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<document id="CA1D9642337215CFA9D42EC4C6E3A1DC" ID-DOI="10.11646/zootaxa.5389.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10421859" ID-ZooBank="45230245-48E8-4BEF-B381-4CB8FCB264C1" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1703234013262" checkinUser="plazi" docAuthor="Gadallah, Neveen S., Edmardash, Yusuf A., Mansour, Amany N. &amp; Imam, Ahmed I." docDate="2023" docId="03BA87BCD777FFE0FF58429DFDA86DC3" docLanguage="en" docName="zootaxa.5389.5.1.pdf" docOrigin="Zootaxa 5389 (5)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5389.5.1/52536" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Praon volucre" docType="treatment" docVersion="3" lastPageNumber="510" masterDocId="FF83FFC4D77EFFE9FFCF420AFFB26E66" masterDocTitle="Parasitoid wasps (Ichneumonoidea) collected from faba bean fields, Kharga Oasis, New Valley, Egypt, with new records and the description of a new species" masterLastPageNumber="544" masterPageNumber="501" pageNumber="510" updateTime="1704282657916" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="2C03D646C6DD7025EA182743A607EF42">Parasitoid wasps (Ichneumonoidea) collected from faba bean fields, Kharga Oasis, New Valley, Egypt, with new records and the description of a new species</mods:title>
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<mods:namePart id="8A8809E698A5CB3A3DC281A83B368458">Gadallah, Neveen S.</mods:namePart>
<mods:affiliation id="E4387306D7D3BFBBC66275DF796C1F3B">Entomology Department, Faculty of Science, Cairo University, Giza, Egypt;</mods:affiliation>
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<mods:namePart id="96DF12988C712C85A77EC56046934E43">Edmardash, Yusuf A.</mods:namePart>
<mods:affiliation id="06BF9AEA5E49A7FC39B85B8951612DB6">Entomology Department, Faculty of Science, Cairo University, Giza, Egypt;</mods:affiliation>
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<mods:namePart id="02EBFD63B1B1A13C7DB2AFD2092C55E5">Mansour, Amany N.</mods:namePart>
<mods:affiliation id="FB574D890B57A44D5009ACAC6858AC9D">Entomology Department, Faculty of Science, Cairo University, Giza, Egypt; &amp; Plant Protection Department, Desert Research Center, Cairo Egypt</mods:affiliation>
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<mods:namePart id="E4DF6483D09170E1DFAFB6AF660B6D3A">Imam, Ahmed I.</mods:namePart>
<mods:affiliation id="97045931BF8F43710A289D0DF0C01A40">Entomology Department, Faculty of Science, Cairo University, Giza, Egypt; &amp; Plant Protection Department, Desert Research Center, Cairo Egypt &amp; Entomology Department, Faculty of Science, Cairo University, Giza, Egypt;</mods:affiliation>
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<subSubSection id="C3096521D777FFE0FF58429DFDBB6ED7" box="[151,521,151,178]" pageId="9" pageNumber="510" type="nomenclature">
<paragraph id="8BAC36AAD777FFE0FF58429DFDBB6ED7" blockId="9.[151,521,151,213]" box="[151,521,151,178]" pageId="9" pageNumber="510">
<heading id="D0E481C6D777FFE0FF58429DFDBB6ED7" bold="true" box="[151,521,151,178]" fontSize="11" level="1" pageId="9" pageNumber="510" reason="1">
<taxonomicName id="4C134D29D777FFE0FF58429DFDBB6ED7" ID-CoL="4M93N" authority="(Haliday, 1833)" baseAuthorityName="Haliday" baseAuthorityYear="1833" box="[151,521,151,178]" class="Insecta" family="Braconidae" genus="Praon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="volucre">
<emphasis id="B967EAB8D777FFE0FF58429DFDBB6ED7" bold="true" box="[151,521,151,178]" pageId="9" pageNumber="510">
<emphasis id="B967EAB8D777FFE0FF58429DFEF16ED4" bold="true" box="[151,323,151,178]" italics="true" pageId="9" pageNumber="510">Praon volucre</emphasis>
(Haliday, 1833)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C3096521D777FFE0FF5842B1FE956EB3" box="[151,295,187,213]" pageId="9" pageNumber="510" type="description">
<paragraph id="8BAC36AAD777FFE0FF5842B1FE956EB3" blockId="9.[151,521,151,213]" box="[151,295,187,213]" pageId="9" pageNumber="510">
<figureCitation id="13282A2FD777FFE0FF5842B1FF6B6EB3" box="[151,217,187,213]" captionStart="FIGURE 4" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[253,1335,181,1919]" captionTargetId="figure-32@8.[253,1335,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Praon volucre (Haliday, 1833) (♂): A, lateral habitus; B, dorsal habitus; C, fore wing." figureDoi="http://doi.org/10.5281/zenodo.10421875" httpUri="https://zenodo.org/record/10421875/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
(AC)
</paragraph>
</subSubSection>
<subSubSection id="C3096521D777FFE0FF584309FE316F00" pageId="9" pageNumber="510" type="materials_examined">
<paragraph id="8BAC36AAD777FFE0FF584309FE316F00" blockId="9.[151,1437,259,933]" pageId="9" pageNumber="510">
<emphasis id="B967EAB8D777FFE0FF584309FE316F7B" bold="true" box="[151,387,259,285]" pageId="9" pageNumber="510">Material examined:</emphasis>
<specimenCount id="9D15FD23D777FFE0FE454309FE026F7B" box="[394,432,259,285]" count="2" pageId="9" pageNumber="510" type="male">2♂</specimenCount>
, Kharga Oasis (
<collectingRegion id="49D7F848D777FFE0FDAD4309FD5A6F7B" box="[610,744,259,285]" country="Egypt" name="Al Wadi al Jadid" pageId="9" pageNumber="510">New Valley</collectingRegion>
) [
<geoCoordinate id="EE27506DD777FFE0FD314309FC2D6F78" box="[766,927,259,286]" degrees="25" direction="north" minutes="33" orientation="latitude" pageId="9" pageNumber="510" precision="1" seconds="41.9" value="25.561638">25°3341.9”N</geoCoordinate>
<geoCoordinate id="EE27506DD777FFE0FC694309FBF46F7B" box="[934,1094,259,285]" degrees="30" direction="east" minutes="37" orientation="longitude" pageId="9" pageNumber="510" precision="1" seconds="41.9" value="30.628305">30°3741.9”E</geoCoordinate>
], May, 2022, white pan trap in
<taxonomicName id="4C134D29D777FFE0FF584322FEBA6F27" box="[151,264,295,321]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="faba">
<emphasis id="B967EAB8D777FFE0FF584322FEBA6F27" box="[151,264,295,321]" italics="true" pageId="9" pageNumber="510">Vicia faba</emphasis>
</taxonomicName>
;
<specimenCount id="9D15FD23D777FFE0FED9432DFE886F27" box="[278,314,295,321]" count="1" pageId="9" pageNumber="510" type="male">1♂</specimenCount>
, same locality and date but [
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<geoCoordinate id="EE27506DD777FFE0FCD6432DFC0A6F27" box="[793,952,295,321]" degrees="30" direction="east" minutes="31" orientation="longitude" pageId="9" pageNumber="510" precision="1" seconds="32.2" value="30.52561">30°3132.2”E</geoCoordinate>
], yellow pan trap in
<taxonomicName id="4C134D29D777FFE0FB574322FABB6F27" box="[1176,1289,295,321]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="faba">
<emphasis id="B967EAB8D777FFE0FB574322FABB6F27" box="[1176,1289,295,321]" italics="true" pageId="9" pageNumber="510">Vicia faba</emphasis>
</taxonomicName>
intercropped with
<taxonomicName id="4C134D29D777FFE0FF1F4346FECE6F03" box="[208,380,332,357]" class="Magnoliopsida" family="Brassicaceae" genus="Brassica" kingdom="Plantae" order="Brassicales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="napus">
<emphasis id="B967EAB8D777FFE0FF1F4346FECE6F03" box="[208,380,332,357]" italics="true" pageId="9" pageNumber="510">Brassica napus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3096521D777FFE0FF084365FCCD6C58" pageId="9" pageNumber="510" type="diagnosis">
<paragraph id="8BAC36AAD777FFE0FF084365FCCD6C58" blockId="9.[151,1437,259,933]" pageId="9" pageNumber="510">
<emphasis id="B967EAB8D777FFE0FF084365FEF06FEF" bold="true" box="[199,322,367,393]" pageId="9" pageNumber="510">Diagnosis.</emphasis>
Head and mesosoma dark brown to black (with lighter pronotum), antenna dark brown, legs clear yellow, metasoma with
<specimenCode id="DBB59ED1D777FFE0FE66439EFE706FD2" box="[425,450,404,436]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="510">
T
<subScript id="179734EFD777FFE0FE7543ACFE706FD2" attach="left" box="[442,450,422,436]" fontSize="6" pageId="9" pageNumber="510">1</subScript>
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honey yellow, rest of segments clear yellow with whitish intersegments; pterostigma waxy white; antenna with 18 antennomers (
<specimenCount id="9D15FD23D777FFE0FDB343BDFD216FB7" box="[636,659,439,465]" count="1" pageId="9" pageNumber="510" type="male"></specimenCount>
), F1 dark brown, with indistinct yellow spot at base; pterostigma 1.3× as long as vein
<specimenCode id="DBB59ED1D777FFE0FE8543D1FEDF6F93" box="[330,365,475,501]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="510">R1</specimenCode>
; vein
<collectingRegion id="49D7F848D777FFE0FE7C43D1FE676F93" box="[435,469,475,501]" country="Bulgaria" name="Ruse" pageId="9" pageNumber="510">RS</collectingRegion>
of fore wing normally developed; vein
<collectingRegion id="49D7F848D777FFE0FC5243D1FC0D6F93" box="[925,959,475,501]" country="Bulgaria" name="Ruse" pageId="9" pageNumber="510">RS</collectingRegion>
+M present, colourless throughout; lateral lobes of mesoscutum densely setose; notauli complete, convergent posteriorly; propodeum smooth, densely setose dorsally; petiole relatively short, 1.2× as long as wide at spiracle;.
</paragraph>
</subSubSection>
<subSubSection id="C3096521D777FFE0FF08404DFF5F6CE3" pageId="9" pageNumber="510" type="distribution">
<paragraph id="8BAC36AAD777FFE0FF08404DFF5F6CE3" blockId="9.[151,1437,259,933]" pageId="9" pageNumber="510">
<emphasis id="B967EAB8D777FFE0FF08404DFDAA6C07" bold="true" box="[199,536,583,609]" pageId="9" pageNumber="510">Distribution in the MENA:</emphasis>
<collectingCountry id="F304763AD777FFE0FDED404DFDCB6C07" box="[546,633,583,609]" name="Algeria" pageId="9" pageNumber="510">Algeria</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FD474042FD7C6C04" box="[648,718,584,610]" name="Egypt" pageId="9" pageNumber="510">Egypt</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FD114042FCBC6C04" box="[734,782,584,610]" name="Iran" pageId="9" pageNumber="510">Iran</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FCD24042FCFF6C04" box="[797,845,584,610]" name="Iraq" pageId="9" pageNumber="510">Iraq</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FC92404DFC2C6C07" box="[861,926,583,609]" name="Israel" pageId="9" pageNumber="510">Israel</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FC61404DFBA16C07" box="[942,1043,583,609]" name="Lebanon" pageId="9" pageNumber="510">Lebanon</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FBED4042FB386C04" box="[1058,1162,584,610]" name="Morocco" pageId="9" pageNumber="510">Morocco</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FB55404DFA816C07" box="[1178,1331,583,609]" name="Saudi Arabia" pageId="9" pageNumber="510">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FA8D404DFA2A6C07" box="[1346,1432,583,609]" name="Tunisia" pageId="9" pageNumber="510">Tunisia</collectingCountry>
,
<collectingCountry id="F304763AD777FFE0FF584061FF5B6CE3" box="[151,233,619,645]" name="Turkey" pageId="9" pageNumber="510">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3096521D777FFE0FF084085FDA86DC3" pageId="9" pageNumber="510" type="discussion">
<paragraph id="8BAC36AAD777FFE0FF084085FDA86DC3" blockId="9.[151,1437,259,933]" pageId="9" pageNumber="510">
<emphasis id="B967EAB8D777FFE0FF084085FEE06CCF" bold="true" box="[199,338,655,681]" pageId="9" pageNumber="510">Comments:</emphasis>
Although the key of
<bibRefCitation id="EF824B5BD777FFE0FD9A4085FCDA6CCF" author="Rakhshani, E. &amp; Barahoei, H. &amp; Ahmad, Z. &amp; Stary, P. &amp; Ghafouri-Moghaddam, M. &amp; Mehrparvar, M. &amp; Kavallieratos, N. G. &amp; Ckrkic, J. &amp; Tomanovic, Z." box="[597,872,655,682]" pageId="9" pageNumber="510" pagination="1 - 132" refId="ref20791" refString="Rakhshani, E., Barahoei, H., Ahmad, Z., Stary, P., Ghafouri-Moghaddam, M., Mehrparvar, M., Kavallieratos, N. G., Ckrkic, J. &amp; Tomanovic, Z. (2019) Review of Aphidiinae parasitoids (Hymenoptera: Braconidae) of the Middle East and north Africa: key to species and host associations. European Journal of Taxonomy, 552, 1 - 132. https: // doi. org / 10.5852 / ejt. 2019.552" type="journal article" year="2019">
Rakhshani
<emphasis id="B967EAB8D777FFE0FD19409AFCBE6CCF" box="[726,780,655,681]" italics="true" pageId="9" pageNumber="510">et al</emphasis>
. (2019)
</bibRefCitation>
is based on females, the characters of the male specimens agree with couplet
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their key, excluding the ovipositor characters. It also agrees with Mescheloff &amp; Rosens description (1989: 94). Although it has been recorded as an endoparasitoid of a wide range of aphid species affecting several economic crops, especially in this instance it has been recorded as a parasitoid of
<taxonomicName id="4C134D29D777FFE0FA9340F1FF676D5F" authorityName="Scopoli" authorityYear="1763" class="Insecta" family="Aphididae" genus="Aphis" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="fabae">
<emphasis id="B967EAB8D777FFE0FA9340F1FF676D5F" italics="true" pageId="9" pageNumber="510">Aphis fabae</emphasis>
</taxonomicName>
attacking
<taxonomicName id="4C134D29D777FFE0FE87412AFE276D5F" box="[328,405,799,825]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="faba">
<emphasis id="B967EAB8D777FFE0FE87412AFE276D5F" box="[328,405,799,825]" italics="true" pageId="9" pageNumber="510">V. faba</emphasis>
</taxonomicName>
in
<collectingCountry id="F304763AD777FFE0FE79412AFE516D5C" box="[438,483,800,826]" name="Iran" pageId="9" pageNumber="510">Iran</collectingCountry>
(
<bibRefCitation id="EF824B5BD777FFE0FE3E4115FD556D5F" author="Rakhshani, E. &amp; Talebi, AA &amp; Manzari, S. &amp; Tomanovic, Z. &amp; Stary, P. &amp; Rezwani, A." box="[497,743,799,826]" pageId="9" pageNumber="510" pagination="19 - 34" refId="ref20640" refString="Rakhshani, E., Talebi, AA., Manzari, S., Tomanovic, Z., Stary, P. &amp; Rezwani, A. (2007) Preliminary taxonomic study of the genus Praon (Hymenoptera: Braconidae: Aphidiinae) and its host associations in Iran. Journal of Entomological Society of Iran, 26 (2), 19 - 34." type="journal article" year="2007">
Rakhshani
<emphasis id="B967EAB8D777FFE0FDA0412AFD2D6D5F" box="[623,671,799,825]" italics="true" pageId="9" pageNumber="510">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EF824B5BD777FFE0FD3D4115FC0E6D5F" author="Nazari, Y. &amp; Zamani, A. A. &amp; Masoumi, S. M. &amp; Rakhshani, E. &amp; Petrovic, O. O. &amp; Tomanovic, S. &amp; Stary, P. &amp; Tomanovic, Z." box="[754,956,799,825]" pageId="9" pageNumber="510" pagination="559 - 584" refId="ref19628" refString="Nazari, Y., Zamani, A. A., Masoumi, S. M., Rakhshani, E., Petrovic, O. O., Tomanovic, S., Stary, P. &amp; Tomanovic, Z. (2012) Diversity and host associations of aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) in the farmlands of western Iran. Acta Entomologica Musei Nationalis Pragae, 52, 559 - 584." type="journal article" year="2012">
Nazari
<emphasis id="B967EAB8D777FFE0FC8D412AFCC86D5F" box="[834,890,799,825]" italics="true" pageId="9" pageNumber="510">et al.</emphasis>
2012
</bibRefCitation>
), and
<collectingCountry id="F304763AD777FFE0FC334115FBEC6D5F" box="[1020,1118,799,825]" name="Lebanon" pageId="9" pageNumber="510">Lebanon</collectingCountry>
(
<bibRefCitation id="EF824B5BD777FFE0FBA34115FAEB6D5C" author="Tremblay, E. &amp; Kawar, N. &amp; Barbagallo, S." box="[1132,1369,799,826]" pageId="9" pageNumber="510" pagination="19 - 32" refId="ref22419" refString="Tremblay, E., Kawar, N. &amp; Barbagallo, S. (1985) Aphids (Homoptera: Aphidoidea) and aphidiines (Hymenoptera: Braconidae) of Lebanon. Bolletino del Laboratoria di Entomologia Agraria ' Filippo Silvestri' Portici, 42, 19 - 32." type="journal article" year="1985">
Tremblay
<emphasis id="B967EAB8D777FFE0FB2F412AFAA56D5F" box="[1248,1303,799,825]" italics="true" pageId="9" pageNumber="510">et al.</emphasis>
1985
</bibRefCitation>
). This is the first record in
<collectingCountry id="F304763AD777FFE0FEB1414EFE736D38" box="[382,449,836,862]" name="Egypt" pageId="9" pageNumber="510">Egypt</collectingCountry>
in association with
<taxonomicName id="4C134D29D777FFE0FD6B414EFD406D3B" box="[676,754,835,861]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="faba">
<emphasis id="B967EAB8D777FFE0FD6B414EFD406D3B" box="[676,754,835,861]" italics="true" pageId="9" pageNumber="510">V. faba</emphasis>
</taxonomicName>
. It should be noted that in
<collectingCountry id="F304763AD777FFE0FBEC4149FBC86D3B" box="[1059,1146,835,861]" name="Algeria" pageId="9" pageNumber="510">Algeria</collectingCountry>
, it has been recorded as a parasitoid of
<taxonomicName id="4C134D29D777FFE0FEE1416DFDAF6DE7" box="[302,541,871,897]" class="Insecta" family="Aphididae" genus="Acyrthosiphon" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="pisum">
<emphasis id="B967EAB8D777FFE0FEE1416DFDAF6DE7" box="[302,541,871,897]" italics="true" pageId="9" pageNumber="510">Acyrthosiphon pisum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C134D29D777FFE0FDE3416DFD5C6DE7" box="[556,750,871,897]" class="Insecta" family="Aphididae" genus="Aphis" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="craccivora">
<emphasis id="B967EAB8D777FFE0FDE3416DFD5C6DE7" box="[556,750,871,897]" italics="true" pageId="9" pageNumber="510">Aphis craccivora</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C134D29D777FFE0FCE1416DFC076DE7" authorityName="Scopoli" authorityYear="1763" box="[814,949,871,897]" class="Insecta" family="Aphididae" genus="Aphis" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="fabae">
<emphasis id="B967EAB8D777FFE0FCE1416DFC076DE7" box="[814,949,871,897]" italics="true" pageId="9" pageNumber="510">Aphis fabae</emphasis>
</taxonomicName>
in association with other crops rather than with
<taxonomicName id="4C134D29D777FFE0FF1F4186FDA46DC3" authority="(Laamari et al. 2012)" baseAuthorityName="Laamari" baseAuthorityYear="2012" box="[208,534,907,933]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="510" phylum="Tracheophyta" rank="species" species="faba">
<emphasis id="B967EAB8D777FFE0FF1F4186FEAF6DC3" box="[208,285,907,933]" italics="true" pageId="9" pageNumber="510">V. faba</emphasis>
(
<bibRefCitation id="EF824B5BD777FFE0FEE34181FDBC6DC3" author="Laamari, M. &amp; Chaouche, S. T. &amp; Halimi, C. W. &amp; Benferhat, S. &amp; Abbas, S. B. &amp; Khenissa, N. &amp; Stary, P." box="[300,526,907,933]" pageId="9" pageNumber="510" pagination="161 - 176" refId="ref18584" refString="Laamari, M., Chaouche, S. T., Halimi, C. W., Benferhat, S., Abbas, S. B., Khenissa, N. &amp; Stary, P. (2012) A review of aphid parasitoids and their associations in Algeria (Hymenoptera: Braconidae: Aphidiinae; Hemiptera: Aphidoidea). African Entomology, 20 (1), 161 - 176. https: // doi. org / 10.4001 / 003.020.0120" type="journal article" year="2012">
Laamari
<emphasis id="B967EAB8D777FFE0FE5D4186FE786DC3" box="[402,458,907,933]" italics="true" pageId="9" pageNumber="510">et al.</emphasis>
2012
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>