246 lines
41 KiB
XML
246 lines
41 KiB
XML
<document id="8ED3939273EAF78F7708090B476FA4D1" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FEB4FEC5FDF5BCFAA203F9BE" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Cheiloneurus bion Noyes 2023, sp.nov." docType="treatment" docUuid="FDC83E81-BF53-43ED-8D18-4F42358F23D2" docUuidSource="ZooBank" docVersion="1" lastPageNumber="353" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="352" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="DC53FCE4E278D6DB7D1D9324458FA6A4">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="70194799D1656DE163740DEE684C5ED7">Noyes, John Stuart</mods:namePart>
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<mods:title id="4DE7E14B727836C0B5E45BE4FD39A9FA">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:part id="6C46B98B6B347FAFFCDE05A339FD9655">
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<mods:date id="9441C2B3C6344CD99FFDE956FB1F582C">2023</mods:date>
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<mods:title id="13F22A26B12854159D2642459E4DBF3E">Oxford, England</mods:title>
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<mods:number id="E646193A62F3EECC08DB586149A088F9">2023-06-30</mods:number>
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<paragraph id="8BAC36B1FEB4FEC6FDF5BCFAA53BFF33" blockId="353.[531,814,222,273]" box="[535,811,222,246]" pageId="353" pageNumber="352">
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<heading id="D0E481DDFEB4FEC6FDF5BCFAA53BFF33" bold="true" box="[535,811,222,246]" centered="true" fontSize="10" level="2" pageId="353" pageNumber="352" reason="2">
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<emphasis id="B967EAA3FEB4FEC6FDF5BCFAA53BFF33" bold="true" box="[535,811,222,246]" pageId="353" pageNumber="352">
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<taxonomicName id="4C134D32FEB4FEC6FDF5BCFAA4C9FF33" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[535,729,222,246]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="353" pageNumber="352" phylum="Arthropoda" rank="species" species="bion" status="sp. nov.">
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<emphasis id="B967EAA3FEB4FEC6FDF5BCFAA4C9FF33" bold="true" box="[535,729,222,246]" italics="true" pageId="353" pageNumber="352">Cheiloneurus bion</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25457D8FEB4FEC6FD02BCFAA53BFF33" box="[736,811,222,246]" pageId="353" pageNumber="352" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFEB4FEC6FDF1BCDDA53EFED4" box="[531,814,249,273]" pageId="353" pageNumber="352" type="description">
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<paragraph id="8BAC36B1FEB4FEC6FDF1BCDDA53EFED4" blockId="353.[531,814,222,273]" box="[531,814,249,273]" pageId="353" pageNumber="352">
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(
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<figureCitation id="13282A34FEB4FEC6FDF8BCDDA4B5FED4" box="[538,677,249,273]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Figs 878-883</figureCitation>
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; Hab. E 128)
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFEB4FEC5FF57BD0AA771FAB4" lastPageId="354" lastPageNumber="353" pageId="353" pageNumber="352" type="diagnosis">
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<paragraph id="8BAC36B1FEB4FEC6FF57BD0AA45AFBD3" blockId="353.[181,1165,302,1046]" pageId="353" pageNumber="352">
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DIAGNOSIS.
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<emphasis id="B967EAA3FEB4FEC6FEB3BD0AA78DFE83" box="[337,413,302,326]" italics="true" pageId="353" pageNumber="352">Female</emphasis>
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(length about
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<quantity id="4CEB9B54FEB4FEC6FDA3BD0AA4A6FE83" box="[577,694,302,326]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="353" pageNumber="352" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.4-2.2mm</quantity>
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): head mostly orange-brown, thorax mostly orange, remainder of body dark to black with a weak to moderate metallic sheen; antenna (
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<figureCitation id="13282A34FEB4FEC6FB81BD6DA6CCFEBE" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 878</figureCitation>
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) with scape white margined dark brown or orange; pedicel and flagellum dark brown; darker area of mesoscutum clothed with silvery setae; fore and hind coxae white, middle coxa white sometimes mixed brown; legs white to pale orange with brown areas on femora and tibiae; fore wing (
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<figureCitation id="13282A34FEB4FEC6FEC9BD97A794FE0E" box="[299,388,435,459]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 878</figureCitation>
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) mostly infuscate with a V-shaped hyaline area proximal to linea calva, a small hyaline area at apex of venation and opposite on posterior wing margin, apex less strongly infuscate; head (
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<figureCitation id="13282A34FEB4FEC6FEBFBDCDA7A4FDC4" box="[349,436,489,513]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 883</figureCitation>
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) about 3.6X as wide as frontovertex, in facial view slightly longer than broad, suboval, genae converging and weakly curved; frontovertex naked medially below anterior ocellus; eye separated from scrobe by about 0.8X diameter of ocellus, area between eye and scrobe completely smooth, scrobal margin carinate; interantennal prominence dorsally rounded with about 40 conspicuous setae; mandible with two teeth and a truncation; antenna (
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<figureCitation id="13282A34FEB4FEC6FBCEBE77A295FDA9" box="[1068,1157,595,620]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 878</figureCitation>
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) with scape about 4.6X as long as broad; funicle with F1 about 2X as long as broad, remaining segments longer than broad or sometimes F5 quadrate and F6 transverse; head width slightly less than length of flagellum; linear sensilla on F4-F6, sometimes also F4; clava 3-segmented, about as long as F4-F6 combined; sensory area large and forming an oblique truncation, extending nearly 0.4X along ventral margin; mesoscutum (
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<figureCitation id="13282A34FEB4FEC6FD4BBEFDA511FD34" box="[681,769,729,753]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 882</figureCitation>
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) with coarse linear sculpture medially in anterior half and laterally, otherwise with polygonally reticulate sculpture; scutellum with a distinct apical tuft of setae; wings fully developed; fore wing (
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<figureCitation id="13282A34FEB4FEC6FCA3BF2AA589FCE3" box="[833,921,782,806]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 880</figureCitation>
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) about 3.0X as long as broad; parastigma very distinctly curved downwards; costal cell dorsally with 1 or 2 setae at apex and ventrally with a single complete line of setae that is broadly interrupted above parastigma; area below proximal part of parastigma with a group of about 20 setae; apices of postmarginal and stigmal veins connected by a naked hyaline area that extends a little way into disc (
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<figureCitation id="13282A34FEB4FEC6FBCABF5DA291FC54" box="[1064,1153,889,913]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 879</figureCitation>
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); apical bristle of postmarginal vein about 0.7X as long as marginal vein; mid tibial spur about as long as basitarsus; propodeum with about 5 setae around each spiracle, side naked; syntergum about as long as mid tibia; ovipositor (
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<figureCitation id="13282A34FEB4FEC6FDA5BFEDA48FFC24" box="[583,671,969,993]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 881</figureCitation>
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) about 2.7X gonostylus or 2.2X as long as mid tibia; gonostylus about 2.3X as long as mid tibial spur; exserted part of ovipositor about 1.5X as long as mid tibial spur.
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<emphasis id="B967EAA3FEB4FEC6FE41BFDAA7C7FBD3" box="[419,471,1022,1046]" italics="true" pageId="353" pageNumber="352">Male</emphasis>
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: unknown.
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</paragraph>
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<paragraph id="8BAC36B1FEB4FEC6FF57B810A293FB89" blockId="353.[181,1165,1076,1687]" box="[181,1155,1076,1100]" pageId="353" pageNumber="352">
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Female (
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<typeStatus id="54A88813FEB4FEC6FEECB810A77DFB89" box="[270,365,1076,1100]" pageId="353" pageNumber="352" type="holotype">holotype</typeStatus>
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): length, including ovipositor,
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<quantity id="4CEB9B54FEB4FEC6FD40B810A4E8FB89" box="[674,760,1076,1100]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="353" pageNumber="352" unit="mm" value="2.24">2.24mm</quantity>
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; excluding ovipositor,
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<quantity id="4CEB9B54FEB4FEC6FC3DB810A223FB89" box="[991,1075,1076,1100]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="353" pageNumber="352" unit="mm" value="1.9">1.90mm</quantity>
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(CPD).
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||
</paragraph>
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<paragraph id="8BAC36B1FEB4FEC5FF39B86AA7B5FEA4" blockId="353.[181,1165,1076,1687]" lastBlockId="354.[181,1165,222,1340]" lastPageId="354" lastPageNumber="353" pageId="353" pageNumber="352">
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||
Head pale orange-brown, darker on frontovertex, temple and gena; a moderate blue sheen on frontovertex, mixed purple anteriorly and behind ocelli; area between eye and scrobe brassy; top of interantennal prominence shining metallic blue, area between toruli green; mouth margin with a slight green lustre; temple largely metallic green or blue-green, mixed brassy; gena with a slight blue sheen, mixed purple; frontovertex with inconspicuous dark brown setae; antenna (
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<figureCitation id="13282A34FEB4FEC6FBCCB89DA295FB14" box="[1070,1157,1209,1233]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 878</figureCitation>
|
||
) with radicle pale orange; scape white, with proximal two thirds of ventral margin narrowly dark brown, upper margin externally pale orange, internally with distal two-thirds dark brown; pedicel and flagellum dark brown; thorax orange with some darker areas; neck of pronotum laterally dark brown; mesoscutum posteriorly dark brown with a weak purple and indigo sheen, the anterior margin of the dark brown area convexly curved and extending a little more than half-way across mesoscutum, darker area clothed in silvery setae that are more dense anteriorly and laterally, setae in paler area pale brown; tegula orange-brown; axilla dusky posteriorly; scutellum largely clothed in pale brown setae, subapical tuft conspicuous and black; metanotum dark brown with a slight purple sheen; mesopleuron anterior slightly dusky and with a slight brassy, green and purple sheen, posterior half with a very weak purple and brassy sheen; fore coxa white; fore femur pale orange with a brown streak externally in distal half connected to an inconspicuous, brown, subapical ring, tibia mostly brown with apices white or pale orange; mid coxa ventrally brown, otherwise dirty white; mid femur white, apex pale orange with a small, external, subapical brown streak, tibia pale orange, dark brown, then white at extreme base and with a elongate, subbasal brown spot; hind coxa white; hind femur pale orange, pale brownish along ventral margin, hind tibia pale orange, but narrowly white basally with a small, faint, brown, subbasal spot externally; tarsi orange, pretarsi slightly dusky; fore wing (
|
||
<figureCitation id="13282A34FEB4FEC6FD22BA5AA50DF952" box="[704,797,1662,1687]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="353" pageNumber="352">Fig. 880</figureCitation>
|
||
) infuscate with a hyaline pattern, especially proximally; submarginal vein pale yellow, otherwise venation brown; propodeum dark purple-brown with a green lustre, medially slightly purplish and brassy; sides with not more than 5 inconspicuous silvery setae near spiracle; gaster dark purple-brown, mixed coppery, purple and brassy, sides and outer plate of ovipositor additionally with some metallic blue and green; gonostylus pale yellow.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF39BD40A7E5FC84" blockId="354.[181,1165,222,1340]" pageId="354" pageNumber="353">
|
||
Head (
|
||
<figureCitation id="13282A34FEB7FEC5FEFCBD40A764FEB9" box="[286,372,356,380]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 883</figureCitation>
|
||
) in facial view about 3.6X as wide as frontovertex, in profile anteriorly almost evenly rounded, shallow, about 2X as high as deep; occipital margin carinate, occiput without a median groove above foramen; ocelli forming an angle of about 85°; frontovertex slightly shiny, with shallow, transversely elongate, reticulate sculpture of mesh size generally greater than diameter of eye facet; scrobal area and mouth margin with shallower sculpture, interantennal prominence dorsally smooth and shiny; temple virtually smooth, with extremely shallow, longitudinally elongate sculpture, gena with similar sculpture to temple but slightly deeper and of smaller cells; scrobes deep, meeting dorsally, dorsal and lateral margins sharp, carinate; antenna as in
|
||
<figureCitation id="13282A34FEB7FEC5FF13BE1DA75BFD94" box="[241,331,569,593]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 878</figureCitation>
|
||
; scape hardly broadened and flattened, about 4.6X as long as broad; all funicle segments clearly longer than broad, F1 relatively longest, about 3X as long as broad, segments broadest distad so that F6 is only about 1.2X as long as broad; linear sensilla present only on F3-F4; clava hardly broadened, about as long as F4-F6 combined, sutures oblique, sensory area enlarged, extending about one-third along clava ventrally, giving it a distinct, obliquely truncate appearance; eye almost reaching occipital margin, separated by less than diameter of facet, appearing naked, clothed with extremely short, inconspicuous setae; inner eye margins slightly diverging anteriorly; malar sulcus absent; mandible with 2 short, acute teeth and a truncation. Relative measurements: HW 79,
|
||
<collectingRegion id="49D7F853FEB7FEC5FDE6BF2BA439FCE3" box="[516,553,783,806]" country="Swaziland" name="Hhohho" pageId="354" pageNumber="353">HH</collectingRegion>
|
||
86, FV 22, OD 6, POL 12, OOL 0.5, OCL 7, AOL 9, EL 58, EW 40, MS 30, SL 39, SW 8.5.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF39BF60A6C5FBA2" blockId="354.[181,1165,222,1340]" pageId="354" pageNumber="353">
|
||
Thorax (
|
||
<figureCitation id="13282A34FEB7FEC5FEDABF60A783FC99" box="[312,403,836,860]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 882</figureCitation>
|
||
) with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size less than diameter facet; sculpture on mesoscutum, axilla and scutellum similar to that on pronotum, but more regular and significantly coarser; extreme apex and sides of scutellum completely smooth and shiny; visible part of mesoscutum about 2X as broad as long; scutellum very slightly broader than long, with a very distinct subapical tuft, setae at least about half as long as scutellum; hind femur about 4X as long as broad; fore wing with venation and setation as in
|
||
<figureCitation id="13282A34FEB7FEC5FE88BFC0A7E8FC39" box="[362,504,996,1020]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Figs 878, 879</figureCitation>
|
||
; costal cell without setae dorsally at apex; apices of postmarginal and stigmal veins connected by a distinct, naked, hyaline streak (
|
||
<figureCitation id="13282A34FEB7FEC5FC8EBFDBA5D9FBD2" box="[876,969,1023,1047]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 879</figureCitation>
|
||
); propodeum (
|
||
<figureCitation id="13282A34FEB7FEC5FB81BFDBA6CCFBF4" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 882</figureCitation>
|
||
) naked but for a few setae anterior and lateral to spiracle, medially about 0.18X as long as scutellum and completely smooth. Relative measurements: FWL 76, FWW 25.5; HWL 68, HWW 17.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF39B84DA455FB17" blockId="354.[181,1165,222,1340]" pageId="354" pageNumber="353">
|
||
Gaster without “gland-like” structures on Gt1 or Gt5; hypopygium reaching about half-way to apex; syntergum about as long as mid tibia, with apex acute; ovipositor (
|
||
<figureCitation id="13282A34FEB7FEC5FC78B8A0A5E1FB59" box="[922,1009,1156,1180]" captionStart="Figs 878-892" captionStartId="800.[181,226,1526,1550]" captionTargetBox="[182,1163,226,1504]" captionTargetId="figure-9@800.[181,1163,223,1508]" captionTargetPageId="800" captionText="Figs 878-892. Cheiloneurus bion – (878) antenna, HT; (879) apex of fore wing venation; (880) fore wing, HT; (881) ovipositor, HT; (882) mesosoma, dorsal view, HT; (883) head, facial view, HT. Cheiloneurus zephyrus – (884) mesosoma, dorsal view; (885) head, facial view; (886) antenna; (887) ovipositor; (888) hypopygium; (889) apex of fore wing venation; (890) fore wing. Cheiloneurus pherus – (891) hypopygium; (892) mesosoma, dorsal view." pageId="354" pageNumber="353">Fig. 881</figureCitation>
|
||
) well exserted, the exserted part about 1.5X as long as mid tibial spur or about 0.6X as long as mid tibia. Relative measurements: OL 98, GL 36 [MT 44].
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF39B8F0A70FFAF9" blockId="354.[181,1165,222,1340]" pageId="354" pageNumber="353">
|
||
Variation. Females vary in overall length from
|
||
<quantity id="4CEB9B54FEB7FEC5FD50B8F0A52DFB29" box="[690,829,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.24" metricValueMin="1.36" pageId="354" pageNumber="353" unit="mm" value="1.8" valueMax="2.24" valueMin="1.36">1.36-2.24mm</quantity>
|
||
, the mid coxa may be completely white or dirty white mixed brown, in smaller specimens F5 may be quadrate and F6 transverse, linear sensilla may be present on F3-F6 or F4-F6, otherwise little variation noted in material examined.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF57B97DA771FAB4" blockId="354.[181,353,1369,1393]" box="[181,353,1369,1393]" pageId="354" pageNumber="353">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEB7FEC5FF57B9ABA7F9FA62" box="[181,489,1423,1447]" pageId="354" pageNumber="353" type="distribution">
|
||
<paragraph id="8BAC36B1FEB7FEC5FF57B9ABA7F9FA62" blockId="354.[181,489,1423,1447]" box="[181,489,1423,1447]" pageId="354" pageNumber="353">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F3047621FEB7FEC5FE91B9ABA7F5FA62" box="[371,485,1423,1447]" name="Costa Rica" pageId="354" pageNumber="353">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEB7FEC5FF57B9E0A76EFA19" box="[181,382,1476,1500]" pageId="354" pageNumber="353" type="biology_ecology">
|
||
<paragraph id="8BAC36B1FEB7FEC5FF57B9E0A76EFA19" blockId="354.[181,382,1476,1500]" box="[181,382,1476,1500]" pageId="354" pageNumber="353">HOSTS. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFEB7FEC5FF57B9DFA203F9BE" pageId="354" pageNumber="353" type="materials_examined">
|
||
<paragraph id="8BAC36B1FEB7FEC5FF57B9DFA7BDF9D5" blockId="354.[181,1165,1531,1659]" box="[181,429,1531,1552]" pageId="354" pageNumber="353">MATERIAL EXAMINED.</paragraph>
|
||
<paragraph id="8BAC36B1FEB7FEC5FF39BA32A203F9BE" blockId="354.[181,1165,1531,1659]" pageId="354" pageNumber="353">
|
||
Type material.
|
||
<materialsCitation id="3B7B3CECFEB7FEC5FE83BA32A411F983" collectingDate="1993-03-15" collectorName="S. Puerto Viejo" country="Costa Rica" elevation="100" location="Heredia" pageId="354" pageNumber="353" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
|
||
<typeStatus id="54A88813FEB7FEC5FE83BA32A7A5F9E9" box="[353,437,1558,1580]" pageId="354" pageNumber="353" type="holotype">Holotype</typeStatus>
|
||
E:
|
||
<emphasis id="B967EAA3FEB7FEC5FE35BA31A471F9EE" bold="true" box="[471,609,1557,1579]" pageId="354" pageNumber="353">
|
||
<collectingCountry id="F3047621FEB7FEC5FE35BA31A471F9EE" box="[471,609,1557,1579]" name="Costa Rica" pageId="354" pageNumber="353">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B967EAA3FEB7FEC5FD89BA31A4A9F9EE" bold="true" box="[619,697,1557,1579]" pageId="354" pageNumber="353">
|
||
<collectingRegion id="49D7F853FEB7FEC5FD89BA31A4A9F9EE" box="[619,697,1557,1579]" country="Costa Rica" name="Heredia" pageId="354" pageNumber="353">Heredia</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<quantity id="4CEB9B54FEB7FEC5FD21BA32A4FBF9E9" box="[707,747,1558,1580]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="354" pageNumber="353" unit="km" value="3.0">3km</quantity>
|
||
<collectorName id="26E65367FEB7FEC5FD12BA32A569F9E9" box="[752,889,1558,1580]" pageId="354" pageNumber="353">S. Puerto Viejo</collectorName>
|
||
, OTS-La Selva,
|
||
<quantity id="4CEB9B54FEB7FEC5FBEFBA32A252F9EE" box="[1037,1090,1558,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="354" pageNumber="353" unit="m" value="100.0">
|
||
<elevation id="003ED182FEB7FEC5FBEFBA32A252F9EE" box="[1037,1090,1558,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="354" pageNumber="353" unit="m" value="100.0">100m</elevation>
|
||
</quantity>
|
||
, canopy fog
|
||
<taxonomicName id="4C134D32FEB7FEC5FF3BBA15A750F983" box="[217,320,1585,1606]" class="Magnoliopsida" family="Meliaceae" genus="Carapa" kingdom="Plantae" order="Sapindales" pageId="354" pageNumber="353" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B967EAA3FEB7FEC5FF3BBA15A70EF983" box="[217,286,1585,1606]" italics="true" pageId="354" pageNumber="353">Carapa</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, FOT/02,
|
||
<date id="FFAD1071FEB7FEC5FE7EBA14A411F983" box="[412,513,1584,1606]" pageId="354" pageNumber="353" value="1993-03-15">
|
||
<collectingDate id="EFE9E999FEB7FEC5FE7EBA14A411F983" box="[412,513,1584,1606]" pageId="354" pageNumber="353" value="1993-03-15">15.iii.1993</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7B3CECFEB7FEC5FDE8BA15A4AAF9A5" collectingDate="2011-02-24" collectingDateMax="2011-02-25" collectingDateMin="2011-02-24" collectorName="J. S. Noyes" country="Costa Rica" elevation="75" latitude="10.433333" location="Estacion Biologica La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="354" pageNumber="353" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<typeStatus id="54A88813FEB7FEC5FDE8BA15A476F983" box="[522,614,1585,1606]" pageId="354" pageNumber="353" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<emphasis id="B967EAA3FEB7FEC5FD8CBA14A4E9F983" bold="true" box="[622,761,1584,1606]" pageId="354" pageNumber="353">
|
||
<collectingCountry id="F3047621FEB7FEC5FD8CBA14A4E9F983" box="[622,761,1584,1606]" name="Costa Rica" pageId="354" pageNumber="353">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
, 1E,
|
||
<emphasis id="B967EAA3FEB7FEC5FCCEBA14A56AF983" bold="true" box="[812,890,1584,1606]" pageId="354" pageNumber="353">
|
||
<collectingRegion id="49D7F853FEB7FEC5FCCEBA14A56AF983" box="[812,890,1584,1606]" country="Costa Rica" name="Heredia" pageId="354" pageNumber="353">Heredia</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="8ECC606AFEB7FEC5FC67BA14A299F983" LSID="urn:lsid:plazi:treatment:03BA87A7FEB4FEC5FDF5BCFAA203F9BE:8ECC606AFEB7FEC5FC67BA14A299F983" box="[901,1161,1584,1606]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Estacion Biologica La Selva" pageId="354" pageNumber="353" stateProvince="Heredia">Estacion Biologica La Selva</location>
|
||
,
|
||
<geoCoordinate id="EE275076FEB7FEC5FF57BA6FA714F9A4" box="[181,260,1611,1633]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="354" pageNumber="353" precision="925" value="10.433333">10°26’N</geoCoordinate>
|
||
<geoCoordinate id="EE275076FEB7FEC5FEE9BA6FA74EF9A5" box="[267,350,1611,1632]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="354" pageNumber="353" precision="925" value="-84.01667">84°01’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B54FEB7FEC5FE8BBA6FA783F9A4" box="[361,403,1611,1633]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="354" pageNumber="353" unit="m" value="75.0">
|
||
<elevation id="003ED182FEB7FEC5FE8BBA6FA783F9A4" box="[361,403,1611,1633]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="354" pageNumber="353" unit="m" value="75.0">75m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFAD1071FEB7FEC5FE7CBA6FA407F9A4" box="[414,535,1611,1633]" pageId="354" pageNumber="353" value="2011-02-24" valueMax="2011-02-25" valueMin="2011-02-24">
|
||
<collectingDate id="EFE9E999FEB7FEC5FE7CBA6FA407F9A4" box="[414,535,1611,1633]" pageId="354" pageNumber="353" value="2011-02-24" valueMax="2011-02-25" valueMin="2011-02-24">24-25.ii.2011</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26E65367FEB7FEC5FDC4BA6FA49CF9A5" box="[550,652,1611,1632]" pageId="354" pageNumber="353">J.S. Noyes</collectorName>
|
||
); 3E
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B7B3CECFEB7FEC5FD24BA6FA7F3F9BE" collectingDate="2015-02-14" collectionCode="INBio" collectorName="J. S. Noyes" country="Costa Rica" latitude="9.983334" location="Parque" longLatPrecision="1298" longitude="-84.13333" municipality="Santo Domingo" pageId="354" pageNumber="353" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<emphasis id="B967EAA3FEB7FEC5FD24BA6FA504F9A4" bold="true" box="[710,788,1611,1633]" pageId="354" pageNumber="353">
|
||
<collectingRegion id="49D7F853FEB7FEC5FD24BA6FA504F9A4" box="[710,788,1611,1633]" country="Costa Rica" name="Heredia" pageId="354" pageNumber="353">Heredia</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<collectingMunicipality id="6BC8ACCBFEB7FEC5FCC3BA6FA5A3F9A4" box="[801,947,1611,1633]" pageId="354" pageNumber="353">Santo Domingo</collectingMunicipality>
|
||
,
|
||
<collectionCode id="ED02AE74FEB7FEC5FC5CBA6FA5E7F9A4" box="[958,1015,1611,1633]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="354" pageNumber="353">INBio</collectionCode>
|
||
<location id="8ECC606AFEB7FEC5FC1FBA6FA22EF9A5" LSID="urn:lsid:plazi:treatment:03BA87A7FEB4FEC5FDF5BCFAA203F9BE:8ECC606AFEB7FEC5FC1FBA6FA22EF9A5" box="[1021,1086,1611,1632]" country="Costa Rica" latitude="9.983334" longLatPrecision="1298" longitude="-84.13333" municipality="Santo Domingo" name="Parque" pageId="354" pageNumber="353" stateProvince="Heredia">Parque</location>
|
||
,
|
||
<geoCoordinate id="EE275076FEB7FEC5FBABBA6FA29CF9A4" box="[1097,1164,1611,1633]" degrees="9" direction="north" minutes="59" orientation="latitude" pageId="354" pageNumber="353" precision="925" value="9.983334">9°59’N</geoCoordinate>
|
||
<geoCoordinate id="EE275076FEB7FEC5FF57BA42A718F9BE" box="[181,264,1638,1659]" degrees="84" direction="west" minutes="08" orientation="longitude" pageId="354" pageNumber="353" precision="925" value="-84.13333">84°08’W</geoCoordinate>
|
||
,
|
||
<date id="FFAD1071FEB7FEC5FEF0BA41A77EF9BE" box="[274,366,1637,1659]" pageId="354" pageNumber="353" value="2015-02-14">
|
||
<collectingDate id="EFE9E999FEB7FEC5FEF0BA41A77EF9BE" box="[274,366,1637,1659]" pageId="354" pageNumber="353" value="2015-02-14">14.ii.2015</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26E65367FEB7FEC5FE9EBA42A7CDF9BE" box="[380,477,1638,1659]" pageId="354" pageNumber="353">J.S. Noyes</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7B3CECFEB7FEC5FE16BA41A200F9BE" box="[500,1040,1637,1659]" collectionCode="NHMUK, MZUCR" country="Costa Rica" pageId="354" pageNumber="353" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
|
||
<typeStatus id="54A88813FEB7FEC5FE16BA41A458F9BE" box="[500,584,1637,1659]" pageId="354" pageNumber="353" type="holotype">Holotype</typeStatus>
|
||
in
|
||
<collectionCode id="ED02AE74FEB7FEC5FD87BA42A4ADF9BE" box="[613,701,1638,1659]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="354" pageNumber="353" type="Museum">NHMUK</collectionCode>
|
||
,
|
||
<typeStatus id="54A88813FEB7FEC5FD24BA42A50DF9BE" box="[710,797,1638,1659]" pageId="354" pageNumber="353" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="ED02AE74FEB7FEC5FCD8BA42A59BF9BE" box="[826,907,1638,1659]" pageId="354" pageNumber="353">MZUCR</collectionCode>
|
||
and
|
||
<collectionCode id="ED02AE74FEB7FEC5FC55BA42A200F9BE" box="[951,1040,1638,1659]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="354" pageNumber="353" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |