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<document id="14A9DD5E8F0335576AB8762A9E40B5C1" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FEC0FEB0FDE2B83DA47EFAF9" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus scolopax Noyes 2023, sp.nov." docType="treatment" docUuid="4512ED6A-F14F-4B45-98AC-8FB3416F0363" docUuidSource="ZooBank" docVersion="2" lastPageNumber="278" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="276" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="44107CF8B64CD0B7F031532110D13278">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="0C28631A632D8F9E40FB1448004E5705">Noyes, John Stuart</mods:namePart>
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<mods:date id="2FA68CE51D3082E95CCB9140E960F115">2023</mods:date>
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<mods:number id="82D327F40EA10CA0041949B01B04AA6F">2023-06-30</mods:number>
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<subSubSection id="C309653AFEC0FEB2FDE2B83DA552FBF4" box="[512,834,1049,1073]" pageId="277" pageNumber="276" type="nomenclature">
<paragraph id="8BAC36B1FEC0FEB2FDE2B83DA552FBF4" blockId="277.[512,834,1049,1100]" box="[512,834,1049,1073]" pageId="277" pageNumber="276">
<heading id="D0E481DDFEC0FEB2FDE2B83DA552FBF4" bold="true" box="[512,834,1049,1073]" centered="true" fontSize="10" level="2" pageId="277" pageNumber="276" reason="2">
<emphasis id="B967EAA3FEC0FEB2FDE2B83DA552FBF4" bold="true" box="[512,834,1049,1073]" pageId="277" pageNumber="276">
<taxonomicName id="4C134D32FEC0FEB2FDE2B83DA4E0FBF4" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[512,752,1049,1073]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="277" pageNumber="276" phylum="Arthropoda" rank="species" species="scolopax" status="sp.nov.">
<emphasis id="B967EAA3FEC0FEB2FDE2B83DA4E0FBF4" bold="true" box="[512,752,1049,1073]" italics="true" pageId="277" pageNumber="276">Cheiloneurus scolopax</emphasis>
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<taxonomicNameLabel id="A25457D8FEC0FEB2FD15B83DA552FBF4" box="[759,834,1049,1073]" pageId="277" pageNumber="276" rank="species">sp.nov.</taxonomicNameLabel>
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<subSubSection id="C309653AFEC0FEB2FDF1B810A53FFB89" box="[531,815,1076,1100]" pageId="277" pageNumber="276" type="description">
<paragraph id="8BAC36B1FEC0FEB2FDF1B810A53FFB89" blockId="277.[512,834,1049,1100]" box="[531,815,1076,1100]" pageId="277" pageNumber="276">
(
<figureCitation id="13282A34FEC0FEB2FDF8B810A4B5FB89" box="[538,677,1076,1100]" captionStart-0="Figs 675-686" captionStart-1="Figs 687-698" captionStartId-0="785.[181,226,1526,1550]" captionStartId-1="786.[181,226,1526,1550]" captionTargetBox-0="[183,1163,224,1505]" captionTargetBox-1="[185,1163,226,1508]" captionTargetId-0="figure-10@785.[181,1163,223,1508]" captionTargetId-1="figure-9@786.[181,1163,223,1508]" captionTargetPageId-0="785" captionTargetPageId-1="786" captionText-0="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." captionText-1="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="277" pageNumber="276">Figs 685-690</figureCitation>
; Hab. E 103)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEC0FEB0FF57B84DA771FC24" lastPageId="279" lastPageNumber="278" pageId="277" pageNumber="276" type="diagnosis">
<paragraph id="8BAC36B1FEC0FEB2FF57B84DA29CF952" blockId="277.[181,1165,1129,1687]" pageId="277" pageNumber="276">
DIAGNOSIS.
<emphasis id="B967EAA3FEC0FEB2FEACB84DA78AFB44" box="[334,410,1129,1153]" italics="true" pageId="277" pageNumber="276">Female</emphasis>
(length about
<quantity id="4CEB9B54FEC0FEB2FDDAB84DA4BCFB44" box="[568,684,1129,1153]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="277" pageNumber="276" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1-1.4mm</quantity>
): body mostly orange, generally with a weak metallic sheen; posterior part of mesoscutum (
<figureCitation id="13282A34FEC0FEB2FD49B8A0A517FB59" box="[683,775,1156,1180]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="277" pageNumber="276">Fig. 690</figureCitation>
) dark brown with a metallic sheen, mesopleuron white with a partial, median, brown band, gaster brown mixed pale orange-brown, darker areas with a stronger metallic sheen; antenna (
<figureCitation id="13282A34FEC0FEB2FD03B89DA52AFB14" box="[737,826,1209,1233]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="277" pageNumber="276">Fig. 687</figureCitation>
) with scape pale orange, ventral margin brown; pedicel white with a dark brown streak; funicle off-white, mixed brown on side; clava dark brown; mesoscutum in posterior half with dense, silvery setae; coxae white; legs mostly white to pale orange but with mid and hind tibia with some brown areas; fore wing (
<figureCitation id="13282A34FEC0FEB2FB81B92DA6CCFAF9" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="277" pageNumber="276">Fig. 685</figureCitation>
) mostly infuscate with basal cell almost completely hyaline, a small hyaline area at apex of venation narrowly connected to a small hyaline area on opposite margin via a narrow hyaline area along wing margin; head (
<figureCitation id="13282A34FEC0FEB2FE25B97DA430FAB4" box="[455,544,1369,1393]" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="277" pageNumber="276">Fig. 686</figureCitation>
) about 9.5X as wide as frontovertex, in facial view slightly broader than long, subrectangular, genae slightly curved but weakly converging, more strongly curved inwards near mouth; frontovertex with two setae medially immediately above scrobes; eye separated from scrobe by slightly less than 2X diameter of anterior ocellus, area between eye and scrobe with deep, distinct sculpture, scrobes quite deep with carinate margins; interantennal prominence with about 60 conspicuous pale setae and dorsally rounded; mandible with two teeth and a truncation, virtually tridentate; antenna (
<figureCitation id="13282A34FEC0FEB2FD3FB9DDA527F9D4" box="[733,823,1529,1553]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="277" pageNumber="276">Fig. 687</figureCitation>
) with scape about 3.2X as long as broad; all funicle segments transverse; linear sensilla on F4-F6; clava 3-segmented, slightly longer than pedicel and funicle combined, apex with a strong oblique truncation, sensory area extending about 0.7X along ventral surface of clava; head width clearly greater than length of funicle; mesoscutum (
<figureCitation id="13282A34FEC0FEB2FE76BA40A7F9F9B9" box="[404,489,1636,1660]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="277" pageNumber="276">Fig. 690</figureCitation>
) in anterior half or so with fine striate to striate reticulate sculpture, posterior part with uniform, polygonally reticulate sculpture; scutellum with a distinct, apical tuft
</paragraph>
<paragraph id="8BAC36B1FEC3FEB1FBBCBCA6A29CFF5A" blockId="278.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="278" pageNumber="277">277</paragraph>
<paragraph id="8BAC36B1FEC3FEB1FF57BCFAA74CFE23" blockId="278.[181,1165,222,486]" pageId="278" pageNumber="277">
of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FEC3FEB1FD8EBCFAA4D2FF33" box="[620,706,222,246]" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="278" pageNumber="277">Fig. 685</figureCitation>
) about 2.6-2.7X as long as broad; parastigma clearly downcurved; costal cell dorsally with one or two apical setae, ventrally with a line of setae that is broadly interrupted above parastigma; area below proximal part of parastigma with a group of about 15 paler setae; apices of postmarginal and stigmal veins connected by a naked, hyaline line that extends a little way into disc; seta at apex of postmarginal vein about 0.6X as long as marginal vein; mid tibial spur slightly longer than basitarsus; propodeum (
<figureCitation id="13282A34FEC3FEB1FC55BD40A21FFEB9" box="[951,1039,356,380]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="278" pageNumber="277">Fig. 690</figureCitation>
) with about 50 setae adjacent to spiracle that extend down side; syntergum about 0.8X as long as mid tibia; ovipositor (
<figureCitation id="13282A34FEC3FEB1FEC8BDBDA791FE74" box="[298,385,409,433]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="278" pageNumber="277">Fig. 689</figureCitation>
) about 3.6X gonostylus or about 1.2X as long as mid tibia; gonostylus about 0.9X as long as mid tibial spur; exserted part of ovipositor about 0.3X as long as mid tibial spur.
<emphasis id="B967EAA3FEC3FEB1FF57BDEAA6F9FE23" box="[181,233,462,486]" italics="true" pageId="278" pageNumber="277">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FEC3FEB1FF57BE20A293FDD9" blockId="278.[181,1165,516,1687]" box="[181,1155,516,540]" pageId="278" pageNumber="277">
Female (
<typeStatus id="54A88813FEC3FEB1FEEFBE20A77CFDD9" box="[269,364,516,540]" pageId="278" pageNumber="277">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FEC3FEB1FD40BE20A4E9FDD9" box="[674,761,516,540]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="278" pageNumber="277" unit="mm" value="1.31">1.31mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FEC3FEB1FC3DBE20A223FDD9" box="[991,1075,516,540]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="278" pageNumber="277" unit="mm" value="1.27">1.27mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FEC3FEB1FF39BE3AA7CAFAF8" blockId="278.[181,1165,516,1687]" pageId="278" pageNumber="277">
Head with occiput pale orange, a dark brown, triangular mark above foramen; frontovertex orange, ocellar area dark brown with metallic blue and purple reflections, other parts of frontovertex with much weaker green, brassy and purple reflections; area between eye and scrobe with a fairly strong blue-green and purple reflections, temple similar but a little less shiny; gena with weaker pale blue, brassy, coppery and purple reflections; scrobal area with weak brassy, pale blue and purple reflections; frontovertex clothed with fairly conspicuous, translucent, pale brown setae along inner eye margins, a line of similar setae below eye; gena and interantennal prominence with similar, but shorter and more dense setae, especially on interantennal prominence; maxillary palpus white; antenna (
<figureCitation id="13282A34FEC3FEB1FE7FBED0A7E2FCC9" box="[413,498,756,780]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="278" pageNumber="277">Fig. 687</figureCitation>
) with radicle pale orange, extreme apex with a narrow brown ring; scape pale orange, apex white, externally with ventral margin narrowly brown proximally and a pale brown subapical band, internally with ventral margin broadly brown proximally almost to dorsal margin and narrowly margined brown subapically; pedicel more or less white with a dark brown streak internally along its length; funicle off-white, brown along sides; clava dark brown; pronotum orange with weak brassy and purple reflections, neck dark brown; prosternum pale orange; mesoscutum orange in anterior half clothed with relatively sparse, brown setae, posterior half dark metallic blue with some purple reflections and clothed with relatively dense, silvery setae, posterior margin purple-brown with a weak purple sheen; tegula orange with apex orange-brown; axilla and scutellum orange, clothed with brown setae, subapical tuft black; metanotum pale orange-brown; mesopleuron white with a weak brassy sheen and an incomplete, vertical median brown stripe; mesosternum very pale orange; fore coxa white; fore femur white, tibia and tarsus pale orange; mid coxa white; mid femur white, apex dusky, very pale orange, tibia pale orange with an incomplete, subbasal brown band, tibial spur and tarsus pale orange; hind coxa white; hind femur white, pale orange at apex, tibia pale orange, externally marked with pale brown in proximal half, tarsus pale orange; fore wing (
<figureCitation id="13282A34FEC3FEB1FCC1B8A0A56BFB59" box="[803,891,1156,1180]" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="278" pageNumber="277">Fig. 685</figureCitation>
) mostly infuscate, hyaline towards base, along anterior margin distad of venation and at apex; submarginal vein partially pale yellow, venation otherwise brown; propodeum pale orange with an extensive group of very dense, translucent, silvery setae surrounding spiracle and extending nearly to posterior margin; gaster largely pale orange with weak purple, green and brassy reflections, basal tergites and apex of syntergum brown with a stronger purple and coppery sheen, Gt1 largely metallic green laterally; gonostylus very pale orange.
</paragraph>
<paragraph id="8BAC36B1FEC3FEB1FF39B91BA29DF952" blockId="278.[181,1165,516,1687]" pageId="278" pageNumber="277">
Head (
<figureCitation id="13282A34FEC3FEB1FEC7B91BA792FA92" box="[293,386,1343,1367]" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="278" pageNumber="277">Fig. 686</figureCitation>
) about 9.5X as wide as frontovertex, in profile 1.6X as high as deep, anteriorly virtually straight from top of scrobes to mouth margin, interantennal prominence slightly protuberant; occipital margin sharp, weakly carinate; ocelli forming an angle of about 40°; frontovertex slightly shiny, with distinct, fairly regular, polygonally reticulate sculpture of mesh size mostly slightly smaller than diameter of eye facet; sculpture immediately above scrobes distinctly transversely elongate; narrowest area between eye and scrobe a little less than 2X as wide as diameter of anterior ocellus and with distinct, transversely elongate, polygonally reticulate sculpture which, when viewed from above, appears similar to a fingerprint; temple and gena with irregular, elongate, reticulate sculpture, that on temple of larger mesh than that on frontovertex; scrobes quite deep, ∩-shaped, meeting and sharply margined dorsally and below eye; interantennal prominence dorsally rounded and with similar sculpture to frontovertex; antenna as in
<figureCitation id="13282A34FEC3FEB1FBCDBA6DA298F9A4" box="[1071,1160,1609,1633]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="278" pageNumber="277">Fig. 687</figureCitation>
; scape broadened and flattened, about 3.2X as long as broad; all funicle segments transverse, F1 subquadrate, distal segments broadest; clava strongly broadened, slightly longer than pedicel and
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF57BCA6A6F3FF5A" blockId="279.[181,227,130,159]" box="[181,227,130,159]" pageId="279" pageNumber="278">
<pageNumber id="BBB83D67FEC2FEB0FF57BCA6A6F3FF5A" ambiguity="12" box="[181,227,130,159]" fuzzyness="0" pageId="279" pageNumber="278" score="12" value="278">
<pageTitle id="CB8CEED6FEC2FEB0FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="279" pageNumber="278">278</pageTitle>
</pageNumber>
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FEB5BCA4A5FBFF5D" blockId="279.[343,1003,128,152]" box="[343,1003,128,152]" pageId="279" pageNumber="278">
<pageTitle id="CB8CEED6FEC2FEB0FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="279" pageNumber="278">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FEC2FEB0FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="279" pageNumber="278" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF57BCFAA580FE52" blockId="279.[181,1165,222,940]" pageId="279" pageNumber="278">
funicle combined, sutures strongly oblique, sensory area enlarged, extending ventrally nearly three-quarters along clava, apex strongly obliquely truncate; eye reaching occipital margin, clothed in fairly conspicuous, short setae, each a little shorter than diameter of facet; inner eye margins converging anteriorly; malar sulcus absent; clypeal margin weakly concave medially; mandible with two acute teeth and a truncation, virtually tridentate; apical segment of maxillary palpus not enlarged, hardly longer than radicle. Relative measurements: HW 62,
<collectingRegion id="49D7F853FEC2FEB0FC0CBD40A203FEBE" box="[1006,1043,356,379]" country="Swaziland" name="Hhohho" pageId="279" pageNumber="278">HH</collectingRegion>
59, FV 6.5, POL 4, OOL 0, OCL 4.5, AOL 9, EL 43, EW 31, MS 30, SL 32, SW 10.
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF39BDBDA21FFD34" blockId="279.[181,1165,222,940]" pageId="279" pageNumber="278">
Thorax (
<figureCitation id="13282A34FEC2FEB0FED6BDBDA79DFE74" box="[308,397,409,433]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="279" pageNumber="278">Fig. 690</figureCitation>
) with pronotum not strongly exposed behind head, about 0.7X as long as mesoscutum, pronotum with polygonally reticulate sculpture that is a little shallower than that in ocellar area; mesoscutum in anterior half with fine striate to striate-reticulate sculpture that is about as deep as sculpture in ocellar area, posterior half with slightly shallower polygonally reticulate sculpture; axilla and scutellum with similar sculpture to pronotum, but slightly coarser and deeper, slightly elongate on side of scutellum; visible part of mesoscutum about 1.7X as broad as long; scutellum nearly 1.2X as broad as long, with a distinct subapical tuft, setae mostly about 0.4X as long as scutellum; mid tibia without a distinct external carina; hind femur about 4X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FEC2FEB0FCEEBE4AA578FD43" box="[780,872,622,646]" captionStart="Figs 675-686" captionStartId="785.[181,226,1526,1550]" captionTargetBox="[183,1163,224,1505]" captionTargetId="figure-10@785.[181,1163,223,1508]" captionTargetPageId="785" captionText="Figs 675-686. Cheiloneurus mardor, HT (675) antenna. Cheiloneurus extillus, HT (676) apex of fore wing venation; (677) fore wing; (678) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus racidia (679) apex of fore wing venation; (680) fore wing; (681) antenna. Cheiloneurus miranda (682) apex of fore wing venation; (683) fore wing; (684) antenna (a, outer aspect of clava; b, inner aspect of clava) (reconstructed). Cheiloneurus scolopax (685) fore wing; (686) head, facial view." pageId="279" pageNumber="278">Figs 685</figureCitation>
,
<figureCitation id="13282A34FEC2FEB0FC96BE4AA58DFD43" box="[884,925,622,646]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="279" pageNumber="278">688</figureCitation>
; costal cell with 1 or 2 setae dorsally at apex; apices of postmarginal and stigmal veins connected by a distinct, naked, hyaline streak (
<figureCitation id="13282A34FEC2FEB0FEBBBE80A7A6FD79" box="[345,438,676,700]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="279" pageNumber="278">Fig. 688</figureCitation>
); propodeum (
<figureCitation id="13282A34FEC2FEB0FDB0BE80A4BCFD79" box="[594,684,676,700]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="279" pageNumber="278">Fig. 690</figureCitation>
) medially about 0.16X as long as scutellum, virtually smooth, but with some extremely shallow, reticulate sculpture; setae on side dense, long and conspicuous. Relative measurements: FWL 56, FWW 21; HWL 49, HWW 12.5.
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF39BED0A444FC84" blockId="279.[181,1165,222,940]" pageId="279" pageNumber="278">Gaster with hypopygium reaching about half-way to apex; syntergum about 0.75X as long as mid tibia, with apex weakly angular; ovipositor hardly exserted, the exserted part about 0.3X as long as mid tibial spur or 0.1X mid tibia.</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF39BF60A29DFCB3" blockId="279.[181,1165,222,940]" pageId="279" pageNumber="278">
<typeStatus id="54A88813FEC2FEB0FF39BF60A727FC99" box="[219,311,836,860]" pageId="279" pageNumber="278">Paratype</typeStatus>
. Funicle with linear sensilla present only on F4-F6; gaster without “gland-like” structures on Gt1 or Gt5; ovipositor
<figureCitation id="13282A34FEC2FEB0FDC0BF7BA469FCB2" box="[546,633,863,887]" captionStart="Figs 687-698" captionStartId="786.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1508]" captionTargetId="figure-9@786.[181,1163,223,1508]" captionTargetPageId="786" captionText="Figs 687-698. Cheiloneurus scolopax (687) antenna (a, outer aspect of clava; b, inner aspect of clava); (688) apex of fore wing venation;(689) ovipositor; (690) mesosoma, dorsal view. Cheiloneurus genys (691) mesosoma, dorsal view; (692) fore wing; (693) antenna (a, outer aspect of clava; b, inner aspect of clava); (694) apex of fore wing venation; (695) head, facial view); (696) ovipositor.Cheiloneurus fabius (697) apex of fore wing venation; (698) mesosoma, dorsal view." pageId="279" pageNumber="278">Fig. 689</figureCitation>
. Relative measurements: OL 45, GL 12.5 [MT 36.5].
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF39BF5DA708FC69" blockId="279.[181,1165,222,940]" pageId="279" pageNumber="278">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FEC2FEB0FD5ABF5DA553FC54" box="[696,835,889,913]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.42" metricValueMin="1.06" pageId="279" pageNumber="278" unit="mm" value="1.24" valueMax="1.42" valueMin="1.06">1.06-1.42mm</quantity>
, otherwise very little in material available.
</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF57BFEDA771FC24" blockId="279.[181,353,969,993]" box="[181,353,969,993]" pageId="279" pageNumber="278">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFEC2FEB0FF57BFDBA7F9FBD2" box="[181,489,1023,1047]" pageId="279" pageNumber="278" type="distribution">
<paragraph id="8BAC36B1FEC2FEB0FF57BFDBA7F9FBD2" blockId="279.[181,489,1023,1047]" box="[181,489,1023,1047]" pageId="279" pageNumber="278">
DISTRIBUTION.
<collectingCountry id="F3047621FEC2FEB0FE91BFDBA7F5FBD2" box="[371,485,1023,1047]" name="Costa Rica" pageId="279" pageNumber="278">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEC2FEB0FF57B810A76EFB89" box="[181,382,1076,1100]" pageId="279" pageNumber="278" type="biology_ecology">
<paragraph id="8BAC36B1FEC2FEB0FF57B810A76EFB89" blockId="279.[181,382,1076,1100]" box="[181,382,1076,1100]" pageId="279" pageNumber="278">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFEC2FEB0FF57B84FA4FEFB2E" pageId="279" pageNumber="278" type="materials_examined">
<paragraph id="8BAC36B1FEC2FEB0FF57B84FA7BDFB45" blockId="279.[181,1165,1131,1259]" box="[181,429,1131,1152]" pageId="279" pageNumber="278">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FEC2FEB0FF39B8A2A4FEFB2E" blockId="279.[181,1165,1131,1259]" pageId="279" pageNumber="278">
Type material.
<materialsCitation id="3B7B3CECFEC2FEB0FE87B8A2A4A7FB73" collectionCode="LN" collectorName="PN Palo Verde &amp; I. Jimenez" country="Costa Rica" elevation="400" location="La Espuela" municipality="Puenta" pageId="279" pageNumber="278" specimenCode="LN 258980" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FEC2FEB0FE87B8A2A7A9FB59" box="[357,441,1158,1180]" pageId="279" pageNumber="278">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FEC2FEB0FE3DB8A1A47BFB5E" bold="true" box="[479,619,1157,1179]" pageId="279" pageNumber="278">
<collectingCountry id="F3047621FEC2FEB0FE3DB8A1A47BFB5E" box="[479,619,1157,1179]" name="Costa Rica" pageId="279" pageNumber="278">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FEC2FEB0FD95B8A1A4F8FB5E" bold="true" box="[631,744,1157,1179]" pageId="279" pageNumber="278">
<collectingRegion id="49D7F853FEC2FEB0FD95B8A1A4F8FB5E" box="[631,744,1157,1179]" country="Costa Rica" name="Guanacaste" pageId="279" pageNumber="278">Guanacaste</collectingRegion>
</emphasis>
,
<collectorName id="26E65367FEC2FEB0FD16B8A2A56DFB59" box="[756,893,1158,1180]" pageId="279" pageNumber="278">PN Palo Verde</collectorName>
,
<quantity id="4CEB9B54FEC2FEB0FC6AB8A2A5ACFB5E" box="[904,956,1158,1179]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="279" pageNumber="278" unit="m" value="400.0">
<elevation id="003ED182FEC2FEB0FC6AB8A2A5ACFB5E" box="[904,956,1158,1179]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="279" pageNumber="278" unit="m" value="400.0">400m</elevation>
</quantity>
E
<collectingMunicipality id="6BC8ACCBFEC2FEB0FC34B8A2A206FB5E" box="[982,1046,1158,1179]" pageId="279" pageNumber="278">Puenta</collectingMunicipality>
,
<location id="8ECC606AFEC2FEB0FBC2B8A2A299FB59" LSID="urn:lsid:plazi:treatment:03BA87A7FEC0FEB0FDE2B83DA47EFAF9:8ECC606AFEC2FEB0FBC2B8A2A299FB59" box="[1056,1161,1158,1180]" country="Costa Rica" municipality="Puenta" name="La Espuela" pageId="279" pageNumber="278" stateProvince="Guanacaste">La Espuela</location>
,
<specimenCode id="DBB59ECAFEC2FEB0FF57B885A70FFB73" box="[181,287,1184,1206]" collectionCode="LN" pageId="279" pageNumber="278">LN 258980</specimenCode>
396500,
<quantity id="4CEB9B54FEC2FEB0FE94B884A7B1FB70" box="[374,417,1184,1205]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="279" pageNumber="278" unit="m" value="10.0">
<elevation id="003ED182FEC2FEB0FE94B884A7B1FB70" box="[374,417,1184,1205]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="279" pageNumber="278" unit="m" value="10.0">10m</elevation>
</quantity>
, #60219, no date (
<collectorName id="26E65367FEC2FEB0FDADB884A4A1FB73" box="[591,689,1184,1206]" pageId="279" pageNumber="278">I. Jiménez</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFEC2FEB0FD23B885A757FB2E" collectingDate="1986-04-13" collectingDateMax="1986-05-04" collectingDateMin="1986-04-13" collectorName="NW Volcan Orosi &amp; Cerro El Hacha" country="Costa Rica" elevation="300" location="Santa" pageId="279" pageNumber="278" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54A88813FEC2FEB0FD23B885A50DFB73" box="[705,797,1185,1206]" pageId="279" pageNumber="278">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FEC2FEB0FCC4B884A5A3FB73" bold="true" box="[806,947,1184,1206]" pageId="279" pageNumber="278">
<collectingCountry id="F3047621FEC2FEB0FCC4B884A5A3FB73" box="[806,947,1184,1206]" name="Costa Rica" pageId="279" pageNumber="278">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FEC2FEB0FC0BB884A24AFB73" bold="true" box="[1001,1114,1184,1206]" pageId="279" pageNumber="278">
<collectingRegion id="49D7F853FEC2FEB0FC0BB884A24AFB73" box="[1001,1114,1184,1206]" country="Costa Rica" name="Guanacaste" pageId="279" pageNumber="278">Guanacaste</collectingRegion>
</emphasis>
,
<collectorName id="26E65367FEC2FEB0FB85B885A720FB14" pageId="279" pageNumber="278">NW Volcan Orosi</collectorName>
,
<collectorName id="26E65367FEC2FEB0FED8B89FA7DBFB14" box="[314,459,1211,1233]" pageId="279" pageNumber="278">Cerro El Hacha</collectorName>
,
<quantity id="4CEB9B54FEC2FEB0FE37B89FA41BFB15" box="[469,523,1211,1232]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="279" pageNumber="278" unit="m" value="300.0">
<elevation id="003ED182FEC2FEB0FE37B89FA41BFB15" box="[469,523,1211,1232]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="279" pageNumber="278" unit="m" value="300.0">300m</elevation>
</quantity>
, 1988; 1E,
<collectingRegion id="49D7F853FEC2FEB0FD9BB89FA4F4FB15" box="[633,740,1211,1232]" country="Costa Rica" name="Guanacaste" pageId="279" pageNumber="278">Guanacaste</collectingRegion>
,
<location id="8ECC606AFEC2FEB0FD0CB89FA530FB15" LSID="urn:lsid:plazi:treatment:03BA87A7FEC0FEB0FDE2B83DA47EFAF9:8ECC606AFEC2FEB0FD0CB89FA530FB15" box="[750,800,1211,1232]" country="Costa Rica" name="Santa" pageId="279" pageNumber="278" stateProvince="Guanacaste">Santa</location>
<taxonomicName id="4C134D32FEC2FEB0FCC5B89FA5C8FB14" authority="NP, Hacienda" authorityName="NP, Hacienda" box="[807,984,1211,1233]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="279" pageNumber="278" phylum="Tracheophyta" rank="genus">Rosa NP, Hacienda</taxonomicName>
1-0,
<date id="FFAD1071FEC2FEB0FBEBB89FA29CFB14" box="[1033,1164,1211,1233]" pageId="279" pageNumber="278" value="1986-04-13" valueMax="1986-05-04" valueMin="1986-04-13">
<collectingDate id="EFE9E999FEC2FEB0FBEBB89FA29CFB14" box="[1033,1164,1211,1233]" pageId="279" pageNumber="278" value="1986-04-13" valueMax="1986-05-04" valueMin="1986-04-13">13.iv-4.v.1986</collectingDate>
</date>
(Janzen, Gauld)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFEC2FEB0FEB2B8F1A4FAFB2E" box="[336,746,1237,1259]" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="279" pageNumber="278" specimenCount="2" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FEC2FEB0FEB2B8F1A7B4FB2E" box="[336,420,1237,1259]" pageId="279" pageNumber="278">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FEC2FEB0FE20B8F2A405FB2E" box="[450,533,1238,1259]" pageId="279" pageNumber="278">MZUCR</collectionCode>
,
<typeStatus id="54A88813FEC2FEB0FDFCB8F2A465FB2E" box="[542,629,1238,1259]" pageId="279" pageNumber="278">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FEC2FEB0FD73B8F2A4FAFB2E" box="[657,746,1238,1259]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="279" pageNumber="278" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFEC2FEB0FF57B92DA47EFAF9" pageId="279" pageNumber="278" type="discussion">
<paragraph id="8BAC36B1FEC2FEB0FF57B92DA47EFAF9" blockId="279.[181,1164,1289,1340]" pageId="279" pageNumber="278">COMMENTS. The sculpture between the eye and scrobe in this species is very unusual in that it has the appearance of parts of a fingerprint.</paragraph>
</subSubSection>
</treatment>
</document>