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<document id="DF6FA82672CAF8206A8D22DFA4C1CEB8" ID-CLB-Dataset="7783" ID-DOI="10.5852/ejt.2021.738.1261" ID-GBIF-Dataset="02cf5931-b18e-4d58-a6ff-9c7b1e37e68d" ID-ISSN="2118-9773" ID-Zenodo-Dep="4594541" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615410255512" checkinUser="plazi" docAuthor="Turner, Ian M. &amp; Cheek, Martin" docDate="2021" docId="355087D8FFD3E44E181DFEA0EC27C559" docLanguage="en" docName="ejt-738_1-60.pdf" docOrigin="European Journal of Taxonomy 738" docSource="http://dx.doi.org/10.5852/ejt.2021.738.1261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chassalia muluensis T. Y. Yu 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="C969FFA0FFF9E4631A5DFFA8EF2BC42F" masterDocTitle="Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species" masterLastPageNumber="60" masterPageNumber="1" pageNumber="43" updateTime="1698932934732" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2A95C32A005C0E42F3F2D27B4BA13C2B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84C4F9CE2261DA5075071E9F5762D6C4">
<mods:title id="6768B58EA761EF45C86B83ACD8967772">Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species</mods:title>
</mods:titleInfo>
<mods:name id="06A51C9857A26A290180EB0C553AE455" type="personal">
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<mods:namePart id="AD56F45E5914B77ED315ED3B2C2697A4">Turner, Ian M.</mods:namePart>
<mods:nameIdentifier id="98CA30818CF84EDB0ADA9150DC4E1C57" type="email">i.turner@kew.org</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="2D619FA3FE8902AC2EFF681291519A12">Cheek, Martin</mods:namePart>
<mods:affiliation id="EC2E67941F3F5A222EBD92A89D066693">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Queen Mary University of London, E 1 4 NS, UK. Singapore Botanical Liaison Officer, Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569, Singapore.</mods:affiliation>
<mods:nameIdentifier id="C99261A6851B9D4FEEF1B2318283598F" type="email">m.cheek@kew.org</mods:nameIdentifier>
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<mods:typeOfResource id="443B481B85F003192892064691A12ACF">text</mods:typeOfResource>
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<mods:title id="3A6C4D12C93C08C1461EC44A5C456C36">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="F5F05DA4D09EC320BF025EA3668DEF76">
<mods:date id="FE5930594C1B606ABE87E99FA061C327">2021</mods:date>
<mods:detail id="C9CBD9CD002EF142D4387F4CBE461472" type="pubDate">
<mods:number id="5A2C097B6025BF32C9FAE924C5E5286D">2021-03-09</mods:number>
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<mods:detail id="A41D656D2A82F6450E1FAE155DD17CE2" type="volume">
<mods:number id="79455DDDC504FAB34240DB0FAE18DFF1">738</mods:number>
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<mods:start id="765FF9D2C17F9134080BE5713F772BC0">1</mods:start>
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<mods:url id="67948EDB9E36E4A6E8A5D83B21BA355D">http://dx.doi.org/10.5852/ejt.2021.738.1261</mods:url>
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<mods:classification id="CBFEF8EC7969D850EFD5D690C91FB625">journal article</mods:classification>
<mods:identifier id="E7EBC03FAB71D53C069D6F0A96D73444" type="CLB-Dataset">7783</mods:identifier>
<mods:identifier id="0500C6AEB9C01B733063CD3EAD843288" type="DOI">10.5852/ejt.2021.738.1261</mods:identifier>
<mods:identifier id="E7B9832BE882E8253CA13D8CDF3C6E14" type="GBIF-Dataset">02cf5931-b18e-4d58-a6ff-9c7b1e37e68d</mods:identifier>
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<treatment id="355087D8FFD3E44E181DFEA0EC27C559" ID-DOI="http://doi.org/10.5281/zenodo.4594529" ID-GBIF-Taxon="180327063" ID-Zenodo-Dep="4594529" LSID="urn:lsid:plazi:treatment:355087D8FFD3E44E181DFEA0EC27C559" httpUri="http://treatment.plazi.org/id/355087D8FFD3E44E181DFEA0EC27C559" lastPageId="45" lastPageNumber="46" pageId="42" pageNumber="43">
<subSubSection id="F5E36545FFD3E449181DFEA0ECDFC50C" box="[576,1012,264,291]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="BD4636CEFFD3E449181DFEA0ECDFC50C" blockId="42.[576,1012,264,364]" box="[576,1012,264,291]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E449181DFEA0ECDFC50C" box="[576,1012,264,291]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="43" reason="2">
<taxonomicName id="7AF94D4DFFD3E449181DFEA0ECB9C50C" ID-CoL="9V47W" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[576,914,264,291]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFD3E449181DFEA0EC11C50C" bold="true" box="[576,826,264,291]" italics="true" pageId="42" pageNumber="43">Chassalia muluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFD3E44919C4FEA1ECDFC50C" box="[921,1012,265,291]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFD3E449181AFE85EC70C543" pageId="42" pageNumber="43" type="description">
<paragraph id="BD4636CEFFD3E449181AFE85ECC6C567" blockId="42.[576,1012,264,364]" box="[583,1005,301,328]" pageId="42" pageNumber="43">urn:lsid:ipni.org:names:77215412-1</paragraph>
<paragraph id="BD4636CEFFD3E4491885FEFAEC70C543" blockId="42.[576,1012,264,364]" box="[728,859,337,364]" pageId="42" pageNumber="43">
<figureCitation id="25C22A4BFFD3E4491885FEFAEC70C543" box="[728,859,337,364]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="43.[189,232,1893,1919]" captionStartId-1="44.[189,232,1999,2025]" captionTargetBox-0="[281,1269,340,1807]" captionTargetBox-1="[271,1328,300,1960]" captionTargetId-0="figure-12@43.[189,1398,262,1857]" captionTargetId-1="figure-14@44.[260,1328,265,1960]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="Fig. 24. Chassalia muluensis T.Y.Yu sp. nov. A. Flowering shoot. B. Flower. C. Longitudinal view of corolla tube showing stamens, band of hairs, style and disc. D. Hypanthium, calyx and disc of a flower. E. Pyrene ventral surface. F. Dry fruit transection. G. Part of infructescence, showing dry fruit. From G. Argent, B. Coppins 1134. Drawn by T.Y. Yu." captionText-1="Fig. 25. Holotype of Chassalia muluensis T.Y.Yu sp. nov. (G. Argent, B. Coppins 1134)." figureDoi-0="http://doi.org/10.5281/zenodo.4594598" figureDoi-1="http://doi.org/10.5281/zenodo.4594600" httpUri-0="https://zenodo.org/record/4594598/files/figure.png" httpUri-1="https://zenodo.org/record/4594600/files/figure.png" pageId="42" pageNumber="43">Figs 24–25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFD3E4491AE0FE3CEE4CC664" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="BD4636CEFFD3E4491AE0FE3CEE1DC581" blockId="42.[189,310,404,430]" box="[189,310,404,430]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E4491AE0FE3CEE1DC581" bold="true" box="[189,310,404,430]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8F8DEADCFFD3E4491AE0FE3CEE1DC581" bold="true" box="[189,310,404,430]" pageId="42" pageNumber="43">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FE6CEE4CC664" blockId="42.[188,1399,452,587]" pageId="42" pageNumber="43">
Differs from other Bornean species of
<taxonomicName id="7AF94D4DFFD3E4491826FE6CEDDBC5F1" box="[635,752,452,478]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8F8DEADCFFD3E4491826FE6CEDDBC5F1" box="[635,752,452,478]" italics="true" pageId="42" pageNumber="43">Chassalia</emphasis>
</taxonomicName>
by having secondary nerves densely distributed, 16–18 on each side of the midrib; stipule broadly flabellate, 5–8 ×
<quantity id="7A019B2BFFD3E44919D3FE40EB29C62C" box="[910,1026,488,515]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="42" pageNumber="43" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
; inflorescence
<quantity id="7A019B2BFFD3E4491EE9FE40EA19C62C" box="[1204,1330,488,515]" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.4" metricValueMin="0.95" pageId="42" pageNumber="43" unit="cm" value="11.75" valueMax="14.0" valueMin="9.5">9.5–14 cm</quantity>
long, longer than the average leaf, but usually shorter than the longest leaf; only found in Gunung Mulu National Park.
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFD3E4491AE0FDDBEBB2C692" box="[189,1177,627,702]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="BD4636CEFFD3E4491AE0FDDBEE6FC6A2" blockId="42.[189,324,627,653]" box="[189,324,627,653]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E4491AE0FDDBEE6FC6A2" bold="true" box="[189,324,627,653]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8F8DEADCFFD3E4491AE0FDDBEE6FC6A2" bold="true" box="[189,324,627,653]" pageId="42" pageNumber="43">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FD0BEBB2C692" blockId="42.[189,1177,675,702]" box="[189,1177,675,702]" pageId="42" pageNumber="43">The epithet alludes to Gunung Mulu National Park to which this species is endemic.</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFD3E4491AE0FD4EEA0BC01C" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="BD4636CEFFD3E4491AE0FD4EEFD1C72F" blockId="42.[189,250,742,768]" box="[189,250,742,768]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E4491AE0FD4EEFD1C72F" bold="true" box="[189,250,742,768]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8F8DEADCFFD3E4491AE0FD4EEFD1C72F" bold="true" box="[189,250,742,768]" pageId="42" pageNumber="43">
<typeStatus id="6242886CFFD3E4491AE0FD4EEFD1C72F" box="[189,250,742,768]" pageId="42" pageNumber="43">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FCBDEDF7C757" blockId="42.[188,1399,788,888]" pageId="42" pageNumber="43">
<materialsCitation id="0D913C93FFD3E4491AE0FCBDEDF3C757" ID-GBIF-Occurrence="3050815305" collectingDate="1978-04-29" collectionCode="G, B, K, KEP" country="Malaysia" elevation="1600" latitude="4.05" location="Camp" longLatPrecision="1307" longitude="114.88333" municipality="Gunung Mulu National Park" pageId="42" pageNumber="43" specimenCode="K001129717, KEP271647" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<collectingCountry id="C5EE765EFFD3E4491AE0FCBDEE7FC700" box="[189,340,789,815]" name="Malaysia" pageId="42" pageNumber="43">MALAYSIA</collectingCountry>
–
<emphasis id="8F8DEADCFFD3E4491B31FCBDED6FC700" bold="true" box="[364,580,789,815]" pageId="42" pageNumber="43">
Borneo,
<collectingRegion id="7F3DF82CFFD3E4491B88FCBDED6FC700" box="[469,580,789,815]" country="Malaysia" name="Sarawak" pageId="42" pageNumber="43">Sarawak</collectingRegion>
</emphasis>
•
<collectingMunicipality id="5D22ACB4FFD3E4491806FCBDEC83C700" box="[603,936,789,815]" pageId="42" pageNumber="43">Gunung Mulu National Park</collectingMunicipality>
, west ridge just below
<location id="B8266015FFD3E4491EF3FCBDEBDFC700" LSID="urn:lsid:plazi:treatment:355087D8FFD3E44E181DFEA0EC27C559:B8266015FFD3E4491EF3FCBDEBDFC700" box="[1198,1268,789,815]" country="Malaysia" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Gunung Mulu National Park" name="Camp" pageId="42" pageNumber="43" stateProvince="Sarawak">Camp</location>
4;
<geoCoordinate id="D8CD5009FFD3E4491F4BFCBCEA59C71F" box="[1302,1394,788,816]" degrees="4" direction="north" minutes="03" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="4.05">4°03′ N</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFD3E4491AE0FC90EE19C77B" box="[189,306,824,852]" degrees="114" direction="east" minutes="53" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="114.88333">114°53′ E</geoCoordinate>
; alt.
<quantity id="7A019B2BFFD3E4491B35FC91EEE9C77B" box="[360,450,825,852]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="42" pageNumber="43" unit="m" value="1600.0">
<elevation id="36D4D1FDFFD3E4491B35FC91EEE9C77B" box="[360,450,825,852]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="42" pageNumber="43" unit="m" value="1600.0">1600 m</elevation>
</quantity>
; submontane ridge forest;
<date id="C947100EFFD3E44918AEFC92ECA7C77B" box="[755,908,826,852]" pageId="42" pageNumber="43" value="1978-04-29">
<collectingDate id="D903E9E6FFD3E44918AEFC92ECA7C77B" box="[755,908,826,852]" pageId="42" pageNumber="43" value="1978-04-29">29 Apr. 1978</collectingDate>
</date>
; fl.;
<emphasis id="8F8DEADCFFD3E44919E0FC92EBD2C77B" box="[957,1273,826,852]" italics="true" pageId="42" pageNumber="43">
<collectionCode id="DBE8AE0BFFD3E44919E0FC92ECFEC77B" box="[957,981,826,852]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="42" pageNumber="43" type="Herbarium">G</collectionCode>
. Argent,
<collectionCode id="DBE8AE0BFFD3E4491E67FC92EB64C77C" box="[1082,1103,826,851]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="42" pageNumber="43" type="Herbarium">B</collectionCode>
. Coppins 1134
</emphasis>
;
<typeStatus id="6242886CFFD3E4491F55FC91EA58C77C" box="[1288,1395,825,851]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
:
<collectionCode id="DBE8AE0BFFD3E4491AE1FCF5EFF9C757" box="[188,210,861,888]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="42" pageNumber="43" type="Herbarium">K</collectionCode>
[
<specimenCode id="ED5F9EB5FFD3E4491A84FCF5EE54C757" box="[217,383,861,888]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="42" pageNumber="43" type="Herbarium">K001129717</specimenCode>
]!; isotype:
<collectionCode id="DBE8AE0BFFD3E4491BA7FCF5ED1FC757" box="[506,564,861,888]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="42" pageNumber="43" type="Herbarium">KEP</collectionCode>
[
<specimenCode id="ED5F9EB5FFD3E4491866FCF5EDFAC757" box="[571,721,861,888]" collectionCode="KEP" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="42" pageNumber="43" type="Herbarium">KEP271647</specimenCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FC08EE9FC795" blockId="42.[189,436,928,954]" box="[189,436,928,954]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E4491AE0FC08EE9FC795" bold="true" box="[189,436,928,954]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8F8DEADCFFD3E4491AE0FC08EE9FC795" bold="true" box="[189,436,928,954]" pageId="42" pageNumber="43">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FC78EA0BC01C" blockId="42.[189,1399,974,1075]" pageId="42" pageNumber="43">
<materialsCitation id="0D913C93FFD3E4491AE0FC78ECFCC020" ID-GBIF-Occurrence="3050815355" collectingDate="1978-03-25" collectionCode="I, K, AAU" country="Malaysia" elevation="1850" latitude="4.0833335" location="Camp" longLatPrecision="1307" longitude="114.916664" municipality="Gunong Mulu National Park" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Sarawak">
<collectingCountry id="C5EE765EFFD3E4491AE0FC78EE7FC7C5" box="[189,340,976,1002]" name="Malaysia" pageId="42" pageNumber="43">MALAYSIA</collectingCountry>
–
<emphasis id="8F8DEADCFFD3E4491B27FC78ED73C7C5" bold="true" box="[378,600,976,1002]" pageId="42" pageNumber="43">
Borneo,
<collectingRegion id="7F3DF82CFFD3E4491BB4FC78ED73C7C5" box="[489,600,976,1002]" country="Malaysia" name="Sarawak" pageId="42" pageNumber="43">Sarawak</collectingRegion>
</emphasis>
•
<collectingMunicipality id="5D22ACB4FFD3E4491826FC78ECF0C7C5" box="[635,987,976,1002]" pageId="42" pageNumber="43">Gunong Mulu National Park</collectingMunicipality>
, ridge west of
<location id="B8266015FFD3E4491EC7FC78EBCBC7C5" LSID="urn:lsid:plazi:treatment:355087D8FFD3E44E181DFEA0EC27C559:B8266015FFD3E4491EC7FC78EBCBC7C5" box="[1178,1248,976,1002]" country="Malaysia" latitude="4.0833335" longLatPrecision="1307" longitude="114.916664" municipality="Gunong Mulu National Park" name="Camp" pageId="42" pageNumber="43" stateProvince="Sarawak">Camp</location>
4;
<geoCoordinate id="D8CD5009FFD3E4491F4DFC66EA59C7C4" box="[1296,1394,974,1003]" degrees="4" direction="north" minutes="05" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="4.0833335">4°05′ N</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFD3E4491AE0FC5BEE1EC020" box="[189,309,1011,1039]" degrees="114" direction="east" minutes="55" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="114.916664">114°55′ E</geoCoordinate>
; alt.
<quantity id="7A019B2BFFD3E4491B2DFC5CEEE6C020" box="[368,461,1012,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="42" pageNumber="43" unit="m" value="1850.0">
<elevation id="36D4D1FDFFD3E4491B2DFC5CEEE6C020" box="[368,461,1012,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="42" pageNumber="43" unit="m" value="1850.0">1850 m</elevation>
</quantity>
;
<date id="C947100EFFD3E4491B86FC5CED55C020" box="[475,638,1012,1039]" pageId="42" pageNumber="43" value="1978-03-25">
<collectingDate id="D903E9E6FFD3E4491B86FC5CED55C020" box="[475,638,1012,1039]" pageId="42" pageNumber="43" value="1978-03-25">25 Mar. 1978</collectingDate>
</date>
; fl.;
<emphasis id="8F8DEADCFFD3E44918EEFC5DEC71C021" box="[691,858,1012,1038]" italics="true" pageId="42" pageNumber="43">
<collectionCode id="DBE8AE0BFFD3E44918EEFC5DEDEBC021" box="[691,704,1013,1038]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="42" pageNumber="43" type="Herbarium">I</collectionCode>
. Nielsen 856
</emphasis>
;
<collectionCode id="DBE8AE0BFFD3E4491937FC5DECADC020" box="[874,902,1013,1039]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="42" pageNumber="43" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFD3E44919CAFC5DECFCC020" box="[919,983,1013,1039]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="42" pageNumber="43">AAU</collectionCode>
</materialsCitation>
•
<materialsCitation id="0D913C93FFD3E44919AFFC5DEA3DC01C" ID-GBIF-Occurrence="3050815371" collectingDate="1978-03-22" collectionCode="K" country="Malaysia" elevation="1800" latitude="4.0833335" location="Camp" longLatPrecision="1307" longitude="114.9" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Borneo">
<location id="B8266015FFD3E44919AFFC5DEB13C020" LSID="urn:lsid:plazi:treatment:355087D8FFD3E44E181DFEA0EC27C559:B8266015FFD3E44919AFFC5DEB13C020" box="[1010,1080,1013,1039]" country="Malaysia" latitude="4.0833335" longLatPrecision="1307" longitude="114.9" name="Camp" pageId="42" pageNumber="43" stateProvince="Borneo">Camp</location>
4;
<geoCoordinate id="D8CD5009FFD3E4491E3DFC5BEBFCC020" box="[1120,1239,1011,1039]" degrees="114" direction="east" minutes="54" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="114.9">114°54′ E</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFD3E4491EB8FC5BEA6FC020" box="[1253,1348,1011,1039]" degrees="4" direction="north" minutes="05" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="4.0833335">4°05′ N</geoCoordinate>
; alt.
<quantity id="7A019B2BFFD3E4491AE0FBB1EE32C01C" box="[189,281,1049,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="43" unit="m" value="1800.0">
<elevation id="36D4D1FDFFD3E4491AE0FBB1EE32C01C" box="[189,281,1049,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="43" unit="m" value="1800.0">1800 m</elevation>
</quantity>
; slope in moss forest in upper montane forest;
<date id="C947100EFFD3E4491960FBB1ECF5C01C" box="[829,990,1049,1075]" pageId="42" pageNumber="43" value="1978-03-22">
<collectingDate id="D903E9E6FFD3E4491960FBB1ECF5C01C" box="[829,990,1049,1075]" pageId="42" pageNumber="43" value="1978-03-22">22 Mar. 1978</collectingDate>
</date>
; fl.;
<emphasis id="8F8DEADCFFD3E4491E4CFBB1EBC1C01C" box="[1041,1258,1049,1075]" italics="true" pageId="42" pageNumber="43">Carlo Hansen 544</emphasis>
;
<collectionCode id="DBE8AE0BFFD3E4491EA4FBB1EA3DC01C" box="[1273,1302,1049,1075]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="42" pageNumber="43" type="Herbarium">K</collectionCode>
</materialsCitation>
!.
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFD3E4491AE0FBF3EEC4C3C6" pageId="42" pageNumber="43" type="description">
<paragraph id="BD4636CEFFD3E4491AE0FBF3EE65C05A" blockId="42.[189,334,1115,1141]" box="[189,334,1115,1141]" pageId="42" pageNumber="43">
<heading id="E60E81A2FFD3E4491AE0FBF3EE65C05A" bold="true" box="[189,334,1115,1141]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8F8DEADCFFD3E4491AE0FBF3EE65C05A" bold="true" box="[189,334,1115,1141]" pageId="42" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD3E4491AE0FB23EEC4C3C6" blockId="42.[187,1401,1163,2025]" pageId="42" pageNumber="43">
Subshrub, shrub or sub-woody epiphyte,
<quantity id="7A019B2BFFD3E44918C5FB23EC2DC08A" box="[664,774,1163,1190]" metricMagnitude="0" metricUnit="m" metricValue="1.375" metricValueMax="2.0" metricValueMin="0.75" pageId="42" pageNumber="43" unit="m" value="1.375" valueMax="2.0" valueMin="0.75">0.75–2 m</quantity>
high, glabrous. Stem terete,
<quantity id="7A019B2BFFD3E4491E12FB23EB84C08A" box="[1103,1199,1163,1189]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
wide. Internodes of the flowering branch
<quantity id="7A019B2BFFD3E4491B81FB07ED1FC0E5" box="[476,564,1199,1226]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="42" pageNumber="43" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long, lower internode terete, upper internode usually drying flattened, canaliculate, broadening at apex, base
<quantity id="7A019B2BFFD3E44918D5FB7CEDC1C0C1" box="[648,746,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="42" pageNumber="43" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
wide, apex
<quantity id="7A019B2BFFD3E4491921FB7BECF5C0C1" box="[892,990,1235,1262]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
wide. Leaves opposite, decussate. Leaf-blades elliptic, 6.5–14.5 ×
<quantity id="7A019B2BFFD3E4491817FB50EDEBC13D" box="[586,704,1272,1299]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.8" metricValueMin="3.0" pageId="42" pageNumber="43" unit="cm" value="4.4" valueMax="5.8" valueMin="3.0">3–5.8 cm</quantity>
; apex acute to acuminate, acumen
<quantity id="7A019B2BFFD3E4491E25FB50EBC6C13D" box="[1144,1261,1272,1299]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="42" pageNumber="43" unit="mm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 mm</quantity>
long; base attenuate; midrib slightly raised above, domatia absent; secondary nerves 16–18 on each side of the midrib, prominent above; tertiary nerves not visible. Petiole flattened, canaliculate,
<quantity id="7A019B2BFFD3E4491ECBFAE9EBC6C174" box="[1174,1261,1345,1371]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
×
<quantity id="7A019B2BFFD3E4491F53FAE9EA59C174" box="[1294,1394,1345,1371]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="42" pageNumber="43" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
. Stipule not sheathing, flabellate, 5–8 ×
<quantity id="7A019B2BFFD3E44918CCFACDEC2DC150" box="[657,774,1381,1408]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="42" pageNumber="43" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
, apex breaking very easily, base drying deep brown, margin light brown. Inflorescence a compound cyme,
<quantity id="7A019B2BFFD3E4491961FA21EC92C18B" box="[828,953,1417,1444]" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.4" metricValueMin="0.95" pageId="42" pageNumber="43" unit="cm" value="11.75" valueMax="14.0" valueMin="9.5">9.5–14 cm</quantity>
long, peduncle
<quantity id="7A019B2BFFD3E4491E2AFA21EBCEC18B" box="[1143,1253,1417,1444]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="42" pageNumber="43" unit="cm" value="7.25" valueMax="8.0" valueMin="6.5">6.5–8 cm</quantity>
long, bracts of first internode of rachis narrowly elliptic, 2–4, 9–12 ×
<quantity id="7A019B2BFFD3E4491927FA05ECCAC1E7" box="[890,993,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, first internode of rachis and first branches
<quantity id="7A019B2BFFD3E4491B73FA7AEEB7C1C3" box="[302,412,1490,1517]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="42" pageNumber="43" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
long, bracts of second internode of rachis and second branches 4, ovate to elliptic, 10 ×
<quantity id="7A019B2BFFD3E4491AA6FA5EEE68C23E" box="[251,323,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="5.0">5 mm</quantity>
, bracts of capitulum 5–6, narrowly elliptic, 5–10 ×
<quantity id="7A019B2BFFD3E44919FDFA5EEB2EC23E" box="[928,1029,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="42" pageNumber="43" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
, second internode of rachis and second branches
<quantity id="7A019B2BFFD3E4491BDBF9B2EEDDC21A" box="[390,502,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="42" pageNumber="43" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long or absent. Flowers
<quantity id="7A019B2BFFD3E4491947F9B3ECA9C21B" box="[794,898,1562,1589]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.572" metricValueMin="3.048" pageId="42" pageNumber="43" unit="in" value="15.0" valueMax="18.0" valueMin="12.0">12–18 in</quantity>
each capitulum, hypanthium 2 ×
<quantity id="7A019B2BFFD3E4491F53F9B3EA59C21A" box="[1294,1394,1563,1589]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="42" pageNumber="43" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, calyx tube 1–1.2 ×
<quantity id="7A019B2BFFD3E4491BFDF997EED5C276" box="[416,510,1599,1626]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="mm" value="1.5">1.5 mm</quantity>
, lobes of calyx tube 0.3 ×
<quantity id="7A019B2BFFD3E4491968F997ECEDC276" box="[821,966,1599,1626]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
, corolla tube 8–12 ×
<quantity id="7A019B2BFFD3E4491E9DF997EA0EC276" box="[1216,1317,1599,1625]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="42" pageNumber="43" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, inside corolla tube a band of hairs
<quantity id="7A019B2BFFD3E4491850F9CBED43C251" box="[525,616,1635,1662]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="0.5">0.5 mm</quantity>
wide at base of stamens,
<quantity id="7A019B2BFFD3E44919C0F9CBECC9C251" box="[925,994,1635,1662]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="mm" value="3.0">3 mm</quantity>
from mouth of corolla tube, hairs erect, colourless,
<quantity id="7A019B2BFFD3E4491BD3F920ED36C28D" box="[398,541,1672,1698]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="42" pageNumber="43" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
long; corolla lobes 2 ×
<quantity id="7A019B2BFFD3E4491962F920EC8DC28D" box="[831,934,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="42" pageNumber="43" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, stamens 2.5 ×
<quantity id="7A019B2BFFD3E4491E3FF92FEBEAC28D" box="[1122,1217,1671,1698]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="0.5">0.5 mm</quantity>
, filament 0.5 ×
<quantity id="7A019B2BFFD3E4491AE0F904EE32C2E9" box="[189,281,1708,1734]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="mm" value="0.1">0.1 mm</quantity>
, style ca 1.5 times as long as corolla,
<quantity id="7A019B2BFFD3E4491891F904EC30C2E9" box="[716,795,1708,1734]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="43" unit="cm" value="1.8">1.8 cm</quantity>
long. Disc cylindric, 0.7 ×
<quantity id="7A019B2BFFD3E4491E05F904EB9EC2E9" box="[1112,1205,1708,1734]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="42" pageNumber="43" unit="mm" value="0.9">0.9 mm</quantity>
. Fruits ellipsoid, dark purple when ripe, 7.5–8 × 4–5.5 ×
<quantity id="7A019B2BFFD3E44918CAF978EC38C2C4" box="[663,787,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="42" pageNumber="43" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4–5.5 mm</quantity>
, pyrenes plano-convex, dorsal surface convex, with two adjacent parallel longitudinal crests, each
<quantity id="7A019B2BFFD3E44918B5F95DEC68C320" box="[744,835,1781,1807]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="mm" value="0.2">0.2 mm</quantity>
wide, curved,
<quantity id="7A019B2BFFD3E44919ABF95DEB7AC320" box="[1014,1105,1781,1807]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="mm" value="0.2">0.2 mm</quantity>
apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, centre slightly elevated, margin with an endocarpal outgrowth
<quantity id="7A019B2BFFD3E449193CF895ECDAC377" box="[865,1009,1853,1880]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="42" pageNumber="43" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
wide and a groove between the edge of endocarp and endocarpal outgrowth
<quantity id="7A019B2BFFD3E449189CF8C9EC66C353" box="[705,845,1889,1916]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="42" pageNumber="43" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
wide. Raphal opening at base of the endocarpal outgrowth, round,
<quantity id="7A019B2BFFD3E4491BCAF82EEEDAC38F" box="[407,497,1926,1952]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="mm" value="0.3">0.3 mm</quantity>
diameter,
<quantity id="7A019B2BFFD3E4491830F82EEDECC38F" box="[621,711,1926,1952]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="mm" value="0.2">0.2 mm</quantity>
from the edge of the pyrene wall, apex with a bony spine, erect, 1 ×
<quantity id="7A019B2BFFD3E4491B67F802EEB1C3EA" box="[314,410,1962,1989]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="mm" value="0.2">0.2 mm</quantity>
. PGS not detected. Seeds brown, hemi-ellipsoid, slightly crested at central part of dorsal side, 3.2 ×
<quantity id="7A019B2BFFD3E4491BD3F866EEC0C3C6" box="[398,491,1998,2025]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="42" pageNumber="43" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="E9866646FFD2E4481AE0F8CDEDC3C3C6" ID-DOI="http://doi.org/10.5281/zenodo.4594598" ID-Zenodo-Dep="4594598" httpUri="https://zenodo.org/record/4594598/files/figure.png" pageId="43" pageNumber="44" startId="43.[189,232,1893,1919]" targetBox="[281,1269,340,1807]" targetPageId="43">
<paragraph id="BD4636CEFFD2E4481AE0F8CDEDC3C3C6" blockId="43.[189,1399,1893,2025]" pageId="43" pageNumber="44">
<emphasis id="8F8DEADCFFD2E4481AE0F8CDEE30C350" bold="true" box="[189,283,1893,1919]" pageId="43" pageNumber="44">Fig. 24.</emphasis>
<taxonomicName id="7AF94D4DFFD2E4481B7EF8CDED5BC3AF" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[291,624,1893,1920]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFD2E4481B7EF8CDED3DC350" box="[291,534,1893,1919]" italics="true" pageId="43" pageNumber="44">Chassalia muluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFD2E4481825F8CEEDFFC3AF" box="[632,724,1894,1920]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="8F8DEADCFFD2E4481880F8CDEDDAC350" bold="true" box="[733,753,1893,1919]" pageId="43" pageNumber="44">A</emphasis>
. Flowering shoot.
<emphasis id="8F8DEADCFFD2E448198FF8CDECCDC350" bold="true" box="[978,998,1893,1919]" pageId="43" pageNumber="44">B</emphasis>
. Flower.
<emphasis id="8F8DEADCFFD2E4481E0AF8CDEB47C350" bold="true" box="[1111,1132,1893,1919]" pageId="43" pageNumber="44">C</emphasis>
. Longitudinal view of corolla tube showing stamens, band of hairs, style and disc.
<emphasis id="8F8DEADCFFD2E4481920F821ECB9C38C" bold="true" box="[893,914,1929,1955]" pageId="43" pageNumber="44">D</emphasis>
. Hypanthium, calyx and disc of a flower.
<emphasis id="8F8DEADCFFD2E4481AE0F804EFFAC3E9" bold="true" box="[189,209,1964,1990]" pageId="43" pageNumber="44">E</emphasis>
. Pyrene ventral surface.
<emphasis id="8F8DEADCFFD2E4481BA6F804ED20C3E9" bold="true" box="[507,523,1964,1990]" pageId="43" pageNumber="44">F</emphasis>
. Dry fruit transection.
<emphasis id="8F8DEADCFFD2E448197DF803EC1EC3EA" bold="true" box="[800,821,1963,1989]" pageId="43" pageNumber="44">G</emphasis>
. Part of infructescence, showing dry fruit. From
<emphasis id="8F8DEADCFFD2E4481AE0F867EED4C3C6" box="[189,511,1999,2025]" italics="true" pageId="43" pageNumber="44">G. Argent, B. Coppins 1134</emphasis>
. Drawn by T.Y. Yu.
</paragraph>
</caption>
<caption id="E9866646FFD5E44F1AE0F867EBEEC3C5" ID-DOI="http://doi.org/10.5281/zenodo.4594600" ID-Zenodo-Dep="4594600" box="[189,1221,1999,2026]" httpUri="https://zenodo.org/record/4594600/files/figure.png" pageId="44" pageNumber="45" startId="44.[189,232,1999,2025]" targetBox="[271,1328,300,1960]" targetPageId="44">
<paragraph id="BD4636CEFFD5E44F1AE0F867EBEEC3C5" blockId="44.[189,1221,1999,2026]" box="[189,1221,1999,2026]" pageId="44" pageNumber="45">
<emphasis id="8F8DEADCFFD5E44F1AE0F867EE31C3C6" bold="true" box="[189,282,1999,2025]" pageId="44" pageNumber="45">Fig. 25.</emphasis>
Holotype of
<taxonomicName id="7AF94D4DFFD5E44F1BE8F867EDD4C3C5" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[437,767,1999,2026]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFD5E44F1BE8F867ED8DC3C6" box="[437,678,1999,2025]" italics="true" pageId="44" pageNumber="45">Chassalia muluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFD5E44F195BF878EC4BC3C5" box="[774,864,2000,2026]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="8F8DEADCFFD5E44F192CF878EB98C3C6" box="[881,1203,1999,2026]" italics="true" pageId="44" pageNumber="45">G. Argent, B. Coppins 1134</emphasis>
).
</paragraph>
</caption>
<subSubSection id="F5E36545FFD4E44E1AE0FEA0EC27C559" pageId="45" pageNumber="46" type="distribution">
<paragraph id="BD4636CEFFD4E44E1AE0FEA0EE94C50D" blockId="45.[189,447,264,290]" box="[189,447,264,290]" pageId="45" pageNumber="46">
<heading id="E60E81A2FFD4E44E1AE0FEA0EE94C50D" bold="true" box="[189,447,264,290]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="8F8DEADCFFD4E44E1AE0FEA0EE94C50D" bold="true" box="[189,447,264,290]" pageId="45" pageNumber="46">Distribution, habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFD4E44E1AE0FE90EC27C559" blockId="45.[188,1399,311,374]" pageId="45" pageNumber="46">
Borneo:
<collectingCountry id="C5EE765EFFD4E44E1B7FFE9FEEBAC57E" box="[290,401,311,337]" name="Malaysia" pageId="45" pageNumber="46">Malaysia</collectingCountry>
,
<collectingCountry id="C5EE765EFFD4E44E1BC6FE9FED28C57E" box="[411,515,311,337]" name="Malaysia" pageId="45" pageNumber="46">Sarawak</collectingCountry>
, Gunung Mulu National Park, around Camp 4. Slope in moss forest in upper sub-montane to montane forest, alt.
<quantity id="7A019B2BFFD4E44E183EFEF3EC23C559" box="[611,776,347,374]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.85" metricValueMin="1.6" pageId="45" pageNumber="46" unit="m" value="1725.0" valueMax="1850.0" valueMin="1600.0">1600–1850 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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