224 lines
38 KiB
XML
224 lines
38 KiB
XML
<document id="35B6BB92372E077F2B2F3641C3172919" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FEDCFEADFDE4BCFAA4C7FA0A" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus mantua Noyes 2023, sp.nov." docType="treatment" docUuid="58B9D290-D81F-4833-8F2B-81A6AE6DD73E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="265" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="264" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="250B1ED91F8D4DA045AB6EA38672A9A4">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="79F55212B48B41A4894ADE60525085FC">Noyes, John Stuart</mods:namePart>
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<mods:title id="1F9D98D1C2B83092FCF2A0495B3F36D5">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="993EE24CAEF090ED34DEF85F2FF1E256">2023</mods:date>
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<mods:title id="81B2CF54F69B1F52015D605366228C05">Oxford, England</mods:title>
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<mods:number id="F1FD84E80545704D1D5122332442E9C9">2023-06-30</mods:number>
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<subSubSection id="C309653AFEDCFEAEFDE4BCFAA52CFF33" box="[518,828,222,246]" pageId="265" pageNumber="264" type="nomenclature">
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<paragraph id="8BAC36B1FEDCFEAEFDE4BCFAA52CFF33" blockId="265.[518,828,222,273]" box="[518,828,222,246]" pageId="265" pageNumber="264">
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<emphasis id="B967EAA3FEDCFEAEFDE4BCFAA52CFF33" bold="true" box="[518,828,222,246]" pageId="265" pageNumber="264">
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<taxonomicName id="4C134D32FEDCFEAEFDE4BCFAA4FAFF30" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[518,746,222,246]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="264" phylum="Arthropoda" rank="species" species="mantua" status="sp.nov.">
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<emphasis id="B967EAA3FEDCFEAEFDE4BCFAA4FAFF30" bold="true" box="[518,746,222,246]" italics="true" pageId="265" pageNumber="264">Cheiloneurus mantua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25457D8FEDCFEAEFD13BCFAA52CFF33" box="[753,828,222,246]" pageId="265" pageNumber="264" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFEDCFEAEFDFBBCDDA538FED4" box="[537,808,248,273]" pageId="265" pageNumber="264" type="description">
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<paragraph id="8BAC36B1FEDCFEAEFDFBBCDDA538FED4" blockId="265.[518,828,222,273]" box="[537,808,248,273]" pageId="265" pageNumber="264">
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(
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<figureCitation id="13282A34FEDCFEAEFDC2BCDDA4BBFED5" box="[544,683,248,273]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Figs 656-660</figureCitation>
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; Hab. E 99)
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFEDCFEADFF57BD0AA771FB68" lastPageId="266" lastPageNumber="265" pageId="265" pageNumber="264" type="diagnosis">
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<paragraph id="8BAC36B1FEDCFEAEFF57BD0AA575FC7A" blockId="265.[181,1165,302,959]" pageId="265" pageNumber="264">
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DIAGNOSIS.
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<emphasis id="B967EAA3FEDCFEAEFEADBD0AA78BFE83" box="[335,411,302,326]" italics="true" pageId="265" pageNumber="264">Female</emphasis>
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(length about
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<quantity id="4CEB9B54FEDCFEAEFDD8BD0AA4BDFE83" box="[570,685,302,326]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="265" pageNumber="264" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5-1.9mm</quantity>
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): body generally dark brown to black with a weak to moderate metallic sheen; antenna (
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<figureCitation id="13282A34FEDCFEAEFD85BD6CA4ADFEA5" box="[615,701,328,352]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 658</figureCitation>
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) with scape pale orange, dorsal margin brown proximally; pedicel mostly pale orange, proximally dark brown; funicle pale orange; clava dark brown; mesoscutum with uniform, silvery setae; coxae white; hind tibia mostly pale orange, legs otherwise white to pale orange with a few brown areas; fore wing (
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<figureCitation id="13282A34FEDCFEAEFC88BDB3A5D2FE6A" box="[874,962,407,431]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 657</figureCitation>
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) almost completely hyaline but weakly infuscate immediately below marginal and stigmal veins, a hint of a weakly infuscate median streak in apical half; head (
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<figureCitation id="13282A34FEDCFEAEFD98BDE8A4C0FE21" box="[634,720,460,484]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 656</figureCitation>
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) about 4X as wide as frontovertex, in facial view nearly 1.1X as broad as long, subcircular, gena weakly and evenly curved; frontovertex naked medially below anterior ocellus; area between eye and scrobe slightly narrower than diameter of anterior ocellus, with distinct sculpture, scrobal margin rounded; interantennal prominence dorsally rounded with only a few long setae; mandible with two long, sharp lower teeth and a very short upper tooth; antenna (
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<figureCitation id="13282A34FEDCFEAEFDE4BE74A44EFDAD" box="[518,606,592,616]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 658</figureCitation>
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) with scape about 4.5X as long as broad; funicle with F1 subquadrate, other segments strongly transverse, becoming larger distad; head about as wide as length of pedicel and flagellum combined; linear sensilla on F5-F6; clava 3-segmented, slightly longer than funicle, sensory area large and forming an oblique truncation, extending about 0.5X along ventral margin; mesoscutum (
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<figureCitation id="13282A34FEDCFEAEFDD2BE9EA499FD17" box="[560,649,698,722]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 660</figureCitation>
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) with polygonally reticulate sculpture; scutellum with a distinct, but very short apical tuft of setae; wings fully developed; fore wing (
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<figureCitation id="13282A34FEDCFEAEFBCEBEF0A295FD29" box="[1068,1157,724,748]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 657</figureCitation>
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) about 2.5X as long as broad; parastigma not curved downwards; costal cell dorsally with 2 or 3 setae at apex and a single, complete line of setae ventrally; area below proximal part of parastigma with a group of about 15 setae; apices of postmarginal and stigmal veins (
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<figureCitation id="13282A34FEDCFEAEFC7EBF07A5E2FCFE" box="[924,1010,803,827]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 659</figureCitation>
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) not connected by a naked hyaline area; apical bristle of postmarginal vein about 0.4X as long as marginal vein; mid tibial spur very nearly as long as basitarsus; propodeum (
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<figureCitation id="13282A34FEDCFEAEFCFEBF7CA562FCB5" box="[796,882,856,880]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 660</figureCitation>
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) with about 15 setae around each spiracle, not extending down side; syntergum about as long as mid tibia; ovipositor about 3.2X gonostylus or 1.6X mid tibia; gonostylus about 1.8X as long as mid tibial spur; exserted part of ovipositor about 0.4X as long as mid tibial spur.
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<emphasis id="B967EAA3FEDCFEAEFD5CBF83A4E2FC7A" box="[702,754,935,959]" italics="true" pageId="265" pageNumber="264">Male</emphasis>
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: unknown.
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</paragraph>
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<paragraph id="8BAC36B1FEDCFEAEFF57BFF8A293FC31" blockId="265.[181,1165,988,1699]" box="[181,1155,988,1012]" pageId="265" pageNumber="264">
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Female (
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<typeStatus id="54A88813FEDCFEAEFEECBFF8A77DFC31" box="[270,365,988,1012]" pageId="265" pageNumber="264">holotype</typeStatus>
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): length, including ovipositor,
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<quantity id="4CEB9B54FEDCFEAEFD40BFF8A4E8FC31" box="[674,760,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="265" pageNumber="264" unit="mm" value="1.84">1.84mm</quantity>
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; excluding ovipositor,
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<quantity id="4CEB9B54FEDCFEAEFC3DBFF8A223FC31" box="[991,1075,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="265" pageNumber="264" unit="mm" value="1.77">1.77mm</quantity>
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||
(CPD).
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||
</paragraph>
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||
<paragraph id="8BAC36B1FEDCFEAEFF39BFD2A29DF967" blockId="265.[181,1165,988,1699]" pageId="265" pageNumber="264">
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||
Head black with weak, blue and purple reflections on frontovertex; ocellar area more strongly purple; scrobal area mostly with a stronger blue-green sheen, mixed with purple, especially on interantennal prominence between toruli; mouth margin narrowly purplish; gena and temple with a brighter, slightly brassy metallic green sheen, strongest immediately below eye, some coppery reflections on lower gena; frontovertex with inconspicuous dark brown setae; antenna (Fig,. 658) with radicle pale orange, apex with a dark brown ring; scape pale orange, proximally with a narrow, dark brown ring, dorsal margin brown in proximal two-thirds or so; pedicel mostly pale orange, proximally dark brown; funicle pale orange; clava dark brown; thorax mostly dark brown to black; pronotum coppery with green and brassy reflections; mesoscutum clothed with uniform, translucent silvery setae and with a metallic green to blue-green sheen mixed with coppery and brassy reflections, posterior margin coppery purple; tegula dark brown; axilla dull, dark purple-brown with coppery reflections; scutellum noticeably less shiny than mesoscutum, but with blue-green and coppery reflections, clothed with dark brown setae, extreme sides and apex shining green mixed coppery; metanotum with slight purple sheen; mesopleuron in posterior half with a coppery-purple and brassy sheen, anterior half mixed blue and brassy, these narrowly separated by a coppery, dark blue and violet sheen; fore coxa white; fore femur white to pale yellow, with a subapical, brown spot externally; fore tibia pale orange with a very small, subbasal, brown spot externally; mid coxa white; mid femur white to pale yellow with a subapical brown spot externally, tibia pale orange with a subbasal brown spot externally; hind coxa white; hind femur white in proximal half or so, otherwise pale orange-brown, tibia mostly pale orange with an incomplete, narrow, subbasal brown ring; tarsi pale orange, pretarsi slightly dusky; fore wing (
|
||
<figureCitation id="13282A34FEDCFEAEFC05BA22A22CF9DB" box="[999,1084,1542,1566]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="265" pageNumber="264">Fig. 657</figureCitation>
|
||
) almost completely hyaline but weakly infuscate immediately below marginal and stigmal veins, a hint of a weakly infuscate median streak distad of pale brown venation; propodeum dark purple-brown; sides metallic green with a few conspicuous silvery setae in spiracular area; gaster dark purple-brown, Gt1 largely metallic blue-green; remainder of dorsum of gaster slightly coppery, mixed purple and brassy, sides of gaster towards apex and outer plates of ovipositor mixed metallic
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFBBCBCA6A29CFF5A" blockId="266.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="266" pageNumber="265">265</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF57BCFBA58AFF32" blockId="266.[181,1165,222,1144]" box="[181,922,222,247]" pageId="266" pageNumber="265">green, coppery and brassy; gonostylus dark brown, extreme apex orange.</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39BCDDA79EFDAD" blockId="266.[181,1165,222,1144]" pageId="266" pageNumber="265">
|
||
Head (
|
||
<figureCitation id="13282A34FEDFFEADFEC1BCDDA76EFED4" box="[291,382,249,273]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="266" pageNumber="265">Fig. 656</figureCitation>
|
||
) about 4X as wide as frontovertex, not subtriangular in profile, occipital margin carinate, occiput without a median groove above foramen; ocelli forming an angle of about 50°; frontovertex slightly shiny, with conspicuous reticulate sculpture of mesh size generally very slightly smaller than diameter of eye facet; scrobal area and mouth margin with shallower sculpture; gena and temple with more elongate sculpture of larger mesh; scrobes shallow, meeting dorsally, dorsal and lateral margins weakly rounded; antenna as in
|
||
<figureCitation id="13282A34FEDFFEADFC4EBD59A213FE50" box="[940,1027,381,405]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="266" pageNumber="265">Fig. 658</figureCitation>
|
||
; scape hardly broadened and flattened, about 4.5X as long as broad; F1 subquadrate, remaining segments strongly transverse and broadest distad; clava moderately broad, about as long as funicle, sutures oblique, sensory area enlarged, extending about one-third along clava ventrally and giving it an obliquely truncate appearance; eye reaching occipital margin, clothed with inconspicuous setae, each of which is about as long as diameter of facet; inner eye margins not converging anteriorly; malar sulcus absent; mandible with two long, acute, lower teeth and a short upper tooth. Relative measurements: HW 71,
|
||
<collectingRegion id="49D7F853FEDFFEADFE4FBE12A7C2FD88" box="[429,466,566,589]" country="Swaziland" name="Hhohho" pageId="266" pageNumber="265">HH</collectingRegion>
|
||
66, FV 18, OD 6, POL 7.5, OOL 1, OCL 6, AOL 9, EL 44, EW 38, MS 28, SL 35, SW 8.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39BE4EA753FC60" blockId="266.[181,1165,222,1144]" pageId="266" pageNumber="265">
|
||
Thorax (
|
||
<figureCitation id="13282A34FEDFFEADFEDABE4EA783FD47" box="[312,403,618,642]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="266" pageNumber="265">Fig. 660</figureCitation>
|
||
) with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size subequal to diameter of facet; sculpture on mesoscutum generally of similar depth to that on pronotum, but more regular and of smaller mesh; sculpture on axilla similar to that on mesoscutum, but slightly deeper; scutellum with conspicuously deeper, polygonally reticulate sculpture; apex and sides narrowly completely smooth; visible part of mesoscutum about 1.7X as broad as long; scutellum slightly broader than long, with a very short subapical tuft, setae about one-quarter as long as scutellum; hind femur about 3.7X as long as broad; fore wing with venation and setation as in
|
||
<figureCitation id="13282A34FEDFFEADFE93BF07A412FCFE" box="[369,514,803,827]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="266" pageNumber="265">Figs 657, 659</figureCitation>
|
||
; costal cell with only 3 setae dorsally at apex; no naked streak present between apices of postmarginal and stigmal veins (
|
||
<figureCitation id="13282A34FEDFFEADFCE7BF19A54DFC90" box="[773,861,829,853]" captionStart="Figs 650-662" captionStartId="783.[181,226,1526,1550]" captionTargetBox="[184,1159,226,1508]" captionTargetId="figure-10@783.[181,1163,223,1508]" captionTargetPageId="783" captionText="Figs 650-662. Cheiloneurus mirabel, HT – (650) antenna (a, outer aspect of clava; b, inner aspect of clava) (clava slightly collapsed); (651) apex of fore wing venation; (652) ovipositor); 653) mesosoma, dorsal view; (654) head, facial view; (655) fore wing. Cheiloneurus mantua – (656) head, facial view; (657) fore wing; (658) antenna (a, outer aspect of clava; b, inner aspect of clava); (659) apex of fore wing venation; (660) mesosoma, dorsal view. Cheiloneurus hamadryas – (661) mesosoma, dorsal view, variant; (662) mesosoma, dorsal view, variant." pageId="266" pageNumber="265">Fig. 659</figureCitation>
|
||
); propodeum with moderately dense setae in spiracular area, side naked, medially less than 0.1X as long as scutellum and with some shallow polygonally reticulate sculpture. Relative measurements: FWL 73, FWW 29; HWL 58, HWW 16.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39BF83A731FC36" blockId="266.[181,1165,222,1144]" pageId="266" pageNumber="265">Gaster with hypopygium reaching about half-way to apex; syntergum about as long as mid tibia and with apex acute; ovipositor slightly exserted, the exserted part about 0.4X as long as mid tibial spur.</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39BFD3A7B6FB86" blockId="266.[181,1165,222,1144]" pageId="266" pageNumber="265">
|
||
<typeStatus id="54A88813FEDFFEADFF39BFD3A727FBCA" box="[219,311,1015,1039]" pageId="266" pageNumber="265">Paratype</typeStatus>
|
||
. Funicle with linear sensilla present only on F5-F6; gaster without “gland-like” structures on Gt1 or Gt5; gonostylus about 1.8X as long as mid tibial spur. Relative measurements: OL 54, GL 17 [MT 33].
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39B861A70FFBBD" blockId="266.[181,1165,222,1144]" pageId="266" pageNumber="265">
|
||
Variation. Females vary in overall length from
|
||
<quantity id="4CEB9B54FEDFFEADFD5BB862A554FB9B" box="[697,836,1094,1118]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" metricValueMax="1.84" metricValueMin="1.48" pageId="266" pageNumber="265" unit="mm" value="1.66" valueMax="1.84" valueMin="1.48">1.48-1.84mm</quantity>
|
||
, otherwise little in
|
||
<typeStatus id="54A88813FEDFFEADFBE5B862A223FB9B" box="[1031,1075,1094,1118]" pageId="266" pageNumber="265">type</typeStatus>
|
||
material examined.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF57B8B1A771FB68" blockId="266.[181,353,1173,1197]" box="[181,353,1173,1197]" pageId="266" pageNumber="265">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEDFFEADFF57B8EEA7F9FB24" box="[181,489,1225,1250]" pageId="266" pageNumber="265" type="distribution">
|
||
<paragraph id="8BAC36B1FEDFFEADFF57B8EEA7F9FB24" blockId="266.[181,489,1225,1250]" box="[181,489,1225,1250]" pageId="266" pageNumber="265">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F3047621FEDFFEADFE91B8EEA7F5FB24" box="[371,485,1225,1250]" name="Costa Rica" pageId="266" pageNumber="265">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEDFFEADFF57B8DAA76EFAD3" box="[181,382,1278,1302]" pageId="266" pageNumber="265" type="biology_ecology">
|
||
<paragraph id="8BAC36B1FEDFFEADFF57B8DAA76EFAD3" blockId="266.[181,382,1278,1302]" box="[181,382,1278,1302]" pageId="266" pageNumber="265">HOSTS. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEDFFEADFF57B911A5A8FA5F" pageId="266" pageNumber="265" type="materials_examined">
|
||
<paragraph id="8BAC36B1FEDFFEADFF57B911A7BDFA8F" blockId="266.[181,1165,1333,1434]" box="[181,429,1333,1354]" pageId="266" pageNumber="265">MATERIAL EXAMINED.</paragraph>
|
||
<paragraph id="8BAC36B1FEDFFEADFF39B974A5A8FA5F" blockId="266.[181,1165,1333,1434]" pageId="266" pageNumber="265">
|
||
Type material.
|
||
<materialsCitation id="3B7B3CECFEDFFEADFE88B96BA6E9FABA" collectingDate="1990-03-18" collectingDateMax="1990-03-22" collectingDateMin="1990-03-18" collectionCode="MT, YPT" collectorName="J. S. Noyes" country="Costa Rica" location="Las Cruces" pageId="266" pageNumber="265" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||
<typeStatus id="54A88813FEDFFEADFE88B96BA7AEFAA0" box="[362,446,1359,1381]" pageId="266" pageNumber="265">Holotype</typeStatus>
|
||
E:
|
||
<emphasis id="B967EAA3FEDFFEADFE0BB96BA467FAA0" bold="true" box="[489,631,1359,1381]" pageId="266" pageNumber="265">
|
||
<collectingCountry id="F3047621FEDFFEADFE0BB96BA467FAA0" box="[489,631,1359,1381]" name="Costa Rica" pageId="266" pageNumber="265">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B967EAA3FEDFFEADFD64B974A4E4FAA0" bold="true" box="[646,756,1360,1381]" pageId="266" pageNumber="265">
|
||
<collectingRegion id="49D7F853FEDFFEADFD64B974A4E4FAA0" box="[646,756,1360,1381]" country="Costa Rica" name="Puntarenas" pageId="266" pageNumber="265">Puntarenas</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="8ECC606AFEDFFEADFCE1B974A57DFAA0" LSID="urn:lsid:plazi:treatment:03BA87A7FEDCFEADFDE4BCFAA4C7FA0A:8ECC606AFEDFFEADFCE1B974A57DFAA0" box="[771,877,1360,1381]" country="Costa Rica" name="Las Cruces" pageId="266" pageNumber="265" stateProvince="Puntarenas">Las Cruces</location>
|
||
,
|
||
<collectionCode id="ED02AE74FEDFFEADFC98B974A58CFAA0" box="[890,924,1360,1381]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="266" pageNumber="265">MT</collectionCode>
|
||
/
|
||
<collectionCode id="ED02AE74FEDFFEADFC41B974A5DEFAA0" box="[931,974,1360,1381]" pageId="266" pageNumber="265">YPT</collectionCode>
|
||
,
|
||
<date id="FFAD1071FEDFFEADFC38B96BA24BFAA0" box="[986,1115,1359,1381]" pageId="266" pageNumber="265" value="1990-03-18" valueMax="1990-03-22" valueMin="1990-03-18">
|
||
<collectingDate id="EFE9E999FEDFFEADFC38B96BA24BFAA0" box="[986,1115,1359,1381]" pageId="266" pageNumber="265" value="1990-03-18" valueMax="1990-03-22" valueMin="1990-03-18">18-22.iii.1990</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26E65367FEDFFEADFB8EB974A6E3FABA" pageId="266" pageNumber="265">J.S. Noyes</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7B3CECFEDFFEADFEE4B94EA402FA5F" collectingDate="1990-03-18" collectingDateMax="1990-03-22" collectingDateMin="1990-03-18" collectionCode="MT, YPT" collectorName="P. Hanson" country="Costa Rica" elevation="1200" location="San Vito" pageId="266" pageNumber="265" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
|
||
<typeStatus id="54A88813FEDFFEADFEE4B94EA772FABA" box="[262,354,1386,1407]" pageId="266" pageNumber="265">Paratypes</typeStatus>
|
||
:
|
||
<emphasis id="B967EAA3FEDFFEADFE8CB94EA7ECFA45" bold="true" box="[366,508,1386,1408]" pageId="266" pageNumber="265">
|
||
<collectingCountry id="F3047621FEDFFEADFE8CB94EA7ECFA45" box="[366,508,1386,1408]" name="Costa Rica" pageId="266" pageNumber="265">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
, 1E,
|
||
<emphasis id="B967EAA3FEDFFEADFDD5B94EA4B5FABA" bold="true" box="[567,677,1386,1407]" pageId="266" pageNumber="265">
|
||
<collectingRegion id="49D7F853FEDFFEADFDD5B94EA4B5FABA" box="[567,677,1386,1407]" country="Costa Rica" name="Puntarenas" pageId="266" pageNumber="265">Puntarenas</collectingRegion>
|
||
</emphasis>
|
||
, same data as holotype; 1E,
|
||
<collectingRegion id="49D7F853FEDFFEADFC20B94EA238FABA" box="[962,1064,1386,1407]" country="Costa Rica" name="Puntarenas" pageId="266" pageNumber="265">Puntarenas</collectingRegion>
|
||
,
|
||
<location id="8ECC606AFEDFFEADFBD7B94EA298FA45" LSID="urn:lsid:plazi:treatment:03BA87A7FEDCFEADFDE4BCFAA4C7FA0A:8ECC606AFEDFFEADFBD7B94EA298FA45" box="[1077,1160,1386,1408]" country="Costa Rica" name="San Vito" pageId="266" pageNumber="265" stateProvince="Puntarenas">San Vito</location>
|
||
,
|
||
<quantity id="4CEB9B54FEDFFEADFF57B9A0A6E7FA5C" box="[181,247,1412,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="266" pageNumber="265" unit="m" value="1200.0">
|
||
<elevation id="003ED182FEDFFEADFF57B9A0A6E7FA5C" box="[181,247,1412,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="266" pageNumber="265" unit="m" value="1200.0">1200m</elevation>
|
||
</quantity>
|
||
, coffee, iii-iv.1996 (
|
||
<collectorName id="26E65367FEDFFEADFE4EB9A0A41BFA5F" box="[428,523,1412,1434]" pageId="266" pageNumber="265">P. Hanson</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7B3CECFEDFFEADFDF9B9A0A5A5FA5F" box="[539,949,1412,1434]" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="266" pageNumber="265" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
|
||
<typeStatus id="54A88813FEDFFEADFDF9B9A0A47FFA5F" box="[539,623,1412,1434]" pageId="266" pageNumber="265">Holotype</typeStatus>
|
||
in
|
||
<collectionCode id="ED02AE74FEDFFEADFD6EB9A0A4CFFA5C" box="[652,735,1412,1433]" pageId="266" pageNumber="265">MZUCR</collectionCode>
|
||
,
|
||
<typeStatus id="54A88813FEDFFEADFD0AB9A1A52FFA5F" box="[744,831,1413,1434]" pageId="266" pageNumber="265">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="ED02AE74FEDFFEADFCBEB9A1A5A5FA5F" box="[860,949,1413,1434]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="266" pageNumber="265" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEDFFEADFF57B993A4C7FA0A" box="[181,727,1463,1487]" pageId="266" pageNumber="265" type="discussion">
|
||
<paragraph id="8BAC36B1FEDFFEADFF57B993A4C7FA0A" blockId="266.[181,727,1463,1487]" box="[181,727,1463,1487]" pageId="266" pageNumber="265">
|
||
COMMENTS. See comments under
|
||
<taxonomicName id="4C134D32FEDFFEADFDC9B993A46AFA0A" authorityName="Noyes" authorityYear="2023" box="[555,634,1463,1487]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="266" pageNumber="265" phylum="Arthropoda" rank="species" species="mirabel">
|
||
<emphasis id="B967EAA3FEDFFEADFDC9B993A46AFA0A" box="[555,634,1463,1487]" italics="true" pageId="266" pageNumber="265">mirabel</emphasis>
|
||
</taxonomicName>
|
||
(p. 263).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |