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<document id="4DF56AD294B34A1E45658C99EBB83D23" ID-DOI="10.11646/zootaxa.5023.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5226112" ID-ZooBank="B08B0A40-66BE-4249-99DB-9BAB750F76DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629427898826" checkinUser="plazi" docAuthor="Yepishin, Viktor, Khalaim, Yevhenii, Budashkin, Yuriy, Zhakov, Oleksandr, Mushynskyi, Vadym &amp; Novytskyi, Sergiy" docDate="2021" docId="03BA87F9E91825650EF7FBCC58C98CF5" docLanguage="en" docName="zootaxa.5023.3.3.pdf" docOrigin="Zootaxa 5023 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bradyrrhoa imperialella" docType="treatment" docVersion="3" lastPageNumber="372" masterDocId="FF83FF81E91D25630E60FF9B5C2A8D24" masterDocTitle="New records of pyraloid moths (Lepidoptera: Pyraloidea) from different regions of Ukraine" masterLastPageNumber="388" masterPageNumber="366" pageNumber="371" updateTime="1699072783881" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DCFA5369CA5B2A639B548ADBCAF16B92">New records of pyraloid moths (Lepidoptera: Pyraloidea) from different regions of Ukraine</mods:title>
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<mods:name id="B5074FBEAB022489BA23871ECF815789" type="personal">
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<mods:namePart id="061F119AEFC5B94D103D3A32E1FC8B15">Yepishin, Viktor</mods:namePart>
<mods:affiliation id="F522CE635201CB685F1B8A10D60F7364">Institute for Evolutionary Ecology of the National Academy of Sciences of Ukraine, 37, Academician Lebediev St., 03143, Kyiv, Ukraine.</mods:affiliation>
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<mods:namePart id="F52F2AEC5C706C4E7B0016F350F48AA1">Khalaim, Yevhenii</mods:namePart>
<mods:affiliation id="587F849D1E4B0D6F68A00531AF5D72F1">Tuzly Lagoons National Nature Park, 2, Partyzanska St., 68100, Tatarbunary, Odesa region, Ukraine.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="EE656B11B2301C5D001C579453087A62">Budashkin, Yuriy</mods:namePart>
<mods:nameIdentifier id="002FAF9E3949D35A2D4E66D04BF67EC1" type="ORCID">0000-0001-7824-8338</mods:nameIdentifier>
<mods:affiliation id="82B578751240498BB72A7A3D6FD80127">T. I. Vyazemsky Karadag Scientific Station - Nature Reserve of RAS, Kurortnoye, Pheodosia, Crimea Republic, 298188. budashkin @ ukr. net; https: // orcid. org / 0000 - 0001 - 7824 - 8338</mods:affiliation>
<mods:nameIdentifier id="0159AEFD8FC15355C1D2809E9CEC4CC9" type="email">budashkin@ukr.net</mods:nameIdentifier>
</mods:name>
<mods:name id="815B6DBE18C9BC7737E2D62F7CFA954C" type="personal">
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<mods:roleTerm id="CD7A60EA8C392815766220B2DB8695D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD597316B2496426C6A4B6235E7C072E">Zhakov, Oleksandr</mods:namePart>
<mods:nameIdentifier id="D52A1366797E4599D8F25C11A12206DF" type="ORCID">0000-0002-6113-8375</mods:nameIdentifier>
<mods:affiliation id="DA7F0B84E15B75C1F298AF3464A1046E">Regional center of tourism and local history for students, 46 A, Nemirovich-Danchenko St., 69091, Zaporizhzhia, Ukraine. a. zhakov @ gmail. com; https: // orcid. org / 0000 - 0002 - 6113 - 8375</mods:affiliation>
<mods:nameIdentifier id="214BE27D686BA97190ACA8BB7DC8790A" type="email">a.zhakov@gmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="349970711C9A80E6DE36DFBEFC2CEECB">Mushynskyi, Vadym</mods:namePart>
<mods:affiliation id="DE3E14649DB39AD2A1C229AB7D69CC91">Zaporizhzhia, Ukraine.</mods:affiliation>
</mods:name>
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<mods:namePart id="03524C254EE43E7BBF5364B369E816F2">Novytskyi, Sergiy</mods:namePart>
<mods:affiliation id="B7AC7AE8D9AFF14EDA2C2041DB38039C">Hradenytsi, Odesa region, Ukraine.</mods:affiliation>
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<mods:title id="D056F484F942E08D5C944160B0D96070">Zootaxa</mods:title>
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<mods:date id="B6B15488315CCAA2B228C0DEA1618CE1">2021</mods:date>
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<mods:number id="F01AAFE2C1A9F9C8A384EFC2B3E3A6FF">5023</mods:number>
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<treatment id="03BA87F9E91825650EF7FBCC58C98CF5" ID-DOI="http://doi.org/10.5281/zenodo.5226141" ID-Zenodo-Dep="5226141" LSID="urn:lsid:plazi:treatment:03BA87F9E91825650EF7FBCC58C98CF5" httpUri="http://treatment.plazi.org/id/03BA87F9E91825650EF7FBCC58C98CF5" lastPageId="6" lastPageNumber="372" pageId="5" pageNumber="371">
<subSubSection id="C3096564E91825660EF7FBCC5EA18956" box="[151,651,1111,1138]" pageId="5" pageNumber="371" type="nomenclature">
<paragraph id="8BAC36EFE91825660EF7FBCC5EA18956" blockId="5.[151,651,1111,1174]" box="[151,651,1111,1138]" pageId="5" pageNumber="371">
<heading id="D0E48183E91825660EF7FBCC5EA18956" bold="true" box="[151,651,1111,1138]" fontSize="11" level="1" pageId="5" pageNumber="371" reason="1">
<taxonomicName id="4C134D6CE91825660EF7FBCC5EA18956" authority="(Ragonot, 1887)" baseAuthorityName="Ragonot" baseAuthorityYear="1887" box="[151,651,1111,1138]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="imperialella">
<emphasis id="B967EAFDE91825660EF7FBCC5EA18956" bold="true" box="[151,651,1111,1138]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EF7FBCC5D978956" bold="true" box="[151,445,1111,1138]" italics="true" pageId="5" pageNumber="371">Bradyrrhoa imperialella</emphasis>
(
<bibRefCitation id="EF824B1EE91825660FADFBC35EA98956" author="Ragonot, E. L." box="[461,643,1112,1138]" pageId="5" pageNumber="371" pagination="225 - 260" refId="ref17817" refString="Ragonot, E. L. (1887) Diagnoses d'especes nouvelles de Phycitidae d'Europe et des Pays limitrophes. Annales de la Societe Entomologique de France, Series 6, 7 (3), 225 - 260." type="journal article" year="1887">Ragonot, 1887</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3096564E91825660EF7FBE75D2789B2" box="[151,269,1148,1174]" pageId="5" pageNumber="371" type="description">
<paragraph id="8BAC36EFE91825660EF7FBE75D2789B2" blockId="5.[151,651,1111,1174]" box="[151,269,1148,1174]" pageId="5" pageNumber="371">
<heading id="D0E48183E91825660EF7FBE75D2789B2" bold="true" box="[151,269,1148,1174]" fontSize="11" level="1" pageId="5" pageNumber="371" reason="1">
<emphasis id="B967EAFDE91825660EF7FBE75D2789B2" bold="true" box="[151,269,1148,1174]" pageId="5" pageNumber="371">
(
<figureCitation id="13282A6AE91825660EFFFBE75D2F89B2" box="[159,261,1148,1174]" captionStart-0="FIGURES 67" captionStart-1="FIGURES 89" captionStartId-0="6.[151,264,1730,1754]" captionStartId-1="7.[151,264,984,1008]" captionTargetBox-0="[151,1399,490,1707]" captionTargetBox-1="[151,1436,309,961]" captionTargetId-0="figure-174@6.[151,1436,489,1707]" captionTargetId-1="figure-90@7.[151,1436,309,961]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 67. Female of Bradyrrhoa imperialella, Crimea. 6. Adult. 7. Female genitalia, prep. no. 435.20." captionText-1="FIGURES 89. Heads of females of Bradyrrhoa spp. 8. B. imperialella, Crimea, prep. no. 435.20. 9. B. gilveolella, Kherson region, prep. no. 449.20. (Black lines indicate the boundaries of the segments of labial palpus, Roman numerals indicate the order of the segments.)" figureDoi-0="http://doi.org/10.5281/zenodo.5226122" figureDoi-1="http://doi.org/10.5281/zenodo.5226124" httpUri-0="https://zenodo.org/record/5226122/files/figure.png" httpUri-1="https://zenodo.org/record/5226124/files/figure.png" pageId="5" pageNumber="371">Figs 68</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3096564E91825660EF7FB5F58208A66" pageId="5" pageNumber="371" type="materials_examined">
<paragraph id="8BAC36EFE91825660EF7FB5F5E828826" blockId="5.[151,1437,1219,2039]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EF7FB5F5D5389FA" bold="true" box="[151,377,1220,1246]" pageId="5" pageNumber="371">Material examined</emphasis>
:
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<specimenCount id="9D15FD66E91825660FE8FB5F5D8189FA" box="[392,427,1220,1246]" pageId="5" pageNumber="371" type="female">1♀</specimenCount>
,
<emphasis id="B967EAFDE91825660FD9FB5F5E3989FA" bold="true" box="[441,531,1220,1246]" pageId="5" pageNumber="371">
<collectingRegion id="49D7F80DE91825660FD9FB5F5E3989FA" box="[441,531,1220,1246]" country="Ukraine" name="Crimea" pageId="5" pageNumber="371">Crimea</collectingRegion>
</emphasis>
,
<location id="8ECC6034E91825660C41FB5F5EE289FA" LSID="urn:lsid:plazi:treatment:03BA87F9E91825650EF7FBCC58C98CF5:8ECC6034E91825660C41FB5F5EE289FA" box="[545,712,1220,1246]" name="Feodosia distr." pageId="5" pageNumber="371" stateProvince="Crimea">Feodosia distr.</location>
, Suuk-Su,
<collectingRegion id="49D7F80DE91825660D21FB5F5F8A89FA" box="[833,928,1220,1246]" country="Ukraine" name="Crimea" pageId="5" pageNumber="371">Crimaea</collectingRegion>
[
<emphasis id="B967EAFDE91825660DD0FB5E5FE589FA" box="[944,975,1221,1246]" italics="true" pageId="5" pageNumber="371">sic</emphasis>
],
<date id="FFAD102FE91825660D86FB5F585D89FA" box="[998,1143,1220,1246]" pageId="5" pageNumber="371" value="1917-08-11">
<collectingDate id="EFE9E9C7E91825660D86FB5F585D89FA" box="[998,1143,1220,1246]" pageId="5" pageNumber="371" value="1917-08-11">11.VIII.1917</collectingDate>
</date>
(
<collectorName id="26E65339E91825660AE7FB5F590089FB" box="[1159,1322,1220,1247]" pageId="5" pageNumber="371">L. Sheljuzhko</collectorName>
), genitalia slide:
<collectionCode id="ED02AE2AE91825660EBAFB725CC48827" box="[218,238,1257,1283]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="371" type="Museum">V</collectionCode>
</materialsCitation>
. Yepishin, prep. no. 435.20
<materialsCitation id="3B7B3CB2E91825660C47FB735E8E8826" box="[551,676,1256,1283]" collectionCode="ZMKU" pageId="5" pageNumber="371" specimenCount="1" specimenCount-female="1" stateProvince="Crimea">
<specimenCount id="9D15FD66E91825660C47FB735E1E8827" box="[551,564,1256,1283]" pageId="5" pageNumber="371" type="female"></specimenCount>
(
<collectionCode id="ED02AE2AE91825660C23FB735EB68826" box="[579,668,1256,1282]" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="5" pageNumber="371">ZMKU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36EFE91825660EA7FA975D9C884A" blockId="5.[151,1437,1219,2039]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EA7FA975E018802" bold="true" box="[199,555,1292,1318]" pageId="5" pageNumber="371">Comparative material studied</emphasis>
.
<taxonomicName id="4C134D6CE91825660C58FA90583B8802" authority="(Treitschke, 1833)" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[568,1041,1291,1318]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91825660C58FA905F108802" bold="true" box="[568,826,1291,1318]" italics="true" pageId="5" pageNumber="371">Bradyrrhoa gilveolella</emphasis>
(
<bibRefCitation id="EF824B1EE91825660D27FA9758228802" author="Treitschke, F." box="[839,1032,1292,1318]" pageId="5" pageNumber="371" refId="ref19300" refString="Treitschke, F. (1833) Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer'schen Werks). Ernst Fleischer, Leipzig, 294 pp." type="book" year="1833">Treitschke, 1833</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="3B7B3CB2E91825660A79FA975D98884A" collectingDate="2019-07-12" collectingDateMax="2019-07-14" collectingDateMin="2019-07-12" collectionCode="V" collectorName="SE Burkuty" latitude="46.393333" location="Skadovsk distr." longLatPrecision="18" longitude="32.810276" pageId="5" pageNumber="371" specimenCount="2" specimenCount-female="2" stateProvince="Kherson">
<specimenCount id="9D15FD66E91825660A79FA9758168802" box="[1049,1084,1292,1318]" pageId="5" pageNumber="371" type="female">1♀</specimenCount>
,
<emphasis id="B967EAFDE91825660A29FA97589A8802" bold="true" box="[1097,1200,1292,1318]" pageId="5" pageNumber="371">
<collectingRegion id="49D7F80DE91825660A29FA97589A8802" box="[1097,1200,1292,1318]" country="Ukraine" name="Kherson" pageId="5" pageNumber="371">Kherson</collectingRegion>
</emphasis>
reg.,
<location id="8ECC6034E91825660A8CFA9759B28802" LSID="urn:lsid:plazi:treatment:03BA87F9E91825650EF7FBCC58C98CF5:8ECC6034E91825660A8CFA9759B28802" box="[1260,1432,1292,1318]" latitude="46.393333" longLatPrecision="18" longitude="32.810276" name="Skadovsk distr." pageId="5" pageNumber="371" stateProvince="Kherson">Skadovsk distr.</location>
, 1,5 km
<collectorName id="26E65339E91825660E8EFAAB5D59886E" box="[238,371,1328,1354]" pageId="5" pageNumber="371">SE Burkuty</collectorName>
,
<geoCoordinate id="EE275028E91825660F1FFAAB5E21886E" box="[383,523,1328,1354]" degrees="46" direction="north" minutes="23" orientation="latitude" pageId="5" pageNumber="371" precision="15" seconds="36" value="46.393333">46°2336”N</geoCoordinate>
,
<geoCoordinate id="EE275028E91825660C77FAAB5EB5886E" box="[535,671,1328,1354]" degrees="32" direction="east" minutes="48" orientation="longitude" pageId="5" pageNumber="371" precision="15" seconds="37" value="32.810276">32°4837”E</geoCoordinate>
, at light,
<date id="FFAD102FE91825660D6BFAAB5F97886E" box="[779,957,1328,1354]" pageId="5" pageNumber="371" value="2019-07-12" valueMax="2019-07-14" valueMin="2019-07-12">
<collectingDate id="EFE9E9C7E91825660D6BFAAB5F97886E" box="[779,957,1328,1354]" pageId="5" pageNumber="371" value="2019-07-12" valueMax="2019-07-14" valueMin="2019-07-12">1214.VII.2019</collectingDate>
</date>
(
<collectionCode id="ED02AE2AE91825660DAEFAAB5FC9886E" box="[974,995,1328,1354]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="371" type="Museum">V</collectionCode>
. Yepishin), genitalia slide:
<collectionCode id="ED02AE2AE91825660B74FAAA5902886F" box="[1300,1320,1329,1355]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="371" type="Museum">V</collectionCode>
. Yepishin, prep. no. 449.20
<specimenCount id="9D15FD66E91825660F36FACF5D49884A" box="[342,355,1364,1390]" pageId="5" pageNumber="371" type="female"></specimenCount>
(VYe)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36EFE91825660EA7FAE35EBA8B06" blockId="5.[151,1437,1219,2039]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EA7FAE35DA288B6" bold="true" box="[199,392,1400,1426]" pageId="5" pageNumber="371">Notes on female</emphasis>
. Sexual dimorphism is not revealed, except for antennae, in males the flagellum is slightly thicker, flattened and sinuate basally, labial palpi are the same. Antennae of female rounded, ventrally brown, as in male not densely covered with very short cilia visible only at very high magnification, dorsally covered with tight-fitting white-yellow scales. For a detailed description of the imago see
<bibRefCitation id="EF824B1EE91825660D8FFA7E593E88DB" author="Ragonot, E. L." box="[1007,1300,1508,1535]" pageId="5" pageNumber="371" pagination="1 - 658" refId="ref17898" refString="Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M., Memoires sur les Lepidopteres, St. Petersburg, VII (7), pp. i - lvi + 1 - 658, pls. 1 - 23." type="book chapter" year="1893">Ragonot (1893: 266277)</bibRefCitation>
. The female genitalia are described here for the first time.
</paragraph>
<paragraph id="8BAC36EFE91825660EA7F9B758208A66" blockId="5.[151,1437,1219,2039]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EA7F9B75DA68B62" bold="true" box="[199,396,1580,1606]" pageId="5" pageNumber="371">Female genitalia</emphasis>
(
<figureCitation id="13282A6AE91825660FFAF9B75DF58B63" box="[410,479,1580,1607]" captionStart="FIGURES 67" captionStartId="6.[151,264,1730,1754]" captionTargetBox="[151,1399,490,1707]" captionTargetId="figure-174@6.[151,1436,489,1707]" captionTargetPageId="6" captionText="FIGURES 67. Female of Bradyrrhoa imperialella, Crimea. 6. Adult. 7. Female genitalia, prep. no. 435.20." figureDoi="http://doi.org/10.5281/zenodo.5226122" httpUri="https://zenodo.org/record/5226122/files/figure.png" pageId="5" pageNumber="371">Fig. 7</figureCitation>
): papillae anales large, covered with long filiform scales, taper abruptly in the middle, so that distal part twice as narrower as the base. Posterior and anterior apophyses of same length. Segment VIII equal in length and width, tergum VIII with a non-sclerotized zone, in the posterior half of the segment, and with small V-shaped anteromedial emargination, sternum VIII with broad anteromedial emargination. Antrum funnel-shaped, very sclerotized, ostium very wide. Ductus bursae short, almost twice shorter than the antrum. Ductus bursae and seminalis equal in width. Ductus seminalis emerges from the base of the bursa copulatrix near ductus bursae.
<collectingRegion id="49D7F80DE91825660B3BF97A59B68BDF" box="[1371,1436,1761,1787]" country="Turkey" name="Bursa" pageId="5" pageNumber="371">Bursa</collectingRegion>
copulatrix oviform. Ductus bursae, ductus seminalis and bursa copulatrix have many needle-like signae. Signae of bursa copulatrix three times longer than signae of ductus bursae and seminalis.
</paragraph>
</subSubSection>
<subSubSection id="C3096564E91825660EA7F8D7589D8AD2" pageId="5" pageNumber="371" type="discussion">
<paragraph id="8BAC36EFE91825660EA7F8D7589D8AD2" blockId="5.[151,1437,1219,2039]" pageId="5" pageNumber="371">
<emphasis id="B967EAFDE91825660EA7F8D75D038A42" bold="true" box="[199,297,1868,1894]" pageId="5" pageNumber="371">Remark</emphasis>
. Non-fresh or pale-coloured specimens can be confused with
<taxonomicName id="4C134D6CE91825660D8EF8D658AE8A42" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[1006,1156,1868,1894]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91825660D8EF8D658AE8A42" box="[1006,1156,1868,1894]" italics="true" pageId="5" pageNumber="371">B. gilveolella</emphasis>
</taxonomicName>
. Antennae in females of these species do not differ, but labial palpi have characteristic differences. In
<taxonomicName id="4C134D6CE91825660A78F8EA589A8AAE" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[1048,1200,1904,1930]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91825660A78F8EA589A8AAE" box="[1048,1200,1904,1930]" italics="true" pageId="5" pageNumber="371">B. gilveolella</emphasis>
</taxonomicName>
II-segment of labial palpus is twice as long as the diameter of eye, cylindrical, directed forward (
<figureCitation id="13282A6AE91825660D83F80F580C8A8A" box="[995,1062,1940,1966]" captionStart="FIGURES 89" captionStartId="7.[151,264,984,1008]" captionTargetBox="[151,1436,309,961]" captionTargetId="figure-90@7.[151,1436,309,961]" captionTargetPageId="7" captionText="FIGURES 89. Heads of females of Bradyrrhoa spp. 8. B. imperialella, Crimea, prep. no. 435.20. 9. B. gilveolella, Kherson region, prep. no. 449.20. (Black lines indicate the boundaries of the segments of labial palpus, Roman numerals indicate the order of the segments.)" figureDoi="http://doi.org/10.5281/zenodo.5226124" httpUri="https://zenodo.org/record/5226124/files/figure.png" pageId="5" pageNumber="371">Fig. 9</figureCitation>
). In
<taxonomicName id="4C134D6CE91825660A38F80E58D58A8A" baseAuthorityName="Ragonot" baseAuthorityYear="1887" box="[1112,1279,1940,1966]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="imperialella">
<emphasis id="B967EAFDE91825660A38F80E58D58A8A" box="[1112,1279,1940,1966]" italics="true" pageId="5" pageNumber="371">B. imperialella</emphasis>
</taxonomicName>
II-segment of labial palpus is not much longer than diameter of eye, fusiform, directed upwards (
<figureCitation id="13282A6AE91825660A50F82358598AF6" box="[1072,1139,1976,2002]" captionStart="FIGURES 89" captionStartId="7.[151,264,984,1008]" captionTargetBox="[151,1436,309,961]" captionTargetId="figure-90@7.[151,1436,309,961]" captionTargetPageId="7" captionText="FIGURES 89. Heads of females of Bradyrrhoa spp. 8. B. imperialella, Crimea, prep. no. 435.20. 9. B. gilveolella, Kherson region, prep. no. 449.20. (Black lines indicate the boundaries of the segments of labial palpus, Roman numerals indicate the order of the segments.)" figureDoi="http://doi.org/10.5281/zenodo.5226124" httpUri="https://zenodo.org/record/5226124/files/figure.png" pageId="5" pageNumber="371">Fig. 8</figureCitation>
), all these traits also apply to males. Other differences between males have already been discussed (
<bibRefCitation id="EF824B1EE91825660DA4F84758868AD2" author="Yepishin, V. &amp; Bidzilya, O. &amp; Budashkin, Yu. &amp; Zhakov, O. &amp; Mushynskyi, V. &amp; Novytskyi, S." box="[964,1196,2012,2039]" pageId="5" pageNumber="371" pagination="101 - 120" refId="ref19552" refString="Yepishin, V., Bidzilya, O., Budashkin, Yu., Zhakov, O., Mushynskyi, V. &amp; Novytskyi, S. (2020) New records of little known pyraloid moths (Lepidoptera: Pyraloidea) from Ukraine. Zootaxa, 4808 (1), 101 - 120. https: // doi. org / 10.11646 / zootaxa. 4808.1.5" type="journal article" year="2020">
Yepishin
<emphasis id="B967EAFDE91825660A50F846584B8AD2" box="[1072,1121,2012,2038]" italics="true" pageId="5" pageNumber="371">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3096564E91B25650EA7FF0D58C98CF5" pageId="6" pageNumber="372" type="distribution">
<paragraph id="8BAC36EFE91B25650EA7FF0D58568DF1" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="372">
<emphasis id="B967EAFDE91B25650EA7FF0D5D738D94" bold="true" box="[199,345,150,176]" pageId="6" pageNumber="372">Distribution</emphasis>
: North
<collectingCountry id="F304767FE91B25650FD4FF0C5E198D95" box="[436,563,151,177]" name="Macedonia" pageId="6" pageNumber="372">Macedonia</collectingCountry>
,
<collectingCountry id="F304767FE91B25650C5FFF0C5EBB8D95" box="[575,657,151,177]" name="Greece" pageId="6" pageNumber="372">Greece</collectingCountry>
,
<collectingCountry id="F304767FE91B25650CFEFF0C5EC58D95" box="[670,751,151,177]" name="Turkey" pageId="6" pageNumber="372">Turkey</collectingCountry>
,
<collectingCountry id="F304767FE91B25650C9BFF0C5F608D95" box="[763,842,151,177]" name="Russia" pageId="6" pageNumber="372">Russia</collectingCountry>
: Siberia (
<bibRefCitation id="EF824B1EE91B25650DD5FF0C58708D95" author="Ragonot, E. L." box="[949,1114,151,177]" pageId="6" pageNumber="372" pagination="1 - 658" refId="ref17898" refString="Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M., Memoires sur les Lepidopteres, St. Petersburg, VII (7), pp. i - lvi + 1 - 658, pls. 1 - 23." type="book chapter" year="1893">Ragonot 1893</bibRefCitation>
;
<bibRefCitation id="EF824B1EE91B25650A07FF0C594E8D95" author="Kocak, A. O. &amp; Kemal, M." box="[1127,1380,151,177]" pageId="6" pageNumber="372" pagination="1 - 487" refId="ref16395" refString="Kocak, A. O. &amp; Kemal, M. (2018) A synonymous and distributional list of the species of the Lepidoptera of Turkey. Centre for Entomological Studies, Memoirs, 8, 1 - 487." type="journal article" year="2018">Koçak &amp; Kemal 2018</bibRefCitation>
) and Caucasus (Sinev
<emphasis id="B967EAFDE91B25650F38FF205DBA8DF1" box="[344,400,187,213]" italics="true" pageId="6" pageNumber="372">et al.</emphasis>
2019);
<collectingCountry id="F304767FE91B25650F87FF205E6D8DF1" box="[487,583,187,213]" name="Bulgaria" pageId="6" pageNumber="372">Bulgaria</collectingCountry>
(
<bibRefCitation id="EF824B1EE91B25650C36FF205EFF8DF2" author="Plant, C. W." box="[598,725,187,214]" pageId="6" pageNumber="372" pagination="1 - 231" refId="ref17619" refString="Plant, C. W. (2016) An Annotated Systematic, Synonymic and Distributional Checklist of the Pyraloidea of Bulgaria (Lepidoptera, Crambidae &amp; Pyralidae). Neue Entomologische Nachrichten, 72, 1 - 231." type="journal article" year="2016">Plant 2016</bibRefCitation>
);
<collectingCountry id="F304767FE91B25650C89FF205F628DF1" box="[745,840,187,213]" name="Ukraine" pageId="6" pageNumber="372">Ukraine</collectingCountry>
:
<collectingRegion id="49D7F80DE91B25650D32FF205F8E8DF1" box="[850,932,187,213]" country="Ukraine" name="Crimea" pageId="6" pageNumber="372">Crimea</collectingRegion>
(
<bibRefCitation id="EF824B1EE91B25650DD2FF20585A8DF1" author="Budashkin, Yu. I." box="[946,1136,187,213]" pageId="6" pageNumber="372" pagination="323 - 366" refId="ref12637" refString="Budashkin, Yu. I. (2004) The results of a twenty-year stationary study of the lepidopteran fauna of the Karadag natural reserve [Itogi dvadtsatiletnego statsionarnogo izucheniya fauny cheshuekrylykh (Lepidoptera) Karadagskogo prirodnogo zapovednika]. In: Karadag. Collection of scientific papers. Book 1. Sonat, Simferopol, pp. 323 - 366. [in Russian]" type="book chapter" year="2004">Budashkin 2004</bibRefCitation>
).
</paragraph>
<paragraph id="8BAC36EFE91B25650EA7FF4458C98CF5" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="372">
<emphasis id="B967EAFDE91B25650EA7FF445D9A8DDD" bold="true" box="[199,432,223,249]" pageId="6" pageNumber="372">Notes on taxonomy</emphasis>
. Having undergone combinations with different genera, this species still remains in the genus
<taxonomicName id="4C134D6CE91B25650E81FE985E438C39" authority="(Yepishin et al. 2020)" baseAuthorityName="Yepishin" baseAuthorityYear="2020" box="[225,617,259,285]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B967EAFDE91B25650E81FE985D4F8C39" box="[225,357,259,285]" italics="true" pageId="6" pageNumber="372">Bradyrrhoa</emphasis>
(
<bibRefCitation id="EF824B1EE91B25650F15FE985E4B8C39" author="Yepishin, V. &amp; Bidzilya, O. &amp; Budashkin, Yu. &amp; Zhakov, O. &amp; Mushynskyi, V. &amp; Novytskyi, S." box="[373,609,259,285]" pageId="6" pageNumber="372" pagination="101 - 120" refId="ref19552" refString="Yepishin, V., Bidzilya, O., Budashkin, Yu., Zhakov, O., Mushynskyi, V. &amp; Novytskyi, S. (2020) New records of little known pyraloid moths (Lepidoptera: Pyraloidea) from Ukraine. Zootaxa, 4808 (1), 101 - 120. https: // doi. org / 10.11646 / zootaxa. 4808.1.5" type="journal article" year="2020">
Yepishin
<emphasis id="B967EAFDE91B25650F82FE985E3F8C39" box="[482,533,259,285]" italics="true" pageId="6" pageNumber="372">et al</emphasis>
. 2020
</bibRefCitation>
)
</taxonomicName>
. However, new data: a simple bursa, a highly sclerotized funnel-shaped antrum in the female genitalia and a specific edeagus with its cornuti in the male genitalia in comparison with those of a typical species—
<taxonomicName id="4C134D6CE91B25650FE8FED05EA28C41" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[392,648,331,357]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91B25650FE8FED05EA28C41" box="[392,648,331,357]" italics="true" pageId="6" pageNumber="372">Bradyrrhoa gilveolella</emphasis>
</taxonomicName>
, the presence of a sinus in the male antennae (absent in
<taxonomicName id="4C134D6CE91B25650A97FED059A68C41" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[1271,1420,331,357]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91B25650A97FED059228C40" box="[1271,1288,331,356]" italics="true" pageId="6" pageNumber="372">B</emphasis>
.
<emphasis id="B967EAFDE91B25650B76FED059A68C41" box="[1302,1420,331,357]" italics="true" pageId="6" pageNumber="372">gilveolella</emphasis>
</taxonomicName>
), as well as a caterpillar host plant—
<taxonomicName id="4C134D6CE91B25650C49FEF45F718CAD" authority="Lebed." authorityName="Lebed." box="[553,859,367,393]" class="Magnoliopsida" family="Boraginaceae" genus="Onosma" kingdom="Plantae" order="Boraginales" pageId="6" pageNumber="372" phylum="Tracheophyta" rank="species" species="polyphylla">
<emphasis id="B967EAFDE91B25650C49FEF45F2F8CAD" box="[553,773,367,393]" italics="true" pageId="6" pageNumber="372">Onosma polyphylla</emphasis>
Lebed.
</taxonomicName>
from the family
<taxonomicName id="4C134D6CE91B25650A7DFEF4589F8CAD" box="[1053,1205,367,393]" class="Magnoliopsida" family="Boraginaceae" kingdom="Plantae" order="Boraginales" pageId="6" pageNumber="372" phylum="Tracheophyta" rank="family">Boraginaceae</taxonomicName>
(famous host plants of
<taxonomicName id="4C134D6CE91B25650ED8FE085D658C89" baseAuthorityName="Treitschke" baseAuthorityYear="1833" box="[184,335,403,429]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="species" species="gilveolella">
<emphasis id="B967EAFDE91B25650ED8FE085CE38C88" box="[184,201,403,428]" italics="true" pageId="6" pageNumber="372">B</emphasis>
.
<emphasis id="B967EAFDE91B25650EB9FE085D658C89" box="[217,335,403,429]" italics="true" pageId="6" pageNumber="372">gilveolella</emphasis>
</taxonomicName>
and other species of the genus
<taxonomicName id="4C134D6CE91B25650CDDFE085F6B8C89" box="[701,833,403,429]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B967EAFDE91B25650CDDFE085F6B8C89" box="[701,833,403,429]" italics="true" pageId="6" pageNumber="372">Bradyrrhoa</emphasis>
</taxonomicName>
—are from the family
<taxonomicName id="4C134D6CE91B25650A23FE0858E88C89" box="[1091,1218,403,429]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="372" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
), cast doubt on the position of
<taxonomicName id="4C134D6CE91B25650F77FE2C5D958CF5" baseAuthorityName="Ragonot" baseAuthorityYear="1887" box="[279,447,439,465]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="species" species="imperialella">
<emphasis id="B967EAFDE91B25650F77FE2C5D028CF4" box="[279,296,439,464]" italics="true" pageId="6" pageNumber="372">B</emphasis>
.
<emphasis id="B967EAFDE91B25650F56FE2C5D958CF5" box="[310,447,439,465]" italics="true" pageId="6" pageNumber="372">imperialella</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="4C134D6CE91B25650C34FE2C5EF28CF5" box="[596,728,439,465]" class="Insecta" family="Pyralidae" genus="Bradyrrhoa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B967EAFDE91B25650C34FE2C5EF28CF5" box="[596,728,439,465]" italics="true" pageId="6" pageNumber="372">Bradyrrhoa</emphasis>
</taxonomicName>
. Further investigation is need for clarification.
</paragraph>
</subSubSection>
</treatment>
</document>