1254]" italics="true" pageId="22" pageNumber="23">C. calamus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFEFE475183EFB64EDE9C0C9" box="[611,706,1228,1254]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
in having bracts usually longer than the flower bud (not shorter), in having a longer peduncle, in having bigger leaves (to 48 ×
<quantity id="7A019B2BFFEFE4751E54FB58EB7DC125" box="[1033,1110,1264,1290]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="cm" value="12.0">12 cm</quantity>
, not to 33 ×
<quantity id="7A019B2BFFEFE4751EADFB58EA17C125" box="[1264,1340,1264,1290]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="cm" value="12.0">12 cm</quantity>
) and more secondary veins (up to 26 on each side of the midrib, not &lt;18), and in having longer (
<quantity id="7A019B2BFFEFE4751F5AFABCEA59C100" box="[1287,1394,1300,1327]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="14.0" metricValueMin="4.0" pageId="22" pageNumber="23" unit="cm" value="9.0" valueMax="14.0" valueMin="4.0">4–14 cm</quantity>
, not &lt;
<quantity id="7A019B2BFFEFE4751AA3FA90EE11C17C" box="[254,314,1336,1363]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="cm" value="5.0">5 cm</quantity>
) petioles.
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFEFE4751AE0FAD3EC4BC1C6" pageId="22" pageNumber="23" type="etymology">
<paragraph id="BD4636CEFFEFE4751AE0FAD3EE6FC1BA" blockId="22.[189,324,1403,1429]" box="[189,324,1403,1429]" pageId="22" pageNumber="23">
<heading id="E60E81A2FFEFE4751AE0FAD3EE6FC1BA" bold="true" box="[189,324,1403,1429]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="8F8DEADCFFEFE4751AE0FAD3EE6FC1BA" bold="true" box="[189,324,1403,1429]" pageId="22" pageNumber="23">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFEFE4751AE0FA02EC4BC1C6" blockId="22.[189,1398,1450,1513]" pageId="22" pageNumber="23">The epithet alludes to the involucrate bracts of rachis and capitulum, which are larger and more conspicuous than in other species of the genus in Borneo.</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFEFE47A1AE0F9B9EA5DC566" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="BD4636CEFFEFE4751AE0F9B9EFD1C204" blockId="22.[189,250,1553,1579]" box="[189,250,1553,1579]" pageId="22" pageNumber="23">
<heading id="E60E81A2FFEFE4751AE0F9B9EFD1C204" bold="true" box="[189,250,1553,1579]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="8F8DEADCFFEFE4751AE0F9B9EFD1C204" bold="true" box="[189,250,1553,1579]" pageId="22" pageNumber="23">
<typeStatus id="6242886CFFEFE4751AE0F9B9EFD1C204" box="[189,250,1553,1579]" pageId="22" pageNumber="23">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFEFE4751AE0F9E9ED9AC28C" blockId="22.[189,1398,1600,1699]" pageId="22" pageNumber="23">
<materialsCitation id="0D913C93FFEFE4751AE0F9E9ED82C28C" ID-GBIF-Occurrence="3050815330" collectingDate="1991-11-22" collectionCode="K" collectorName="Carlo Hansen" country="Brunei" county="Kuala Belalong" latitude="4.55" location="Batu Apoi Forest Reserve" longLatPrecision="1306" longitude="115.15" municipality="Batu Apoi Forest Reserve" pageId="22" pageNumber="23" specimenCode="K001129724" specimenCount="1" stateProvince="Temburong district" typeStatus="holotype">
<collectingCountry id="C5EE765EFFEFE4751AE0F9E9EE01C274" box="[189,298,1601,1627]" name="Brunei" pageId="22" pageNumber="23">BRUNEI</collectingCountry>
•
<collectingRegion id="7F3DF82CFFEFE4751B17F9E8ED1AC275" box="[330,561,1600,1626]" country="Brunei" name="Temburong" pageId="22" pageNumber="23">Temburong district</collectingRegion>
,
<collectingCounty id="54274E42FFEFE475181DF9E8EDD5C275" box="[576,766,1600,1626]" pageId="22" pageNumber="23">Kuala Belalong</collectingCounty>
,
<collectingMunicipality id="5D22ACB4FFEFE4751953F9E9EB61C274" box="[782,1098,1600,1627]" pageId="22" pageNumber="23">Batu Apoi Forest Reserve</collectingMunicipality>
,
<locationDeviation id="ACF42FECFFEFE4751E07F9E9ECA5C250" pageId="22" pageNumber="23">at the Sungai Engkiang River, towards the Sungai Belalong, below Pondok Engkiang</locationDeviation>
;
<geoCoordinate id="D8CD5009FFEFE47519C7F9CBEB24C250" box="[922,1039,1635,1663]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="115.15">115°09′ E</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFEFE4751E46F9CBEB52C250" box="[1051,1145,1635,1663]" degrees="4" direction="north" minutes="33" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="4.55">4°33′ N</geoCoordinate>
;
<date id="C947100EFFEFE4751ED8F9CDEA0DC250" box="[1157,1318,1637,1663]" pageId="22" pageNumber="23" value="1991-11-22">
<collectingDate id="D903E9E6FFEFE4751ED8F9CDEA0DC250" box="[1157,1318,1637,1663]" pageId="22" pageNumber="23" value="1991-11-22">22 Nov. 1991</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFEFE4751F6FF9CDEE72C28C" italics="true" pageId="22" pageNumber="23">
<collectorName id="100C5318FFEFE4751F6FF9CDEE3CC28C" pageId="22" pageNumber="23">Carlo Hansen</collectorName>
1608
</emphasis>
;
<typeStatus id="6242886CFFEFE4751B35F921EEF8C28C" box="[360,467,1673,1699]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
:
<collectionCode id="DBE8AE0BFFEFE4751B83F920EEDFC28C" box="[478,500,1672,1699]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
[
<specimenCode id="ED5F9EB5FFEFE4751BA6F920ED89C28C" box="[507,674,1672,1699]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K001129724</specimenCode>
]
</materialsCitation>
!.
</paragraph>
<paragraph id="BD4636CEFFEFE4751AE0F963EE9FC2CA" blockId="22.[189,436,1739,1765]" box="[189,436,1739,1765]" pageId="22" pageNumber="23">
<heading id="E60E81A2FFEFE4751AE0F963EE9FC2CA" bold="true" box="[189,436,1739,1765]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="8F8DEADCFFEFE4751AE0F963EE9FC2CA" bold="true" box="[189,436,1739,1765]" pageId="22" pageNumber="23">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFEFE4751AE0F953EE57C371" blockId="22.[189,1400,1787,1886]" pageId="22" pageNumber="23">
<materialsCitation id="0D913C93FFEFE4751AE0F953EE25C315" ID-GBIF-Occurrence="3050815320" collectingDate="1959-02-11" collectionCode="BRUN, K, SING" collectorName="P. S. Ashton" country="Brunei" location="BRUNEI" municipality="Belalong" pageId="22" pageNumber="23" specimenCode="BRUN 5227" specimenCount="1">
<collectingCountry id="C5EE765EFFEFE4751AE0F953EE01C33A" box="[189,298,1787,1813]" name="Brunei" pageId="22" pageNumber="23">BRUNEI</collectingCountry>
•
<locationDeviation id="ACF42FECFFEFE4751B1FF953ED97C33A" box="[322,700,1787,1813]" pageId="22" pageNumber="23">S. Belalong below Kuala Ropan</locationDeviation>
;
<date id="C947100EFFEFE475189AF953EC49C339" box="[711,866,1787,1814]" pageId="22" pageNumber="23" value="1959-02-11">
<collectingDate id="D903E9E6FFEFE475189AF953EC49C339" box="[711,866,1787,1814]" pageId="22" pageNumber="23" value="1959-02-11">11 Feb. 1959</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFEFE4751933F953EBADC33A" box="[878,1158,1787,1813]" italics="true" pageId="22" pageNumber="23">
<collectorName id="100C5318FFEFE4751933F953ECDBC33A" box="[878,1008,1787,1813]" pageId="22" pageNumber="23">P.S. Ashton</collectorName>
<specimenCode id="ED5F9EB5FFEFE47519A5F954EBADC33A" box="[1016,1158,1787,1813]" collectionCode="BRUN" country="Brunei" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12726" lsid="urn:lsid:biocol.org:col:12726" name="Brunei Forestry Centre" pageId="22" pageNumber="23" type="Herbarium">BRUN 5227</specimenCode>
</emphasis>
;
<collectionCode id="DBE8AE0BFFEFE4751EC9F953EBC3C33A" box="[1172,1256,1787,1813]" country="Brunei" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12726" lsid="urn:lsid:biocol.org:col:12726" name="Brunei Forestry Centre" pageId="22" pageNumber="23" type="Herbarium">BRUN</collectionCode>
,
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!,
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!,
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•
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, LZ 298,
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;
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;
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.
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.
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484
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;
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!,
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</materialsCitation>
!.
</paragraph>
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<materialsCitation id="0D913C93FFEFE4751AE0F82EEA5CC3EB" ID-GBIF-Occurrence="3051662301" collectingDate="1983-10-11" collectionCode="K, MSC, UKMS" collectorName="John Beaman" country="Malaysia" county="Tambunan District" elevation="1250" latitude="5.7666664" location="Borneo" longLatPrecision="1305" longitude="-116.35" municipality="Crocker Range" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Sabah">
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–
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<location id="B8266015FFEFE4751B21F82EEEF2C38F" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181:B8266015FFEFE4751B21F82EEEF2C38F" box="[380,473,1926,1952]" country="Malaysia" name="Borneo" pageId="22" pageNumber="23" stateProvince="Sabah">Borneo</location>
,
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</emphasis>
•
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,
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,
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;
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,
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; alt.
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</quantity>
;
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<collectingDate id="D903E9E6FFEFE4751953F802EC8CC3EB" box="[782,935,1962,1988]" pageId="22" pageNumber="23" value="1983-10-11">11 Oct. 1983</collectingDate>
</date>
;
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<collectorName id="100C5318FFEFE47519EFF802EB79C3EB" box="[946,1106,1962,1988]" pageId="22" pageNumber="23">John Beaman</collectorName>
7268
</emphasis>
;
<collectionCode id="DBE8AE0BFFEFE4751EFFF802EB95C3EB" box="[1186,1214,1962,1988]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFEFE4751E92F802EA3BC3EB" box="[1231,1296,1962,1988]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15587" lsid="urn:lsid:biocol.org:col:15587" name="Michigan State University" pageId="22" pageNumber="23" type="Herbarium">MSC</collectionCode>
,
<collectionCode id="DBE8AE0BFFEFE4751F47F802EA58C3EB" box="[1306,1395,1962,1988]" collectionName="Sudan, Khartoum, University of Khartoum, Sudan Natural History Museum" country="Sudan" httpUri="http://grbio.org/cool/vy3j-vjek" name="Sudan Natural History Museum" pageId="22" pageNumber="23">UKMS</collectionCode>
.
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–
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<location id="B8266015FFEFE4751A8FF866EE04C3C7" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181:B8266015FFEFE4751A8FF866EE04C3C7" box="[210,303,1998,2024]" country="Malaysia" county="Tambunan District" latitude="5.7666664" longLatPrecision="1305" longitude="-116.35" municipality="Crocker Range" name="Borneo" pageId="22" pageNumber="23" stateProvince="Sarawak">Borneo</location>
,
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</emphasis>
•
<collectingMunicipality id="5D22ACB4FFEFE4751B9FF866EDA2C3C7" box="[450,649,1998,2024]" pageId="22" pageNumber="23">Kuching District</collectingMunicipality>
,
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12
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mile Penrissen Road
</locationDeviation>
;
<date id="C947100EFFEFE4751998F866EB49C3C7" box="[965,1122,1998,2024]" pageId="22" pageNumber="23" value="1966-09-19">
<collectingDate id="D903E9E6FFEFE4751998F866EB49C3C7" box="[965,1122,1998,2024]" pageId="22" pageNumber="23" value="1966-09-19">19 Sep. 1966</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFEFE47A1E30F867ED21C50C" italics="true" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
Banyeng ak Nudong et Benang ak Bubong,
<accessionNumber id="A2AAAB2DFFE0E47A1BF6FEA1ED21C50C" box="[427,522,265,291]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S25474" pageId="25" pageNumber="26" type="EnaNcbi">S 25474</accessionNumber>
</emphasis>
;
<collectionCode id="DBE8AE0BFFE0E47A1845FEA1ED1BC50C" box="[536,560,265,291]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="25" pageNumber="26" type="Herbarium">A</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A1861FEA1ED4CC50C" box="[572,615,265,291]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="25" pageNumber="26" type="Herbarium">BO</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A182EFEA1EDA4C50C" box="[627,655,265,291]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="25" pageNumber="26" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFE0E47A18FDFEA2ED9FC50B" box="[672,692,266,292]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="25" pageNumber="26" type="Herbarium">L</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A189CFEA1EDD4C50C" box="[705,767,265,291]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="25" pageNumber="26" type="Herbarium">SAN</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A1957FEA1EC7AC50C" box="[778,849,265,291]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="25" pageNumber="26" type="Herbarium">SING</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A1900FEA1ECB0C50C" box="[861,923,265,291]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="25" pageNumber="26" type="Herbarium">SAN</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A19F8FEA1EC96C50C" box="[933,957,265,291]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="25" pageNumber="26" type="Herbarium">A</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A1994FEA1ECD9C50C" box="[969,1010,265,291]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="25" pageNumber="26" type="Herbarium">BO</collectionCode>
</materialsCitation>
•
<materialsCitation id="0D913C93FFE0E47A1E57FEA2EA59C566" ID-GBIF-Occurrence="3050815366" accessionNumber="S 78122" collectingDate="1998-06-04" collectionCode="K, KEP, L" collectorName="Kit Pearce, Awg Enjah" location="Ulu Engkari" municipality="Nanga Segera Base Camp" pageId="25" pageNumber="26" specimenCount="1">
<collectingMunicipality id="5D22ACB4FFE0E47A1E57FEA2EA14C50C" box="[1034,1343,265,292]" pageId="25" pageNumber="26">Nanga Segera Base Camp</collectingMunicipality>
,
<location id="B8266015FFE0E47A1F17FEA1EE36C567" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181:B8266015FFE0E47A1F17FEA1EE36C567" municipality="Nanga Segera Base Camp" name="Ulu Engkari" pageId="25" pageNumber="26">Ulu Engkari</location>
,
<location id="B8266015FFE0E47A1B75FE86EECDC567" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181:B8266015FFE0E47A1B75FE86EECDC567" box="[296,486,302,328]" municipality="Nanga Segera Base Camp" name="Lanjak Entimau" pageId="25" pageNumber="26">Lanjak Entimau</location>
,
<location id="B8266015FFE0E47A1BACFE86EDFAC567" LSID="urn:lsid:plazi:treatment:355087D8FFEFE47A1864FC28EA6EC181:B8266015FFE0E47A1BACFE86EDFAC567" box="[497,721,302,328]" municipality="Nanga Segera Base Camp" name="Wildlife Sanctuary" pageId="25" pageNumber="26">Wildlife Sanctuary</location>
;
<date id="C947100EFFE0E47A1881FE86EC4DC567" box="[732,870,302,329]" pageId="25" pageNumber="26" value="1998-06-04">
<collectingDate id="D903E9E6FFE0E47A1881FE86EC4DC567" box="[732,870,302,329]" pageId="25" pageNumber="26" value="1998-06-04">4 Jun. 1998</collectingDate>
</date>
;
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<collectorName id="100C5318FFE0E47A192DFE87EB52C567" box="[880,1145,302,329]" pageId="25" pageNumber="26">Kit Pearce, Awg Enjah</collectorName>
<accessionNumber id="A2AAAB2DFFE0E47A1EDDFE87EBF5C567" box="[1152,1246,302,329]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S78122" pageId="25" pageNumber="26" type="EnaNcbi">S 78122</accessionNumber>
</emphasis>
;
<collectionCode id="DBE8AE0BFFE0E47A1EB3FE86EA21C567" box="[1262,1290,302,328]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="25" pageNumber="26" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFE0E47A1F46FE87EA79C566" box="[1307,1362,303,329]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="25" pageNumber="26" type="Herbarium">KEP</collectionCode>
,
<collectionCode id="DBE8AE0BFFE0E47A1F00FE87EA59C566" box="[1373,1394,303,329]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="25" pageNumber="26" type="Herbarium">L</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="E9866646FFEEE4741AE3F8EAED40C3C6" ID-DOI="http://doi.org/10.5281/zenodo.4594571" ID-Zenodo-Dep="4594571" httpUri="https://zenodo.org/record/4594571/files/figure.png" pageId="23" pageNumber="24" startId="23.[190,233,1858,1884]" targetBox="[224,1317,329,1810]" targetPageId="23">
<paragraph id="BD4636CEFFEEE4741AE3F8EAED40C3C6" blockId="23.[188,1399,1858,2025]" pageId="23" pageNumber="24">
<emphasis id="8F8DEADCFFEEE4741AE3F8EAEE0AC373" bold="true" box="[190,289,1858,1884]" pageId="23" pageNumber="24">Fig. 12.</emphasis>
<taxonomicName id="7AF94D4DFFEEE4741B73F8EAEDB3C372" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[302,664,1858,1885]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="involucrata" status="sp. nov.">
<emphasis id="8F8DEADCFFEEE4741B73F8EAED12C373" box="[302,569,1858,1884]" italics="true" pageId="23" pageNumber="24">Chassalia involucrata</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFEEE47418F8F8EBEC2DC372" box="[677,774,1859,1885]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="8F8DEADCFFEEE474194EF8EAEC03C373" bold="true" box="[787,808,1858,1884]" pageId="23" pageNumber="24">A</emphasis>
. Flowering shoot.
<emphasis id="8F8DEADCFFEEE4741E44F8EAEB06C373" bold="true" box="[1049,1069,1858,1884]" pageId="23" pageNumber="24">B</emphasis>
. Inflorescence.
<emphasis id="8F8DEADCFFEEE4741EAEF8EAEA23C373" bold="true" box="[1267,1288,1858,1884]" pageId="23" pageNumber="24">C</emphasis>
. Flower.
<emphasis id="8F8DEADCFFEEE4741AE3F8CDEFF8C350" bold="true" box="[190,211,1893,1919]" pageId="23" pageNumber="24">D</emphasis>
. Longitudinal view of corolla showing stamens, band of hairs, stigma, style and disc.
<emphasis id="8F8DEADCFFEEE4741EF4F8CDEB96C350" bold="true" box="[1193,1213,1893,1919]" pageId="23" pageNumber="24">E</emphasis>
. Infructescence with stem node showing fresh fruit.
<emphasis id="8F8DEADCFFEEE474182DF821EDAAC38C" bold="true" box="[624,641,1929,1955]" pageId="23" pageNumber="24">F</emphasis>
. Dry fruit.
<emphasis id="8F8DEADCFFEEE4741957F820EC34C38D" bold="true" box="[778,799,1928,1954]" pageId="23" pageNumber="24">G</emphasis>
. Pyrene ventral surface. A, C–D, F–G from
<emphasis id="8F8DEADCFFEEE4741F6FF820EE70C3EA" italics="true" pageId="23" pageNumber="24">Carlo Hansen 1608</emphasis>
; B, E from photographs of living plants in Ulu Temburong National Park, Brunei, taken by Mr Y.W. Low. Drawn by T.Y. Yu.
</paragraph>
</caption>
<caption id="E9866646FFE1E47B1AE0F867EB50C3C5" ID-DOI="http://doi.org/10.5281/zenodo.4594574" ID-Zenodo-Dep="4594574" box="[189,1147,1999,2026]" httpUri="https://zenodo.org/record/4594574/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1999,2025]" targetBox="[267,1321,265,1961]" targetPageId="24">
<paragraph id="BD4636CEFFE1E47B1AE0F867EB50C3C5" blockId="24.[189,1147,1999,2026]" box="[189,1147,1999,2026]" pageId="24" pageNumber="25">
<emphasis id="8F8DEADCFFE1E47B1AE0F867EE32C3C5" bold="true" box="[189,281,1999,2026]" pageId="24" pageNumber="25">Fig. 13.</emphasis>
Holotype of
<taxonomicName id="7AF94D4DFFE1E47B1BE8F878EC3AC3C5" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[437,785,2000,2026]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="involucrata" status="sp. nov.">
<emphasis id="8F8DEADCFFE1E47B1BE8F878ED93C3C5" box="[437,696,2000,2026]" italics="true" pageId="24" pageNumber="25">Chassalia involucrata</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFE1E47B1945F878EC59C3C5" box="[792,882,2000,2026]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="8F8DEADCFFE1E47B19DEF878EB40C3C5" box="[899,1131,2000,2026]" italics="true" pageId="24" pageNumber="25">Carlo Hansen 1608</emphasis>
).
</paragraph>
</caption>
<subSubSection id="F5E36545FFE0E47A1AE0FED9EE0CC116" pageId="25" pageNumber="26" type="description">
<paragraph id="BD4636CEFFE0E47A1AE0FED9EE65C5A4" blockId="25.[189,334,369,395]" box="[189,334,369,395]" pageId="25" pageNumber="26">
<heading id="E60E81A2FFE0E47A1AE0FED9EE65C5A4" bold="true" box="[189,334,369,395]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="8F8DEADCFFE0E47A1AE0FED9EE65C5A4" bold="true" box="[189,334,369,395]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE0E47A1AE0FE0AEE0CC116" blockId="25.[188,1401,418,1337]" pageId="25" pageNumber="26">
Herb, semi-woody shrub to small tree,
<quantity id="7A019B2BFFE0E47A18CCFE0AEDF6C592" box="[657,733,418,445]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">1–4 m</quantity>
tall, glabrous. Stem strong and thick,
<quantity id="7A019B2BFFE0E47A1EF5FE0AEA06C592" box="[1192,1325,418,445]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="25" pageNumber="26" unit="cm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 cm</quantity>
wide, slightly hollowed or not. Internodes
<quantity id="7A019B2BFFE0E47A1827FE6FEDD1C5CD" box="[634,762,455,482]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="13.5" metricValueMin="3.0" pageId="25" pageNumber="26" unit="cm" value="8.25" valueMax="13.5" valueMin="3.0">3–13.5 cm</quantity>
long, thick, sometimes slightly canaliculate, slightly broadening at apex. Stipules big, flabellate, 1–1.5 ×
<quantity id="7A019B2BFFE0E47A1969FE45ECBAC628" box="[820,913,493,519]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
, base usually divided into 2 parts. Leafblades large, broadly elliptic, 13–36 ×
<quantity id="7A019B2BFFE0E47A18C8FDBAEC2AC603" box="[661,769,530,557]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="25" pageNumber="26" unit="cm" value="10.5" valueMax="15.0" valueMin="6.0">6–15 cm</quantity>
, midrib slightly raised both abaxially and adaxially, secondary veins 14–26 on each side, apex acuminate, acumen
<quantity id="7A019B2BFFE0E47A19CEFD9FEB28C67D" box="[915,1027,567,594]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.8" metricValueMin="0.5" pageId="25" pageNumber="26" unit="mm" value="11.5" valueMax="18.0" valueMin="5.0">5–18 mm</quantity>
long, base cuneate or attenuate, petiole long,
<quantity id="7A019B2BFFE0E47A1B0BFDF4EE97C658" box="[342,444,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="14.0" metricValueMin="4.0" pageId="25" pageNumber="26" unit="cm" value="9.0" valueMax="14.0" valueMin="4.0">4–14 cm</quantity>
×
<quantity id="7A019B2BFFE0E47A1B87FDF4ED7FC658" box="[474,596,604,631]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="25" pageNumber="26" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
. Peduncle (5–)20–25 ×
<quantity id="7A019B2BFFE0E47A1937FDF4ECCFC658" box="[874,996,604,631]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="25" pageNumber="26" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
, bracts of rachis and first branches 2, opposite, usually connate to half the length, each bract 1.5–2 ×
<quantity id="7A019B2BFFE0E47A19BCFD2AEB78C6B3" box="[993,1107,642,668]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="25" pageNumber="26" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
, sheath 0.5–0.7 ×
<quantity id="7A019B2BFFE0E47A1F6EFD2AEA59C6B3" box="[1331,1394,642,668]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="cm" value="2.0">2 cm</quantity>
, rachis and secondary branches very short to absent,
<quantity id="7A019B2BFFE0E47A197BFD0EECA2C6EE" box="[806,905,678,705]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="25" pageNumber="26" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 mm</quantity>
long, bracts of secondary branches 2, not sheathing, 12–15 ×
<quantity id="7A019B2BFFE0E47A1BF4FD64ED07C6C9" box="[425,556,716,742]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="25" pageNumber="26" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
, bracts of each capitulum elliptic, 8–12 ×
<quantity id="7A019B2BFFE0E47A1E7FFD64EBBCC6C9" box="[1058,1175,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="25" pageNumber="26" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
. Flowers sessile in each capitulum,
<quantity id="7A019B2BFFE0E47A1BDCFD59EEFEC723" box="[385,469,753,780]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="25" pageNumber="26" unit="mm" value="16.0">16 mm</quantity>
long. Hypanthium 2 ×
<quantity id="7A019B2BFFE0E47A18B3FD59EC67C723" box="[750,844,753,780]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
, calyx tube 0.5 ×
<quantity id="7A019B2BFFE0E47A1E7EFD59EBAAC723" box="[1059,1153,753,780]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="mm" value="1.3">1.3 mm</quantity>
, calyx lobes broadly triangular, 0.2 ×
<quantity id="7A019B2BFFE0E47A1BD9FCBEEEC9C71E" box="[388,482,790,817]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
. Corolla tube 20 ×
<quantity id="7A019B2BFFE0E47A189BFCBEEC25C71E" box="[710,782,790,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="1.0">1 mm</quantity>
, broadening to
<quantity id="7A019B2BFFE0E47A1998FCBEEB22C71E" box="[965,1033,790,817]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="26" unit="mm" value="3.0">3 mm</quantity>
from midway, glabrous except inside corolla tube a band of hairs
<quantity id="7A019B2BFFE0E47A1802FC94ED8EC779" box="[607,677,828,854]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="2.0">2 mm</quantity>
wide at base of stamens,
<quantity id="7A019B2BFFE0E47A1981FC94EB09C779" box="[988,1058,828,854]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="mm" value="7.0">7 mm</quantity>
from mouth of corolla tube, hairs erect, colourless,
<quantity id="7A019B2BFFE0E47A1B91FCC9ED72C754" box="[460,601,865,891]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
long; corolla lobes 5, triangular, 4.5 ×
<quantity id="7A019B2BFFE0E47A1E7DFCC9EB4CC754" box="[1056,1127,865,891]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="2.0">2 mm</quantity>
. Stamens
<quantity id="7A019B2BFFE0E47A1E81FCC8EA1DC754" box="[1244,1334,864,891]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="26" unit="mm" value="2.5">2.5 mm</quantity>
long, slightly concave, base bifurcate, inserted
<quantity id="7A019B2BFFE0E47A18F8FC2EEDC1C78F" box="[677,746,902,928]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="2.0">2 mm</quantity>
from mouth of corolla, filaments very short and broad, less than
<quantity id="7A019B2BFFE0E47A1B6CFC03EEA5C7E9" box="[305,398,939,966]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="0.1">0.1 mm</quantity>
long. Style probably exserted, stigma bilobed, lobes orbiculate, 1 ×
<quantity id="7A019B2BFFE0E47A1E8FFC03EA19C7E9" box="[1234,1330,939,966]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="mm" value="0.8">0.8 mm</quantity>
. Disc cylindric, 1 ×
<quantity id="7A019B2BFFE0E47A1B35FC78EE9AC7C4" box="[360,433,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="1.0">1 mm</quantity>
, apex flat. Fruit sessile, ellipsoid, 7 × 5 ×
<quantity id="7A019B2BFFE0E47A19EDFC78ECD1C7C4" box="[944,1018,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="4.0">4 mm</quantity>
, ripening blue, calyx persistent, not accrescent, 0.5 ×
<quantity id="7A019B2BFFE0E47A1BE6FC5DED32C03F" box="[443,537,1013,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
, disc persistent, not accrescent, 0.7 ×
<quantity id="7A019B2BFFE0E47A1986FC5EEB12C03F" box="[987,1081,1014,1040]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="mm" value="0.7">0.7 mm</quantity>
. Pyrene plano-convex, 6 × 5 ×
<quantity id="7A019B2BFFE0E47A1AB0FBB3EE1EC01A" box="[237,309,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="2.0">2 mm</quantity>
, dorsal surface convex, with two adjacent parallel longitudinal crests,
<quantity id="7A019B2BFFE0E47A1E2EFBB3EBE6C01A" box="[1139,1229,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="0.4">0.4 mm</quantity>
wide,
<quantity id="7A019B2BFFE0E47A1F40FBB3EA53C01A" box="[1309,1400,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="0.4">0.4 mm</quantity>
apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, central part slightly elevated, margin with an endocarpal outgrowth
<quantity id="7A019B2BFFE0E47A1EFCFBCDEA05C0AF" box="[1185,1326,1125,1152]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="25" pageNumber="26" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
wide, with a groove between the edge of endocarp and endocarpal outgrowth
<quantity id="7A019B2BFFE0E47A1E4DFB22EB47C08A" box="[1040,1132,1162,1189]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="0.2">0.2 mm</quantity>
wide. Raphal opening at base of the endocarpal outgrowth, round,
<quantity id="7A019B2BFFE0E47A1893FB18EC02C0E5" box="[718,809,1200,1226]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="0.1">0.1 mm</quantity>
in diameter,
<quantity id="7A019B2BFFE0E47A199AFB07EB08C0E5" box="[967,1059,1199,1226]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
from the edge of the pyrene wall, apex with a bony spine, spine triangular, erect, 0.5 ×
<quantity id="7A019B2BFFE0E47A1934FB7DECEEC0C0" box="[873,965,1237,1263]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="0.2">0.2 mm</quantity>
. PGS not detected. Seed light brown, hemi-ellipsoid, 0.5 ×
<quantity id="7A019B2BFFE0E47A1BE7FB52ED29C13B" box="[442,514,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="5.0">5 mm</quantity>
, longitudinally slightly crested at midline of dorsal surface, endosperm not detected.
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE0E47A1AE0FACAEA6EC181" pageId="25" pageNumber="26" type="distribution">
<paragraph id="BD4636CEFFE0E47A1AE0FACAEE94C153" blockId="25.[189,447,1378,1404]" box="[189,447,1378,1404]" pageId="25" pageNumber="26">
<heading id="E60E81A2FFE0E47A1AE0FACAEE94C153" bold="true" box="[189,447,1378,1404]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="8F8DEADCFFE0E47A1AE0FACAEE94C153" bold="true" box="[189,447,1378,1404]" pageId="25" pageNumber="26">Distribution, habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE0E47A1AE0FA3CEA6EC181" blockId="25.[189,1349,1427,1454]" box="[189,1349,1427,1454]" pageId="25" pageNumber="26">
Borneo:
<collectingCountry id="C5EE765EFFE0E47A1B7EFA3BEEB9C182" box="[291,402,1427,1453]" name="Malaysia" pageId="25" pageNumber="26">Malaysia</collectingCountry>
,
<collectingCountry id="C5EE765EFFE0E47A1BC0FA3BED2EC182" box="[413,517,1427,1453]" name="Malaysia" pageId="25" pageNumber="26">Sarawak</collectingCountry>
,
<collectingRegion id="7F3DF82CFFE0E47A184CFA3BED77C182" box="[529,604,1427,1453]" country="Malaysia" name="Sabah" pageId="25" pageNumber="26">Sabah</collectingRegion>
;
<collectingCountry id="C5EE765EFFE0E47A1835FA3BED97C182" box="[616,700,1427,1453]" name="Brunei" pageId="25" pageNumber="26">Brunei</collectingCountry>
. River banks in lowland forest, alt. 100–350(–1250) m.
</paragraph>
</subSubSection>
</treatment>
</document>
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