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<document id="48A5BDDCEB11C023E5311F53480842D2" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FEF6FE82FDE6B9FAA4D2F957" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Cheiloneurus marcius Noyes 2023, sp.nov." docType="treatment" docUuid="C31AF8E2-B23B-4854-AF4A-15E321E0AEA6" docUuidSource="ZooBank" docVersion="1" lastPageNumber="292" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="290" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="97D604EDBE7C1A34112D43C400B59BCB">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="5E6F883145DB71B71D8C0EEE1A16A2EC">Noyes, John Stuart</mods:namePart>
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<mods:date id="8E41CC9B461B6E0C9529B627FBD91BC5">2023</mods:date>
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<subSubSection id="C309653AFEF6FE84FDE6B9FAA52EFA32" box="[516,830,1502,1527]" pageId="291" pageNumber="290" type="nomenclature">
<paragraph id="8BAC36B1FEF6FE84FDE6B9FAA52EFA32" blockId="291.[516,830,1502,1553]" box="[516,830,1502,1527]" pageId="291" pageNumber="290">
<heading id="D0E481DDFEF6FE84FDE6B9FAA52EFA32" bold="true" box="[516,830,1502,1527]" centered="true" fontSize="10" level="2" pageId="291" pageNumber="290" reason="2">
<emphasis id="B967EAA3FEF6FE84FDE6B9FAA52EFA32" bold="true" box="[516,830,1502,1527]" pageId="291" pageNumber="290">
<taxonomicName id="4C134D32FEF6FE84FDE6B9FAA4FCFA33" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[516,748,1502,1526]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="291" pageNumber="290" phylum="Arthropoda" rank="species" species="marcius" status="sp. nov.">
<emphasis id="B967EAA3FEF6FE84FDE6B9FAA4FCFA33" bold="true" box="[516,748,1502,1526]" italics="true" pageId="291" pageNumber="290">Cheiloneurus marcius</emphasis>
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<taxonomicNameLabel id="A25457D8FEF6FE84FD11B9FBA52EFA32" box="[755,830,1503,1527]" pageId="291" pageNumber="290" rank="species">sp.nov.</taxonomicNameLabel>
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<subSubSection id="C309653AFEF6FE84FDB7B9DDA4FCF9D4" box="[597,748,1529,1553]" pageId="291" pageNumber="290" type="description">
<paragraph id="8BAC36B1FEF6FE84FDB7B9DDA4FCF9D4" blockId="291.[516,830,1502,1553]" box="[597,748,1529,1553]" pageId="291" pageNumber="290">
(
<figureCitation id="13282A34FEF6FE84FDBEB9DDA4F5F9D4" box="[604,741,1529,1553]" captionStart-0="Figs 713-724" captionStart-1="Figs 725-736" captionStartId-0="788.[181,226,1526,1550]" captionStartId-1="789.[181,226,1526,1550]" captionTargetBox-0="[186,1163,228,1508]" captionTargetBox-1="[183,1163,224,1502]" captionTargetId-0="figure-9@788.[181,1163,223,1508]" captionTargetId-1="figure-10@789.[181,1163,223,1508]" captionTargetPageId-0="788" captionTargetPageId-1="789" captionText-0="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." captionText-1="Figs 725-736. Cheiloneurus marcius, HT (725) fore wing. Cheiloneurus castalia (726) apex of fore wing venation; (727) antenna; (728) ovipositor; (729) head, facial view; (730) fore wing. Cheiloneurus dipsas (731) antenna; (732) ovipositor; (733) apex of fore wing venation; (734) fore wing. Cheiloneurus medusa (735) head, facial view; (736) hypopygium." pageId="291" pageNumber="290">Figs 720-725</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEF6FE82FF57BA0BA771FAB8" lastPageId="293" lastPageNumber="292" pageId="291" pageNumber="290" type="diagnosis">
<paragraph id="8BAC36B1FEF6FE83FF57BA0BA476FCB4" blockId="291.[181,1165,1582,1687]" lastBlockId="292.[181,1165,222,881]" lastPageId="292" lastPageNumber="291" pageId="291" pageNumber="290">
DIAGNOSIS.
<emphasis id="B967EAA3FEF6FE84FEAEBA0AA788F983" box="[332,408,1582,1606]" italics="true" pageId="291" pageNumber="290">Female</emphasis>
(length about
<quantity id="4CEB9B54FEF6FE84FDCDBA0BA46BF982" box="[559,635,1583,1607]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="291" pageNumber="290" unit="mm" value="1.4">1.4mm</quantity>
): body generally orange, gaster mostly dark brown, paler areas with a weak metallic sheen, darker areas with a stronger metallic sheen; antenna (
<figureCitation id="13282A34FEF6FE84FB81BA6DA6CCF9B9" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="291" pageNumber="290">Fig. 722</figureCitation>
) with scape mostly pale orange, ventrally margined brown; pedicel very pale orange with a dark brown streak; funicle with proximally brown, distally orange; clava dark brown; posterior half of mesoscutum with dense silvery setae; fore and hind coxae white, mid coxa very pale orange; legs mostly white to pale orange; fore wing (Fg. 725) mostly infuscate with basal cell mostly hyaline, a small hyaline area at apex of venation which continues more broadly around wing apex to opposite margin; head (
<figureCitation id="13282A34FEF1FE83FDCCBD15A494FE8C" box="[558,644,305,329]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 720</figureCitation>
) about 5.3X as wide as frontovertex, in facial view slightly broader than long, subrectangular, genae weakly curved, hardly converging, but more strongly curved inwards near mouth; frontovertex with about 4 setae medially between anterior ocellus and scrobes; eye separated from scrobe by about 1.5X diameter of anterior ocellus, area between eye and scrobe completely smooth, scrobes deep and sharply margined; interantennal prominence with more than 100 conspicuous setae and dorsally rounded; mandible tridentate; antenna (
<figureCitation id="13282A34FEF1FE83FEF1BDF3A77AFE2A" box="[275,362,471,495]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 722</figureCitation>
) with scape about 1.8X as long as broad; funicle segments subquadrate; linear sensilla on F4-F6; clava 3-segmented, about as long as funicle, sensory area enlarged, extending about 0.5X along ventral surface of clava giving it a strongly obliquely truncate appearance; head width about equal to length of pedicel and flagellum combined; mesoscutum (
<figureCitation id="13282A34FEF1FE83FC24BE0DA20CFD87" box="[966,1052,553,578]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 721</figureCitation>
) anteriorly with coarse lineolate sculpture, posteriorly with polygonally reticulate sculpture; scutellum with a distinct, apical tuft of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FEF1FE83FCA8BE45A5B1FDBD" box="[842,929,608,633]" captionStart="Figs 725-736" captionStartId="789.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1502]" captionTargetId="figure-10@789.[181,1163,223,1508]" captionTargetPageId="789" captionText="Figs 725-736. Cheiloneurus marcius, HT (725) fore wing. Cheiloneurus castalia (726) apex of fore wing venation; (727) antenna; (728) ovipositor; (729) head, facial view; (730) fore wing. Cheiloneurus dipsas (731) antenna; (732) ovipositor; (733) apex of fore wing venation; (734) fore wing. Cheiloneurus medusa (735) head, facial view; (736) hypopygium." pageId="292" pageNumber="291">Fig. 725</figureCitation>
) about 2.6X as long as broad; parastigma slightly downcurved; costal cell dorsally naked or with a single apical seta, ventrally more or less with a complete line of setae; area below proximal part of parastigma with a line of about 8-10, paler setae that curves towards fore wing base; apices of postmarginal and stigmal veins connected by a naked, hyaline area that extends a little way into disc (
<figureCitation id="13282A34FEF1FE83FC19BEEBA244FD22" box="[1019,1108,719,743]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 724</figureCitation>
); seta at apex of postmarginal vein about 0.8X as long as marginal vein; mid tibial spur slightly longer than basitarsus; propodeum (
<figureCitation id="13282A34FEF1FE83FE3EBF22A423FCDB" box="[476,563,774,798]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 721</figureCitation>
) with only about five setae adjacent to spiracle; syntergum hardly longer than mid tibia; ovipositor (
<figureCitation id="13282A34FEF1FE83FDACBF06A4B4FCFF" box="[590,676,802,826]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 723</figureCitation>
) about 3.2X gonostylus or about 1.7X as long as mid tibia; gonostylus about 1.4X as long as mid tibial spur; exserted part of ovipositor about 0.5X as long as mid tibial spur.
<emphasis id="B967EAA3FEF1FE83FE5DBF7DA7E3FCB4" box="[447,499,857,881]" italics="true" pageId="292" pageNumber="291">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FEF1FE83FF57BFB4A293FC6D" blockId="292.[181,1165,912,1681]" box="[181,1155,912,936]" pageId="292" pageNumber="291">
<materialsCitation id="3B7B3CECFEF1FE83FF57BFB4A26EFC6D" box="[181,1150,912,936]" collectionCode="CPD" pageId="292" pageNumber="291" specimenCount="1" typeStatus="holotype">
Female (
<typeStatus id="54A88813FEF1FE83FEEFBFB4A77CFC6D" box="[269,364,912,936]" pageId="292" pageNumber="291" type="holotype">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FEF1FE83FD40BFB4A4E9FC6D" box="[674,761,912,936]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="292" pageNumber="291" unit="mm" value="1.37">1.37mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FEF1FE83FC3DBFB4A223FC6D" box="[991,1075,912,936]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="292" pageNumber="291" unit="mm" value="1.29">1.29mm</quantity>
(
<collectionCode id="ED02AE74FEF1FE83FBA3BFB4A268FC6D" box="[1089,1144,912,936]" pageId="292" pageNumber="291">CPD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36B1FEF1FE82FF39BF88A5E5FEA0" blockId="292.[181,1165,912,1681]" lastBlockId="293.[181,1165,222,1350]" lastPageId="293" lastPageNumber="292" pageId="292" pageNumber="291">
Head generally orange; occiput pale orange, slightly dusky near foramen; an oblique, brown stripe across temple towards torulus; frontovertex with a weak metallic blue sheen, stronger metallic green between posterior ocelli, purple immediately above scrobes; temple with a slight metallic blue and purple sheen; shining metallic green and purple between eye and scrobe; gena relatively dull green and brassy with some coppery and purple reflections; top of scrobes delimited by a slightly curved, metallic green band; frontovertex with inconspicuous pale brown setae along inner eye margins; a line of slightly more conspicuous, translucent, white setae below eye; interantennal prominence clothed with dense, conspicuous, silvery setae; maxillary palpus yellow; antenna (
<figureCitation id="13282A34FEF1FE83FE81B8ACA7A8FB64" box="[355,440,1160,1185]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="292" pageNumber="291">Fig. 722</figureCitation>
) with radicle orange-brown with a narrow brown apical ring; scape pale orange, apical one-third white, extreme apex brown, ventral margin dark brown, more broadly so proximally, upper margin pale brown proximally, a transverse brown band between apical white area and proximal pale orange area; pedicel very pale orange with a dark brown streak along inner surface; F1 brown, F2-F5 brown dorsally mixed yellow, distal segments more yellow, F6 yellow, clava dark brown; pronotum orange with weak brassy and coppery reflections, neck and area above coxa brown; prosternum pale orange, brown anterior to coxa; mesoscutum orange in anterior half with some very weak coppery and brassy reflections, clothed in relatively sparse golden brown and dark brown setae, posterior half and side brown with a moderately strong, dark blue sheen with some purple reflections, clothed in conspicuous, relatively dense silvery setae, posterior margin purple-brown with a purple sheen; tegula orange with apex orange-brown; axilla and scutellum orange, clothed with golden brown and brown setae, subapical tuft on scutellum black; metanotum orange-brown; mesopleuron orange with a weak brassy sheen, slightly dusky posteriorly with a weak purple and brassy sheen; mesosternum orange; fore coxa and proximal half of fore femur white, fore leg otherwise pale orange; mid coxa slightly dusky, very pale orange; mid femur white in proximal half, apex pale orange, tibia mostly pale orange, white at base and an incomplete, brown subbasal ring, spur and tarsus pale orange; hind coxa white; hind femur pale orange, slightly dusky along margins towards apex, tibia and tarsus pale orange; fore wing (
<figureCitation id="13282A34FEF1FE83FE95BA5DA7DDF954" box="[375,461,1657,1681]" captionStart="Figs 725-736" captionStartId="789.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1502]" captionTargetId="figure-10@789.[181,1163,223,1508]" captionTargetPageId="789" captionText="Figs 725-736. Cheiloneurus marcius, HT (725) fore wing. Cheiloneurus castalia (726) apex of fore wing venation; (727) antenna; (728) ovipositor; (729) head, facial view; (730) fore wing. Cheiloneurus dipsas (731) antenna; (732) ovipositor; (733) apex of fore wing venation; (734) fore wing. Cheiloneurus medusa (735) head, facial view; (736) hypopygium." pageId="292" pageNumber="291">Fig. 725</figureCitation>
) mostly infuscate, hyaline towards base, along anterior margin distad of venation and at apex; submarginal vein pale orange, venation otherwise brown; propodeum orange medially, brown laterally with a dark metallic blue and purple sheen, side metallic blue, about 3 or 4 long, translucent, white setae in a line outside spiracle; gaster dark brown with a coppery sheen, Gt1 largely with a metallic blue and green sheen, side and venter largely with green and blue reflections, syntergum pale orange-brown; gonostylus pale orange.
</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF39BD4CA4A6FC90" blockId="293.[181,1165,222,1350]" pageId="293" pageNumber="292">
Head (
<figureCitation id="13282A34FEF0FE82FEFCBD4CA764FE45" box="[286,372,360,384]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 720</figureCitation>
) about 5.3X as wide as frontovertex, in profile 1.5X as high as deep, anteriorly quite straight from top of scrobes to mouth margin, interantennal prominence slightly protuberant; occipital margin sharp, not carinate; ocelli forming an angle of about 50°; frontovertex quite shiny, with shallow, polygonally reticulate sculpture of mesh size much smaller than diameter of facet; narrowest point between eye and scrobe about 1.5X diameter of anterior ocellus and completely smooth; temple quite smooth, but with shallow, irregular, elongate, reticulate sculpture; gena with slightly deeper more regular, polygonally reticulate sculpture; scrobes deep, ∩-shaped, meeting, sharply margined; interantennal prominence dorsally rounded and with shallow, polygonally reticulate sculpture; antenna as in
<figureCitation id="13282A34FEF0FE82FDEEBE61A473FD98" box="[524,611,581,605]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 722</figureCitation>
; scape strongly broadened and flattened, about 1.8X as long as broad, broadest in middle; funicle segments subquadrate, larger and broader distad; linear sensilla present only on F4-F6; clava clearly broadened, about as long as funicle, sutures slightly oblique, sensory area enlarged, extending ventrally nearly half length of clava, apex appearing strongly obliquely truncate; eye reaching occipital margin, appearing naked, but clothed in sparse, inconspicuous, fine, very short setae, each much shorter than diameter of facet; inner eye margins subparallel, converging slightly anteriorly; malar sulcus absent; clypeal margin very slightly convex medially; mandible with three acute; apical segment of maxillary palpus not enlarged, about as long as radicle. Relative measurements: HW 63,
<collectingRegion id="49D7F853FEF0FE82FCE6BF06A539FCFC" box="[772,809,802,825]" country="Swaziland" name="Hhohho" pageId="293" pageNumber="292">HH</collectingRegion>
60, FV 12, POL 5.5, OOL 0, OCL 5, AOL 7, EL 41, EW 32, MS 30, SL 31.5, SW 18.
</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF39BF7DA410FB79" blockId="293.[181,1165,222,1350]" pageId="293" pageNumber="292">
Thorax (
<figureCitation id="13282A34FEF0FE82FED1BF7DA79BFCB4" box="[307,395,857,881]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 721</figureCitation>
) with pronotum short and hardly exposed behind head, about 0.6X as long as mesoscutum, pronotum with polygonally reticulate sculpture that is a little deeper than that on frontovertex; mesoscutum (
<figureCitation id="13282A34FEF0FE82FE0FBFB4A454FC6D" box="[493,580,912,936]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 721</figureCitation>
) in anterior half with coarse lineolate sculpture, posterior half with polygonally reticulate sculpture of similar depth; axilla and scutellum with polygonally reticulate sculpture similar to that on pronotum, but perhaps slightly coarser, especially on scutellum where it is longitudinally elongate laterally; visible part of mesoscutum about 2.4X as broad as long; scutellum about as long as broad with a distinct subapical tuft, setae mostly about 0.4X as long as scutellum; mid tibia with a distinct external carina for about half its length; hind femur about 4.2X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FEF0FE82FC5FB812A205FB8B" box="[957,1045,1078,1102]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Figs 724</figureCitation>
,
<figureCitation id="13282A34FEF0FE82FBFDB811A258FB88" box="[1055,1096,1077,1101]" captionStart="Figs 725-736" captionStartId="789.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1502]" captionTargetId="figure-10@789.[181,1163,223,1508]" captionTargetPageId="789" captionText="Figs 725-736. Cheiloneurus marcius, HT (725) fore wing. Cheiloneurus castalia (726) apex of fore wing venation; (727) antenna; (728) ovipositor; (729) head, facial view; (730) fore wing. Cheiloneurus dipsas (731) antenna; (732) ovipositor; (733) apex of fore wing venation; (734) fore wing. Cheiloneurus medusa (735) head, facial view; (736) hypopygium." pageId="293" pageNumber="292">725</figureCitation>
; costal cell with 2 or 3 setae dorsally at apex; apices of postmarginal and stigmal veins connected by a distinct, naked, hyaline streak (
<figureCitation id="13282A34FEF0FE82FE19B849A447FB40" box="[507,599,1133,1157]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 724</figureCitation>
); propodeum medially (
<figureCitation id="13282A34FEF0FE82FCB2B849A5B9FB40" box="[848,937,1133,1157]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 721</figureCitation>
) about 0.12X as long as scutellum and virtually smooth, setae on side conspicuous. Relative measurements: FWL 66, FWW 25; HWL 50, HWW 13.5.
</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF39B8E4A4A3FAEF" blockId="293.[181,1165,222,1350]" pageId="293" pageNumber="292">
Gaster without “gland-like” structures on Gt1 or Gt5; hypopygium reaching about 0.4X to apex; syntergum about 0.9X as long as mid tibia, with apex acute; ovipositor (
<figureCitation id="13282A34FEF0FE82FC38B8FFA221FB36" box="[986,1073,1243,1267]" captionStart="Figs 713-724" captionStartId="788.[181,226,1526,1550]" captionTargetBox="[186,1163,228,1508]" captionTargetId="figure-9@788.[181,1163,223,1508]" captionTargetPageId="788" captionText="Figs 713-724. Cheiloneurus santaros, HT (713) ovipositor. Cheiloneurus simus (714) apex of fore wing venation; (715) fore wing; (716) head, facial view; (717) ovipositor; (718) antenna; (719) mesosoma, dorsal view. Cheiloneurus marcius, HT (720) head, facial view; (721) mesosoma, dorsal view; (722) antenna (a, outer aspect of clava; b, inner aspect of clava); (723) ovipositor; (724) apex of fore wing venation." pageId="293" pageNumber="292">Fig. 723</figureCitation>
) slightly exserted, the exserted part about half as long as mid tibial spur or nearly 0.2X as long as mid tibia. Relative measurements: OL 55, GL 17 [MT 32.5].
</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF39B90AA457FA83" blockId="293.[181,1165,222,1350]" box="[219,583,1326,1350]" pageId="293" pageNumber="292">
Variation. Only
<typeStatus id="54A88813FEF0FE82FE9CB90AA7C7FA83" box="[382,471,1326,1350]" pageId="293" pageNumber="292" type="holotype">holotype</typeStatus>
examined.
</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF57B941A771FAB8" blockId="293.[181,353,1381,1405]" box="[181,353,1381,1405]" pageId="293" pageNumber="292">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFEF0FE82FF57B9B8A7F9FA71" box="[181,489,1436,1460]" pageId="293" pageNumber="292" type="distribution">
<paragraph id="8BAC36B1FEF0FE82FF57B9B8A7F9FA71" blockId="293.[181,489,1436,1460]" box="[181,489,1436,1460]" pageId="293" pageNumber="292">
DISTRIBUTION.
<collectingCountry id="F3047621FEF0FE82FE91B9B8A7F5FA71" box="[371,485,1436,1460]" name="Costa Rica" pageId="293" pageNumber="292">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEF0FE82FF57B9F0A76EFA2E" box="[181,382,1491,1516]" pageId="293" pageNumber="292" type="biology_ecology">
<paragraph id="8BAC36B1FEF0FE82FF57B9F0A76EFA2E" blockId="293.[181,382,1491,1516]" box="[181,382,1491,1516]" pageId="293" pageNumber="292">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFEF0FE82FF57BA29A522F99F" pageId="293" pageNumber="292" type="materials_examined">
<paragraph id="8BAC36B1FEF0FE82FF57BA29A7BDF9E7" blockId="293.[181,1164,1549,1626]" box="[181,429,1549,1570]" pageId="293" pageNumber="292">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FEF0FE82FF39BA0DA522F99F" blockId="293.[181,1164,1549,1626]" pageId="293" pageNumber="292">
Type material.
<materialsCitation id="3B7B3CECFEF0FE82FE81BA0DA44BF99F" collectorName="Send. Los Gigantes, LS" country="Costa Rica" elevation="1460" location="Puntarenas" pageId="293" pageNumber="292" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54A88813FEF0FE82FE81BA0DA7A7F9FA" box="[355,439,1577,1599]" pageId="293" pageNumber="292" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FEF0FE82FE38BA0CA475F9FB" bold="true" box="[474,613,1576,1598]" pageId="293" pageNumber="292">
<collectingCountry id="F3047621FEF0FE82FE38BA0CA475F9FB" box="[474,613,1576,1598]" name="Costa Rica" pageId="293" pageNumber="292">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FEF0FE82FD92BA0DA4CEF9FB" bold="true" box="[624,734,1577,1598]" pageId="293" pageNumber="292">
<collectingRegion id="49D7F853FEF0FE82FD92BA0DA4CEF9FB" box="[624,734,1577,1598]" country="Costa Rica" name="Puntarenas" pageId="293" pageNumber="292">Puntarenas</collectingRegion>
</emphasis>
, Est. Altamira,
<collectorName id="26E65367FEF0FE82FC8EBA0DA252F9FB" box="[876,1090,1577,1599]" pageId="293" pageNumber="292">Send. Los Gigantes, LS</collectorName>
321800 572100,
<quantity id="4CEB9B54FEF0FE82FEE7BA60A756F99F" box="[261,326,1604,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="293" pageNumber="292" unit="m" value="1460.0">
<elevation id="003ED182FEF0FE82FEE7BA60A756F99F" box="[261,326,1604,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="293" pageNumber="292" unit="m" value="1460.0">1460m</elevation>
</quantity>
, #62126, iii-iv.2001 (
<taxonomicName id="4C134D32FEF0FE82FDEBBA60A445F99F" box="[521,597,1604,1626]" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="293" pageNumber="292" phylum="Arthropoda" rank="species" species="rubi">D. Rubí</taxonomicName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFEF0FE82FD86BA60A53EF99F" box="[612,814,1604,1626]" collectionCode="NHMUK" country="Costa Rica" pageId="293" pageNumber="292" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54A88813FEF0FE82FD86BA60A4A8F99F" box="[612,696,1604,1626]" pageId="293" pageNumber="292" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FEF0FE82FD37BA61A53EF99F" box="[725,814,1605,1626]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="293" pageNumber="292" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEF0FE82FF57BA5EA4D2F957" box="[181,706,1658,1682]" pageId="293" pageNumber="292" type="discussion">
<paragraph id="8BAC36B1FEF0FE82FF57BA5EA4D2F957" blockId="293.[181,706,1658,1682]" box="[181,706,1658,1682]" pageId="293" pageNumber="292">
COMMENTS. See comments under
<taxonomicName id="4C134D32FEF0FE82FDC9BA5EA475F954" authorityName="Noyes" authorityYear="2023" box="[555,613,1658,1681]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="293" pageNumber="292" phylum="Arthropoda" rank="species" species="simus">
<emphasis id="B967EAA3FEF0FE82FDC9BA5EA475F954" box="[555,613,1658,1681]" italics="true" pageId="293" pageNumber="292">simus</emphasis>
</taxonomicName>
(p. 290).
</paragraph>
</subSubSection>
</treatment>
</document>