258 lines
29 KiB
XML
258 lines
29 KiB
XML
<document id="6979A17C80AE23F0FA15508DE355C72C" ID-CLB-Dataset="50618" ID-DOI="10.5281/zenodo.172933" ID-GBIF-Dataset="fcd3c991-5b05-4d3c-ac16-7e91f3cf61ec" ID-ISSN="11755326" ID-Zenodo-Dep="172933" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459765723628" checkinUser="plazi" docAuthor="A, Socorro Ávila & Flowers, R. Wills" docDate="2006" docId="03BA87A4FF99FD2CFEF7FBB9FE16FA0D" docLanguage="en" docName="zt01245p068.pdf" docOrigin="Zootaxa 1245" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Choroterpes gregoryi Avila and Flowers, new species" docType="treatment" docVersion="10" lastPageNumber="63" masterDocId="FF83FFDCFF98FD28FFFFFFD8FFE9FFB3" masterDocTitle="Two new species of Choroterpes Eaton (Ephemeroptera: Leptophlebiidae) from Costa Rica" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698222416575" updateUser="ExternalLinkService">
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<mods:mods id="95DF2B699C7CF649636976A3781C70C9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A2F274D2A354DE77578F8324208AB2DD">
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<mods:title id="EC6165140C650278F19A181C916B73CD">Two new species of Choroterpes Eaton (Ephemeroptera: Leptophlebiidae) from Costa Rica</mods:title>
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<mods:name id="C469FC176B87132F3903FF45EAF34EC8" type="personal">
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<mods:roleTerm id="CD0536C9D0DB4CD455F1AC991C52C8C7">Author</mods:roleTerm>
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<mods:namePart id="8D0F4EDE1A981450C8B04264195D7F52">A, Socorro Ávila</mods:namePart>
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</mods:name>
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<mods:name id="BAB8454E9DB5C1DD42A3EEE1E83CB441" type="personal">
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<mods:roleTerm id="FF9250F2DF3458AD73FC872B9F4680C4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AF27432997FCF1D85DAEDF840026976E">Flowers, R. Wills</mods:namePart>
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<mods:typeOfResource id="35E9AC27621167B41B616D8BDA6706C2">text</mods:typeOfResource>
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<mods:title id="32C0D2BF69B095BC6311A586D93F3E7B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C286BC23CC93643CF84B85F170815CC8">
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<mods:date id="A1C2D10B58F60CEE95CDFECB4869931F">2006</mods:date>
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<mods:detail id="DFE2CD85C8BF0A3C0F45EDFE84A826B8" type="volume">
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<mods:number id="26A777E43A4D30B0A6D96ED722453F03">1245</mods:number>
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<mods:start id="05BD36B9AA764BC9423C8E7BBD9B613B">59</mods:start>
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<mods:end id="E0FC24F82C8EEA9F542F96CA5BB3E0A1">68</mods:end>
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<mods:classification id="E0F86BF195087659DAC196F6A6C6A4B9">journal article</mods:classification>
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<mods:identifier id="138901C86703B432ADD5FDFAA51C307C" type="CLB-Dataset">50618</mods:identifier>
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<mods:identifier id="7B5D50448758A7A207415E60B0454F56" type="DOI">10.5281/zenodo.172933</mods:identifier>
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<mods:identifier id="6CBCADDEDDE3032C017E982C91075E7D" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="2CE53D318235A9D3D0F8A617483F6D0B" type="Zenodo-Dep">172933</mods:identifier>
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<treatment id="03BA87A4FF99FD2CFEF7FBB9FE16FA0D" ID-DOI="http://doi.org/10.5281/zenodo.6263554" ID-GBIF-Taxon="119345960" ID-Zenodo-Dep="6263554" LSID="urn:lsid:plazi:treatment:03BA87A4FF99FD2CFEF7FBB9FE16FA0D" httpUri="http://treatment.plazi.org/id/03BA87A4FF99FD2CFEF7FBB9FE16FA0D" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60">
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<subSubSection id="C3096539FF99FD29FEF7FBB9FCCAFB2D" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="8BAC36B2FF99FD29FEF7FBB9FC7DFBC8" blockId="1.[264,916,1121,1182]" box="[264,916,1121,1147]" pageId="1" pageNumber="60">
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<heading id="D0E481DEFF99FD29FEF7FBB9FC7DFBC8" bold="true" box="[264,916,1121,1147]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
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<emphasis id="B967EAA0FF99FD29FEF7FBB9FC7DFBC8" bold="true" box="[264,916,1121,1147]" pageId="1" pageNumber="60">
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<taxonomicName id="4C134D31FF99FD29FEF7FBB9FD1FFBC8" ID-CoL="5YF2Y" authority="Avila and Flowers" authorityName="Avila and Flowers" box="[264,758,1121,1147]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="gregoryi" status="sp. nov.">
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<emphasis id="B967EAA0FF99FD29FEF7FBB9FDE1FBC8" bold="true" box="[264,520,1121,1147]" italics="true" pageId="1" pageNumber="60">Choroterpes gregoryi</emphasis>
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Ávila and Flowers
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25457DBFF99FD29FCFCFBB9FC7DFBC8" box="[771,916,1121,1147]" pageId="1" pageNumber="60" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAC36B2FF99FD29FEF7FB5CFCCAFB2D" blockId="1.[264,916,1121,1182]" box="[264,803,1156,1182]" pageId="1" pageNumber="60">(Figs. 1, 3–5, 9, 10, 13, 15, 16, 19, 20, 23, 25)</paragraph>
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</subSubSection>
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<subSubSection id="C3096539FF99FD29FEF7FB0CFE64FB5E" box="[264,397,1236,1261]" pageId="1" pageNumber="60" type="description">
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<paragraph id="8BAC36B2FF99FD29FEF7FB0CFE64FB5E" blockId="1.[264,1325,1236,1902]" box="[264,397,1236,1261]" pageId="1" pageNumber="60">
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<emphasis id="B967EAA0FF99FD29FEF7FB0CFE64FB5E" box="[264,397,1236,1261]" italics="true" pageId="1" pageNumber="60">Discription</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3096539FF99FD2AFEC7FB24FBBFFD75" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60" type="materials_examined">
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<paragraph id="8BAC36B2FF99FD29FEC7FB24FAC0F8DD" blockId="1.[264,1325,1236,1902]" pageId="1" pageNumber="60">
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<emphasis id="B967EAA0FF99FD29FEC7FB24FE58FAA5" bold="true" box="[312,433,1276,1302]" pageId="1" pageNumber="60">
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<typeStatus id="54A88810FF99FD29FEC7FB24FE45FAA5" box="[312,428,1276,1302]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male imago (in alcohol, genitalia and one set of wings on slides): Length: body
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<quantity id="4CEB9B57FF99FD29FEAFFAFCFE4CFA8D" box="[336,421,1316,1342]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="60" unit="mm" value="4.7">4.7mm</quantity>
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, forewing
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<quantity id="4CEB9B57FF99FD29FDD1FAFCFD6DFA8D" box="[558,644,1316,1342]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="60" unit="mm" value="4.8">4.8mm</quantity>
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. Head: brown, washed with black, antenna translucent yellowish brown, scape and pedicel washed with reddish brown. Upper portion of eyes olive brown, stalk very short, lower portion dark gray. Ocelli whitish, their bases dark brown. Thorax: pronotum yellowish brown washed with black laterally and on basal margin; meso and metanota chestnut brown, margins and carinae darker. Pleura orangebrown washed with dark brown, a diagonal area above mesocoxae washed with black. Thoracic sterna orange brown. Forewing (Fig. 1): membrane hyaline, translucent yellowish brown at base; longitudinal veins translucent brown, cross veins hyaline.
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<collectingCountry id="F3047622FF99FD29FB0FF9E4FAC2F9E5" box="[1264,1323,1596,1622]" name="India" pageId="1" pageNumber="60">Hind</collectingCountry>
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wing (Fig. 1): membrane hyaline, veins C and Sc translucent yellow in basal half, remaining veins hyaline. Foreleg: yellowish brown, tarsus lighter; femur with a dark brown spot at midlength; tarsal formula: 0.18:1.0 (
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<quantity id="4CEB9B57FF99FD29FC8CF96CFC35F97D" box="[883,988,1716,1742]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="60" unit="mm" value="0.56">0.56mm</quantity>
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): 0.75:0.39:0.18.
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<collectingCountry id="F3047622FF99FD29FB49F96CFB18F97D" box="[1206,1265,1716,1742]" name="India" pageId="1" pageNumber="60">Hind</collectingCountry>
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legs translucent yellow, femora with dark brown spots at midlength and apically, tibiae darker brown at the femoral joint. Abdomen (Fig. 3, 5): terga translucent yellowish brown, washed with dark brown, with pale median, submedian and lateral markings. Abdominal sterna translucent yellowish brown, sternum IX with brown basal and lateral margins.
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</paragraph>
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<paragraph id="8BAC36B2FF9AFD2AFEF7FEC4FC4FFE35" blockId="2.[264,1325,284,1630]" pageId="2" pageNumber="61">Genitalia (Fig. 4): forceps expanded in basal 1/3; styliger plate whitish, forceps brown, penes whitish. Penis lobes elongate, membranous, with a tiny pair of spines in apical third along midline. Caudal filaments broken off and missing.</paragraph>
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<paragraph id="8BAC36B2FF9AFD2AFEC7FE4BFBBFFD75" blockId="2.[264,1325,284,1630]" pageId="2" pageNumber="61">
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<emphasis id="B967EAA0FF9AFD2AFEC7FE4BFE40FE1E" bold="true" box="[312,425,403,429]" pageId="2" pageNumber="61">Allotype.</emphasis>
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Female subimago (reared with nymphal exuviae). Length: body
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<quantity id="4CEB9B57FF9AFD2AFB2DFE4CFAC1FE1D" box="[1234,1320,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="61" unit="mm" value="4.6">4.6mm</quantity>
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, forewing
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<quantity id="4CEB9B57FF9AFD2AFE86FE64FE26FE65" box="[377,463,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="61" unit="mm" value="5.1">5.1mm</quantity>
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. Coloration as in male imago except: head yellowish white marked with dark brown between ocelli and on vertex; antenna whitish. Thorax yellowish brown, meso and metanota darker brown laterally, membranes whitish washed with grey. Wing membrane whitish, veins translucent. Legs yellowish white, dark spots as in male imago except fore femur has an additional apical dark spot. Abdomen yellowish white with dark brown markings as in male imago. Subanal plate yellowish white, coneshaped, without apical notch. Caudal filaments white with narrow dark bands on annuli.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3096539FF9AFD2AFEC7FD0CFB3CF9ED" pageId="2" pageNumber="61" type="description">
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<paragraph id="8BAC36B2FF9AFD2AFEC7FD0CFB3CF9ED" blockId="2.[264,1325,284,1630]" pageId="2" pageNumber="61">
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<emphasis id="B967EAA0FF9AFD2AFEC7FD0CFE15FD5D" bold="true" box="[312,508,724,750]" pageId="2" pageNumber="61">Mature nymph</emphasis>
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(in alcohol): Body length
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<quantity id="4CEB9B57FF9AFD2AFC9DFD0CFC05FD5D" box="[866,1004,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="2" pageNumber="61" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">4.6–5.1mm</quantity>
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. Head: yellowish brown washed with darker brown, pale spot anterior to the median ocellus, and pale areas between eyes and lateral ocelli. Vertex and area between ocelli washed with black. Antenna yellowish white. Mouthparts: labrum (Fig. 13) with maximum length slightly less than maximum width, anteromedian emargination well developed with three irregular denticles, a row of short dorsal subapical seta present immediately behind anteromedian emargination, and a second row of long dorsal present on apical third, length of this row slightly more than one third the total width of labrum; mandibles (Fig. 15, 16) with outer margins obtusely angled, a tuft of setae at angle; maxilla (Fig. 19) with anteromedian tooth obsolete, galealacina with inner subapical row of 17–18 pectinate setae; hypopharynx as in Fig. 20; segment 2 of labial palpi 3/4 the length of segment 1, segment 3 slightly shorter than segment 2, (Fig. 23) with 5 large dorsal seta on segment 3; labrum, basal 2/3 of mandibles, outer margin of maxilla brown, labium yellowish brown. Thorax: terga yellowish tan, pronotum with crescentshaped submedian spots and lateral margins dark brown; mesonotum washed with brown on lateral margins; pleura washed with dark brown, sterna yellowish brown. Legs: yellowish white, femora with dark brown median and apical spots, tibiae and tarsi pale yellowish brown, tarsi paler at base and apex; claw with denticles divided into two sections of 8–10+4–5 denticles (Fig. 9) with denticles in both sections increasing in size toward apex. Abdomen: terga washed with dark brown, pale markings as in male imago; sterna yellowish brown, male with dark brown at base of each segment in lateral 1/4; female with sterna washed with orangebrown; posterolateral spines present on segments IV–IX. Gills (Fig. 10) whitish grey, trachea grayviolet, secondary tracheae lacking side branches. Caudal filaments pale yellowish brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3096539FF9AFD2AFEF7F94CFE40F895" pageId="2" pageNumber="61" type="etymology">
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<paragraph id="8BAC36B2FF9AFD2AFEF7F94CFE6CF91E" blockId="2.[264,1321,1684,1830]" box="[264,389,1684,1709]" pageId="2" pageNumber="61">
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<emphasis id="B967EAA0FF9AFD2AFEF7F94CFE6CF91E" box="[264,389,1684,1709]" italics="true" pageId="2" pageNumber="61">Etymology</emphasis>
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</paragraph>
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<paragraph id="8BAC36B2FF9AFD2AFEC7F964FE40F895" blockId="2.[264,1321,1684,1830]" pageId="2" pageNumber="61">
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<taxonomicName id="4C134D31FF9AFD2AFEC7F964FE75F966" box="[312,412,1724,1749]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="gregoryi">
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<emphasis id="B967EAA0FF9AFD2AFEC7F964FE75F966" box="[312,412,1724,1749]" italics="true" pageId="2" pageNumber="61">gregoryi</emphasis>
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</taxonomicName>
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, masculine. This species is dedicated to the senior author’s youngest son, Gregory Juseppy Porras, who at six years old is already an enthusiastic aquatic entomologist.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3096539FF9BFD2CFEF7FA2BFDEBFC3D" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="description">
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<paragraph id="8BAC36B2FF9BFD2BFEF7FA2BFB38F9CF" blockId="3.[264,1323,1523,1660]" pageId="3" pageNumber="62">
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<emphasis id="B967EAA0FF9BFD2BFEF7FA2BFE2FF9B8" bold="true" box="[264,454,1523,1547]" pageId="3" pageNumber="62">FIGURES 1–12.</emphasis>
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<taxonomicName id="4C134D31FF9BFD2BFE2DFA2BFDBFF9B9" box="[466,598,1523,1546]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="B967EAA0FF9BFD2BFE2DFA2BFDBFF9B9" box="[466,598,1523,1546]" italics="true" pageId="3" pageNumber="62">Choroterpes</emphasis>
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</taxonomicName>
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. 1, 3–5, 9–10.
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<taxonomicName id="4C134D31FF9BFD2BFCF6FA2BFC61F9B9" box="[777,904,1523,1546]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="gregoryi">
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<emphasis id="B967EAA0FF9BFD2BFCF6FA2BFC61F9B9" box="[777,904,1523,1546]" italics="true" pageId="3" pageNumber="62">C. gregoryi</emphasis>
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</taxonomicName>
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; 1, wings; 3, tergites II–VII of male abdomen; 4, male genitalia; 5, segments II–VII of male abdomen, lateral; 9, gill IV; 10, foreclaw of nymph. 2, 6–8, 11–12
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<taxonomicName id="4C134D31FF9BFD2BFE04F9E6FD9BF9E6" box="[507,626,1598,1621]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="mairena">
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<emphasis id="B967EAA0FF9BFD2BFE04F9E6FD9BF9E6" box="[507,626,1598,1621]" italics="true" pageId="3" pageNumber="62">C. mairena</emphasis>
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</taxonomicName>
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; 2, wings; 6, tergites II–VII of male abdomen; 7, male genitalia; 8, segments II–VII of male abdomen, lateral; 11, gill IV of nymph; 12, foreclaw of nymph.
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</paragraph>
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<paragraph id="8BAC36B2FF9BFD2BFEF7F96BFE15F97F" blockId="3.[264,1323,1715,1901]" box="[264,508,1715,1740]" pageId="3" pageNumber="62">
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||
<emphasis id="B967EAA0FF9BFD2BFEF7F96BFE15F97F" box="[264,508,1715,1740]" italics="true" pageId="3" pageNumber="62">Specimens examined</emphasis>
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</paragraph>
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<paragraph id="8BAC36B2FF9BFD2CFEC7F903FE01FDC5" blockId="3.[264,1323,1715,1901]" lastBlockId="4.[264,1323,283,630]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
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Male imago
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<typeStatus id="54A88810FF9BFD2BFE24F903FD9FF946" box="[475,630,1755,1781]" pageId="3" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
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labeled
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<collectingCountry id="F3047622FF9BFD2BFD17F903FC74F946" box="[744,925,1755,1781]" name="Costa Rica" pageId="3" pageNumber="62">COSTA RICA</collectingCountry>
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, Guanacaste, Nandayure, Queb. Maquenco. Fecha:
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<date id="FFAD1072FF9BFD2BFE0EF8DBFD6CF8AE" box="[497,645,1795,1821]" pageId="3" pageNumber="62" value="2005-12-16">16XII2005</date>
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Elev.
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<specimenCount id="9D15FD3BFF9BFD2BFD27F8DBFCCBF8AE" box="[728,802,1795,1821]" pageId="3" pageNumber="62" type="generic">
|
||
<quantity id="4CEB9B57FF9BFD2BFD27F8DBFCF7F8AE" box="[728,798,1795,1821]" metricMagnitude="2" metricUnit="m" metricValue="3.92" pageId="3" pageNumber="62" unit="m" value="392.0">392m</quantity>
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.
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</specimenCount>
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<geoCoordinate id="EE275075FF9BFD2BFCD1F8DBFC02F8AE" box="[814,1003,1795,1821]" direction="north" orientation="latitude" pageId="3" pageNumber="62" precision="1" value="9.972633">N 9° 58’21.48”</geoCoordinate>
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<geoCoordinate id="EE275075FF9BFD2BFC08F8DBFB23F8AE" box="[1015,1226,1795,1821]" direction="west" orientation="longitude" pageId="3" pageNumber="62" precision="1" value="-85.259865">W 85° 15’35.52”</geoCoordinate>
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Col. S. Ávila, R. W. Flowers. Female subimago ALLOTYPE labeled
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<collectingCountry id="F3047622FF9BFD2BFC16F8F3FB7EF8F6" box="[1001,1175,1835,1861]" name="Costa Rica" pageId="3" pageNumber="62">COSTA RICA</collectingCountry>
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, Puntarenas, Golfito, Pavones, Queb. El Macho, Fecha:
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<date id="FFAD1072FF9BFD2BFCF7F88BFC7BF8DE" box="[776,914,1875,1901]" pageId="3" pageNumber="62" value="2005-04-16">16IV2005</date>
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, Elev.
|
||
<specimenCount id="9D15FD3BFF9BFD2BFC1CF88BFBF6F8DE" box="[995,1055,1875,1901]" pageId="3" pageNumber="62" type="generic">
|
||
<quantity id="4CEB9B57FF9BFD2BFC1CF88BFBF2F8DE" box="[995,1051,1875,1901]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="62" unit="m" value="40.0">40m</quantity>
|
||
.
|
||
</specimenCount>
|
||
<geoCoordinate id="EE275075FF9BFD2BFBD8F88BFB3CF8DE" box="[1063,1237,1875,1901]" direction="north" orientation="latitude" pageId="3" pageNumber="62" precision="1" value="8.344383">N8° 20’39.78”</geoCoordinate>
|
||
<geoCoordinate id="EE275075FF9BFD2CFB22F88BFE86FE85" direction="west" lastPageId="4" lastPageNumber="63" orientation="longitude" pageId="3" pageNumber="62" precision="1" value="-83.136">W 83° 08’09.6”</geoCoordinate>
|
||
Col. S. vila, Cria
|
||
<date id="FFAD1072FF9CFD2CFDBBFEC4FD26FE85" box="[580,719,284,310]" pageId="4" pageNumber="63" value="2005-04-21">21IV2005</date>
|
||
.
|
||
<typeStatus id="54A88810FF9CFD2CFD1DFEC4FC6DFE85" box="[738,900,284,310]" pageId="4" pageNumber="63" type="paratype">PARATYPES</typeStatus>
|
||
(3 ɗɗ, 1 ΨΨ, 10 nymphs): 2 ɗɗ, 5 nymphs (2 nymphs, MZUCR; 2 nymphs, INBio) same data as
|
||
<typeStatus id="54A88810FF9CFD2CFBF7FE9CFB9BFEED" box="[1032,1138,324,350]" pageId="4" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
; 1 ɗ subimago (reared), 1 Ψ subimago (reared), 1 nymph (INBio), same data as allotype; 3 nymphs (INBio), same locality as allotype, Fecha:
|
||
<date id="FFAD1072FF9CFD2CFCF8FE4CFC96FE1D" box="[775,895,404,430]" pageId="4" pageNumber="63" value="2005-01-18">18I2005</date>
|
||
, S. Ávila, R. W. Flowers; 1 nymph (INBio), Puntarenas Province, Parrita, Queb. Pirrís,
|
||
<specimenCount id="9D15FD3BFF9CFD2CFC98FE64FC4AFE65" box="[871,931,444,470]" pageId="4" pageNumber="63" type="generic">
|
||
<quantity id="4CEB9B57FF9CFD2CFC98FE64FC76FE65" box="[871,927,444,470]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="63" unit="m" value="47.0">47m</quantity>
|
||
.
|
||
</specimenCount>
|
||
<geoCoordinate id="EE275075FF9CFD2CFC55FE64FBBCFE65" box="[938,1109,444,470]" direction="north" orientation="latitude" pageId="4" pageNumber="63" precision="1" value="9.600284">N 9° 36’ 1.02”</geoCoordinate>
|
||
<geoCoordinate id="EE275075FF9CFD2CFBA4FE64FAC2FE65" box="[1115,1323,444,470]" direction="west" orientation="longitude" pageId="4" pageNumber="63" precision="1" value="-84.32197">W 84° 19’ 19.08”</geoCoordinate>
|
||
Fecha:
|
||
<date id="FFAD1072FF9CFD2CFE9EFE3CFE00FE4D" box="[353,489,484,510]" pageId="4" pageNumber="63" value="2005-02-13">13/02/2005</date>
|
||
; 3 nymphs (INBio), San José Province, La Cangreja, Queb. Grande,
|
||
<quantity id="4CEB9B57FF9CFD2CFEF7FDD4FEA7FD95" box="[264,334,524,550]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="63" unit="m" value="200.0">200m</quantity>
|
||
,
|
||
<date id="FFAD1072FF9CFD2CFEA6FDD4FE33FD95" box="[345,474,524,550]" pageId="4" pageNumber="63" value="2005-02-13">13II2005</date>
|
||
, S. Ávila, R. W. Flowers. Nymphs from La Cangreja had a paler ground color on the abdomen and correspondingly darker lateral markings than the nymphs from the other localities.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B2FF9CFD2CFEF7FD74FE81FD76" blockId="4.[264,1323,684,910]" box="[264,360,684,709]" pageId="4" pageNumber="63">
|
||
<emphasis id="B967EAA0FF9CFD2CFEF7FD74FE81FD76" box="[264,360,684,709]" italics="true" pageId="4" pageNumber="63">Ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAC36B2FF9CFD2CFEC7FD0CFDEBFC3D" blockId="4.[264,1323,684,910]" pageId="4" pageNumber="63">
|
||
This species was collected from leaf packs in pools and in slowflowing areas of streams along the southern Pacific coast of
|
||
<collectingCountry id="F3047622FF9CFD2CFCFEFD24FC6CFCA5" box="[769,901,764,790]" name="Costa Rica" pageId="4" pageNumber="63">Costa Rica</collectingCountry>
|
||
. Associated mayfly species include
|
||
<taxonomicName id="4C134D31FF9CFD2CFEF7FCFCFD06FC8D" authority="Avila & Flowers" authorityName="Avila & Flowers" box="[264,751,804,830]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="acosa">
|
||
<emphasis id="B967EAA0FF9CFD2CFEF7FCFCFE10FC8E" box="[264,505,804,829]" italics="true" pageId="4" pageNumber="63">Ulmeritoides acosa</emphasis>
|
||
Ávila & Flowers
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D31FF9CFD2CFCF8FCFCFB98FC8D" authority="Traver" authorityName="Traver" box="[775,1137,804,830]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="atramentum">
|
||
<emphasis id="B967EAA0FF9CFD2CFCF8FCFCFC10FC8E" box="[775,1017,804,829]" italics="true" pageId="4" pageNumber="63">Tikuna atramentum</emphasis>
|
||
(Traver)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D31FF9CFD2CFB76FCFCFB1CFC8E" box="[1161,1269,804,829]" class="Insecta" family="Leptophlebiidae" genus="Farrodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFB76FCFCFB1CFC8E" box="[1161,1269,804,829]" italics="true" pageId="4" pageNumber="63">Farrodes</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
<taxonomicName id="4C134D31FF9CFD2CFEF7FC94FE64FCD6" box="[264,397,844,869]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFEF7FC94FE64FCD6" box="[264,397,844,869]" italics="true" pageId="4" pageNumber="63">Thraulodes</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
<taxonomicName id="4C134D31FF9CFD2CFE3CFC94FDFCFCD6" box="[451,533,844,869]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFE3CFC94FDFCFCD6" box="[451,533,844,869]" italics="true" pageId="4" pageNumber="63">Caenis</emphasis>
|
||
</taxonomicName>
|
||
sp., and
|
||
<taxonomicName id="4C134D31FF9CFD2CFD7BFC94FCCEFCD6" box="[644,807,844,869]" class="Insecta" family="Leptohyphidae" genus="Tricorythodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFD7BFC94FCCEFCD6" box="[644,807,844,869]" italics="true" pageId="4" pageNumber="63">Tricorythodes</emphasis>
|
||
</taxonomicName>
|
||
sp. All streams in which this species was found are permanent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3096539FF9CFD2CFEF7FC1CFE16FA0D" pageId="4" pageNumber="63" type="diagnosis">
|
||
<paragraph id="8BAC36B2FF9CFD2CFEF7FC1CFE96FC6E" blockId="4.[264,1324,964,1470]" box="[264,383,964,989]" pageId="4" pageNumber="63">
|
||
<emphasis id="B967EAA0FF9CFD2CFEF7FC1CFE96FC6E" box="[264,383,964,989]" italics="true" pageId="4" pageNumber="63">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAC36B2FF9CFD2CFEC7FC34FE16FA0D" blockId="4.[264,1324,964,1470]" pageId="4" pageNumber="63">
|
||
<taxonomicName id="4C134D31FF9CFD2CFEC7FC34FDD2FBB6" box="[312,571,1004,1029]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="gregoryi">
|
||
<emphasis id="B967EAA0FF9CFD2CFEC7FC34FDD2FBB6" box="[312,571,1004,1029]" italics="true" pageId="4" pageNumber="63">Choroterpes gregoryi</emphasis>
|
||
</taxonomicName>
|
||
can be separated from other Middle and Central American species of
|
||
<taxonomicName id="4C134D31FF9CFD2CFE73FBCCFDF7FB9E" box="[396,542,1044,1069]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFE73FBCCFDF7FB9E" box="[396,542,1044,1069]" italics="true" pageId="4" pageNumber="63">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters. In the imago: (1) abdomen dark with color pattern as in Figs. 3,5; (2) male genitalia with basal expansion of forceps 1/3 of their total length; (3) apices of penes rounded. In the nymph: (1) mandibles lacking anterolateral expansion basal to incisors; (2) maxilla with 17–18 apical pectinate setae and lacking an anteromedian tooth; (3) abdominal gills (Fig.) with outer and inner processes of both lamellae pointed. Of the other Western Hemisphere
|
||
<taxonomicName id="4C134D31FF9CFD2CFB93FB04FB14FB46" box="[1132,1277,1244,1269]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAA0FF9CFD2CFB93FB04FB14FB46" box="[1132,1277,1244,1269]" italics="true" pageId="4" pageNumber="63">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D31FF9CFD2CFAEFFB04FE85FAAE" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="gregoryi">
|
||
<emphasis id="B967EAA0FF9CFD2CFAEFFB04FE85FAAE" italics="true" pageId="4" pageNumber="63">C. gregoryi</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles dark forms of
|
||
<taxonomicName id="4C134D31FF9CFD2CFCCCFADCFBE4FAAD" authority="Banks" authorityName="Banks" box="[819,1037,1284,1310]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="basalis">
|
||
<emphasis id="B967EAA0FF9CFD2CFCCCFADCFC40FAAE" box="[819,937,1284,1309]" italics="true" pageId="4" pageNumber="63">C. basalis</emphasis>
|
||
(Banks)
|
||
</taxonomicName>
|
||
from the eastern
|
||
<collectingCountry id="F3047622FF9CFD2CFB24FADCFEB9FAF5" name="United States of America" pageId="4" pageNumber="63">United States</collectingCountry>
|
||
, but can be distinguished by the following combination of characters: In the imago the penes of
|
||
<taxonomicName id="4C134D31FF9CFD2CFE5AFA8CFDF5FADE" box="[421,540,1364,1389]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="basalis">
|
||
<emphasis id="B967EAA0FF9CFD2CFE5AFA8CFDF5FADE" box="[421,540,1364,1389]" italics="true" pageId="4" pageNumber="63">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
are more elongate than those of
|
||
<taxonomicName id="4C134D31FF9CFD2CFC52FA8CFBDFFADE" box="[941,1078,1364,1389]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="gregoryi">
|
||
<emphasis id="B967EAA0FF9CFD2CFC52FA8CFBDFFADE" box="[941,1078,1364,1389]" italics="true" pageId="4" pageNumber="63">C. gregoryi</emphasis>
|
||
</taxonomicName>
|
||
and their apices are slanted outward. In the nymph, the labrum of
|
||
<taxonomicName id="4C134D31FF9CFD2CFCCDFAA4FC41FA26" box="[818,936,1404,1429]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="basalis">
|
||
<emphasis id="B967EAA0FF9CFD2CFCCDFAA4FC41FA26" box="[818,936,1404,1429]" italics="true" pageId="4" pageNumber="63">C. basalis</emphasis>
|
||
</taxonomicName>
|
||
lacks the low irregular denticles found in
|
||
<taxonomicName id="4C134D31FF9CFD2CFE8DFA7CFE11FA0E" box="[370,504,1444,1469]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="gregoryi">
|
||
<emphasis id="B967EAA0FF9CFD2CFE8DFA7CFE11FA0E" box="[370,504,1444,1469]" italics="true" pageId="4" pageNumber="63">C. gregoryi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |