169 lines
18 KiB
XML
169 lines
18 KiB
XML
<document id="A3E1E479CC18D58B8758FEB1B86E31B4" ID-DOI="10.5281/zenodo.158702" ID-GBIF-Dataset="64a4a578-1107-4369-b2c4-67921dd0664c" ID-ISSN="11755326" ID-Zenodo-Dep="158702" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461028075636" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2004" docId="03BA879A5D71FFD7FECCD6A97910FA77" docLanguage="en" docName="zt00662.pdf" docOrigin="Zootaxa 662" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Aspasiola steineri Erwin, new species" docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="FF83FFE25D60FFC5FFC4D42D7D5BFF8C" masterDocTitle="The beetle family Carabidae of Costa Rica: The genera of the Cryptobatida group of subtribe Agrina, tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="18" updateTime="1698572776705" updateUser="plazi">
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<mods:titleInfo id="429338B87545810675384AAFA741122F">
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<mods:title id="19E0B348CC9FDD428795698321E85AC1">The beetle family Carabidae of Costa Rica: The genera of the Cryptobatida group of subtribe Agrina, tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="098F9ECD788C5AB844476FDD036FFEEE">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="669C791C1FCF32CE734D102710BFA7F1">text</mods:typeOfResource>
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<mods:title id="0E56575FC8B2516CD076403FF46C1F9C">Zootaxa</mods:title>
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<mods:part id="C94E841DB1B9F3E247D02F1B6901BF98">
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<mods:date id="B3A3B922A7A4AEC0F650708E4A6846AA">2004</mods:date>
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<mods:detail id="707C8C0B157430ABDA4EE2C9F0D591B4" type="volume">
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<mods:number id="28379D5A604932F0EF9F3FE4977AB239">662</mods:number>
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<mods:classification id="510C4DAC490B4427F862376F9C71EBE5">journal article</mods:classification>
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<mods:identifier id="B894CB4327D6A632500047CDBBA1F598" type="DOI">10.5281/zenodo.158702</mods:identifier>
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<mods:identifier id="6B577057274BF76C4F78F100F983B09A" type="GBIF-Dataset">64a4a578-1107-4369-b2c4-67921dd0664c</mods:identifier>
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<mods:identifier id="E9385BA954AD403C4BE2E8BE64780E7C" type="ISSN">11755326</mods:identifier>
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</mods:mods>
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<treatment id="03BA879A5D71FFD7FECCD6A97910FA77" ID-DOI="http://doi.org/10.5281/zenodo.6271464" ID-GBIF-Taxon="119607028" ID-Zenodo-Dep="6271464" LSID="urn:lsid:plazi:treatment:03BA879A5D71FFD7FECCD6A97910FA77" httpUri="http://treatment.plazi.org/id/03BA879A5D71FFD7FECCD6A97910FA77" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection id="C30965075D71FFD4FECCD6A97C8AFD4D" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BAC368C5D71FFD4FECCD6A97F89FD12" blockId="17.[264,722,644,705]" box="[264,722,644,670]" pageId="17" pageNumber="18">
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<heading id="D0E481E05D71FFD4FECCD6A97F89FD12" bold="true" box="[264,722,644,670]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="B967EA9E5D71FFD4FECCD6A97F89FD12" bold="true" box="[264,722,644,670]" pageId="17" pageNumber="18">
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<taxonomicName id="4C134D0F5D71FFD4FECCD6A97F6EFD12" ID-CoL="HBPG" authority="Erwin" authorityName="Erwin" box="[264,565,644,670]" class="Insecta" family="Carabidae" genus="Aspasiola" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="steineri" status="sp. nov.">
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<emphasis id="B967EA9E5D71FFD4FECCD6A97C85FD12" bold="true" box="[264,478,644,670]" italics="true" pageId="17" pageNumber="18">Aspasiola steineri</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25457E55D71FFD4FD86D6A97F89FD12" box="[578,722,644,670]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAC368C5D71FFD4FECCD68A7C8AFD4D" blockId="17.[264,722,644,705]" box="[264,465,679,705]" pageId="17" pageNumber="18">(Figs. 11, 12, 33)</paragraph>
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</subSubSection>
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<subSubSection id="C30965075D71FFD4FECCD6DB7CB5FBA5" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="8BAC368C5D71FFD4FECCD6DB7E81FCED" blockId="17.[264,1325,758,1865]" pageId="17" pageNumber="18">
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<emphasis id="B967EA9E5D71FFD4FECCD6DB7CDAFC9C" bold="true" box="[264,385,758,784]" pageId="17" pageNumber="18">
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<typeStatus id="54A8882E5D71FFD4FECCD6DB7C27FC9C" box="[264,380,758,784]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male,
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<collectingCountry id="F304761C5D71FFD4FE10D6DA7FD8FC9D" box="[468,643,759,785]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
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. HEREDIA, Estación Biológica La
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<taxonomicName id="4C134D0F5D71FFD4FBDAD6DA7939FC9D" box="[1054,1122,759,785]" class="Insecta" family="Carabidae" genus="Aspasiola" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="29" phylum="Arthropoda" rank="species" species="selva">Selva</taxonomicName>
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, 3.0 km S Puerto Viejo, Finca La
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<taxonomicName id="4C134D0F5D71FFD4FE0BD7327F48FCB5" box="[463,531,799,825]" class="Insecta" family="Carabidae" genus="Aspasiola" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="29" phylum="Arthropoda" rank="species" species="selva">Selva</taxonomicName>
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,
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<quantity id="4CEB9B695D71FFD4FDE6D7327FC4FCB5" box="[546,671,799,825]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="17" pageNumber="18" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
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,
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<geoCoordinate id="EE27504B5D71FFD4FD6AD7327E06FCB5" box="[686,861,799,825]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="10.431945">10° 25' 55” N</geoCoordinate>
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,
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<geoCoordinate id="EE27504B5D71FFD4FCA8D7327976FCB5" box="[876,1069,799,825]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-84.00889">084° 00' 32” W</geoCoordinate>
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, LN535500, 268000, March (W.E. Steiner, J.M. Hill, et al.)(
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<collectionCode id="ED02AE495D71FFD4FD02D76A7E71FCED" box="[710,810,839,865]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">NMNH</collectionCode>
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: ADP006027).
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</paragraph>
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<paragraph id="8BAC368C5D71FFD4FEFCD7437CB5FBA5" blockId="17.[264,1325,758,1865]" pageId="17" pageNumber="18">
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<emphasis id="B967EA9E5D71FFD4FEFCD7437FCEFC04" bold="true" box="[312,661,878,904]" pageId="17" pageNumber="18">Derivation of specific name.</emphasis>
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The specific epithet,
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<emphasis id="B967EA9E5D71FFD4FC57D7427EA9FC04" box="[915,1010,879,904]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="4C134D0F5D71FFD4FC57D7427EB5FC04" box="[915,1006,879,904]" class="Insecta" family="Carabidae" genus="Aspasiola" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="steineri">steineri</taxonomicName>
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,
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</emphasis>
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is used as a noun in apposition based on the last name of Warren Steiner, who has contributed significantly to our field knowledge of the species richness in
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<taxonomicName id="4C134D0F5D71FFD4FCCCD7927ED7FC55" box="[776,908,959,985]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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. The
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<collectionCode id="ED02AE495D71FFD4FC14D7927971FC55" box="[976,1066,959,985]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">NMNH</collectionCode>
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collection, where his collection is deposited, vouchers his many years of efforts and includes the
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<typeStatus id="54A8882E5D71FFD4FB46D7CA79EEFB8D" box="[1154,1205,999,1025]" pageId="17" pageNumber="18">type</typeStatus>
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specimen of this new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30965075D71FFD4FEFCD01B7ED2FBDD" box="[312,905,1078,1105]" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BAC368C5D71FFD4FEFCD01B7ED2FBDD" blockId="17.[264,1325,758,1865]" box="[312,905,1078,1105]" pageId="17" pageNumber="18">
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<emphasis id="B967EA9E5D71FFD4FEFCD01B7CA5FBDC" bold="true" box="[312,510,1078,1104]" pageId="17" pageNumber="18">Common name.</emphasis>
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Steiner's colorful fungus dweller.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30965075D71FFD4FEFCD073794CFB45" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="8BAC368C5D71FFD4FEFCD073794CFB45" blockId="17.[264,1325,758,1865]" pageId="17" pageNumber="18">
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<emphasis id="B967EA9E5D71FFD4FEFCD0737CE3FBF4" bold="true" box="[312,440,1118,1144]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
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With the attributes of the genus as described above and elytron bright rufous, with humeral area darkly pigmented with bluish reflections, legs bicolored, femur testaceous, tibia and tarsomeres piceous, apex of femur infuscated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30965075D71FFD7FEFCD0FB7910FA77" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BAC368C5D71FFD4FEFCD0FB7E39F8C5" blockId="17.[264,1325,758,1865]" pageId="17" pageNumber="18">
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<emphasis id="B967EA9E5D71FFD4FEFCD0FB7C94FB7C" bold="true" box="[312,463,1238,1264]" pageId="17" pageNumber="18">Description.</emphasis>
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(Fig. 11). Size small: ABL =
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<quantity id="4CEB9B695D71FFD4FCF5D0FA7EB3FB7D" box="[817,1000,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="4.71" metricValueMax="4.92" metricValueMin="4.5" pageId="17" pageNumber="18" unit="mm" value="4.71" valueMax="4.92" valueMin="4.5">4.5 to 4.92 mm</quantity>
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, SBL =
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<quantity id="4CEB9B695D71FFD4FB8DD0FA79AAFB7D" box="[1097,1265,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="17" pageNumber="18" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2 to 4.5 mm</quantity>
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, TW = 1.83 to 2.0 mm.
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<emphasis id="B967EA9E5D71FFD4FE30D0D27F19FA94" box="[500,578,1279,1304]" italics="true" pageId="17" pageNumber="18">Color:</emphasis>
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Dorsum and venter rufescent, humeri infuscated with purple highlights; palpi, four basal 4 antennomeres dorsally, tibiae and tarsi, antennomeres 5–11 infuscatedblackish, femora and basal 4 antennomeres ventrally testaceous. Metathorasic wings smoky grey.
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<emphasis id="B967EA9E5D71FFD4FE2ED15A7F64FA1C" box="[490,575,1399,1424]" italics="true" pageId="17" pageNumber="18">Luster:</emphasis>
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Ve r y s h i n y.
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<emphasis id="B967EA9E5D71FFD4FD0BD15A7ED7FA1C" box="[719,908,1399,1424]" italics="true" pageId="17" pageNumber="18">Microsculpture:</emphasis>
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Absent.
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<emphasis id="B967EA9E5D71FFD4FC37D15A7967FA1C" box="[1011,1084,1399,1424]" italics="true" pageId="17" pageNumber="18">Head:</emphasis>
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Moderately broader across eyes than pronotum; frontal furrows shallowly impressed, very short; eyes large, produced; frons slightly convex, vertex moderately convex; ultimate labial palpomere slightly securiform; antennae of moderate length, reaching just posterior of humerus.
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<emphasis id="B967EA9E5D71FFD4FB32D1C27C04F9BC" italics="true" pageId="17" pageNumber="18">Prothorax:</emphasis>
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Quadrate, moderately convex, side margin narrowly explanate and slightly produced at level of anterior setigerous pore, slightly sinuate anterior to slightly obtuse hind angle. Base markedly lobed posteriorly.
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<emphasis id="B967EA9E5D71FFD4FD30D24A7ED2F90C" box="[756,905,1639,1664]" italics="true" pageId="17" pageNumber="18">Pterothorax:</emphasis>
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Normal for Agrina, fully winged.
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<emphasis id="B967EA9E5D71FFD4FECCD2A27C12F924" box="[264,329,1679,1704]" italics="true" pageId="17" pageNumber="18">Legs:</emphasis>
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Normal for Agrina.
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<emphasis id="B967EA9E5D71FFD4FD8CD2A27F9BF924" box="[584,704,1679,1704]" italics="true" pageId="17" pageNumber="18">Abdomen:</emphasis>
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Sterna normal for Agrina, mostly glabrous, except normal paired ambulatory setae on sterna 3–5; males and females with two pairs long setae on sternum 6.
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<emphasis id="B967EA9E5D71FFD4FE6BD2F27F3EF974" box="[431,613,1759,1784]" italics="true" pageId="17" pageNumber="18">Male genitalia:</emphasis>
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Phallus (
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<figureCitation id="13282A095D71FFD4FD16D2F27E7CF975" box="[722,807,1759,1785]" captionStart="FIGURE 12" captionStartId="18.[264,368,1107,1131]" captionTargetBox="[310,1264,329,1096]" captionTargetId="figure@18.[275,1312,308,1102]" captionTargetPageId="18" captionText="FIGURE 12. Phallus, dorsal, ventral, left lateral aspects of Aspasiola steineri Erwin n. sp., specimen # CRI 000981559." httpUri="https://zenodo.org/record/158708/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
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) with ostium of 1/3 its length, catopic, apex markedly elongate, rounded; endophallus with flagellum, flagellum without barb. Parameres asymmetric, right very small, left larger.
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</paragraph>
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<caption id="DF6C66045D72FFD7FECCD07E7CA6FB1D" httpUri="https://zenodo.org/record/158708/files/figure.png" pageId="18" pageNumber="19" targetBox="[310,1264,329,1096]" targetPageId="18">
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<paragraph id="8BAC368C5D72FFD7FECCD07E7CA6FB1D" blockId="18.[264,1323,1107,1169]" pageId="18" pageNumber="19">
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<emphasis id="B967EA9E5D72FFD7FECCD07E7CC2FBE7" bold="true" box="[264,409,1107,1131]" pageId="18" pageNumber="19">FIGURE 12.</emphasis>
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Phallus, dorsal, ventral, left lateral aspects of
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<taxonomicName id="4C134D0F5D72FFD7FC54D07E79C1FBE6" authority="Erwin" authorityName="Erwin" box="[912,1178,1107,1130]" class="Insecta" family="Carabidae" genus="Aspasiola" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="steineri" status="sp. nov.">
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<emphasis id="B967EA9E5D72FFD7FC54D07E790AFBE6" box="[912,1105,1107,1130]" italics="true" pageId="18" pageNumber="19">Aspasiola steineri</emphasis>
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Erwin
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</taxonomicName>
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<taxonomicNameLabel id="A25457E55D72FFD7FB66D07E7985FBE6" box="[1186,1246,1107,1130]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
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, specimen # CRI000981559.
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</paragraph>
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</caption>
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<paragraph id="8BAC368C5D72FFD7FEFCD0DD795FFA5F" blockId="18.[264,1325,1264,1531]" pageId="18" pageNumber="19">
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<emphasis id="B967EA9E5D72FFD7FEFCD0DD7C9DFA86" bold="true" box="[312,454,1264,1290]" pageId="18" pageNumber="19">Way of life.</emphasis>
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The known altitudinal range of this species is between 50 and 150 meters above sea level. They have been collected in March and October hence they occur as adults in both the dry and rainy season in the lowlands east of the Cordillera Central.
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<emphasis id="B967EA9E5D72FFD7FEFCD1457FC9FA0E" bold="true" box="[312,658,1384,1410]" pageId="18" pageNumber="19">Other specimens examined.</emphasis>
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<typeStatus id="54A8882E5D72FFD7FD5FD1447E5DFA0F" box="[667,774,1385,1411]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
|
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:
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<collectingCountry id="F304761C5D72FFD7FCD6D1447E99FA0F" box="[786,962,1385,1411]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
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.
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<specimenCount id="9D15FD055D72FFD7FC0BD1447979FA0F" box="[975,1058,1385,1411]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
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, LIMÓN, Cariari, Sector Cerro Cocori, 30.0 km N Finca de E. Rojas,
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<quantity id="4CEB9B695D72FFD7FC97D1BC7EF8FA27" box="[851,931,1425,1451]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="m" value="150.0">150 m</quantity>
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,
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<geoCoordinate id="EE27504B5D72FFD7FC75D1BC7907FA27" box="[945,1116,1425,1451]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="15" value="10.591389">10° 35' 29” N</geoCoordinate>
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,
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<geoCoordinate id="EE27504B5D72FFD7FBAED1BC7872FA27" box="[1130,1321,1425,1451]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="15" value="-83.70805">083° 42' 29” W</geoCoordinate>
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, LN285700,568400, October (E. Rojas)(INBIO: CRI000981559).
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</paragraph>
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<paragraph id="8BAC368C5D72FFD7FEFCD1CD7910FA77" blockId="18.[264,1325,1264,1531]" box="[312,1099,1504,1531]" pageId="18" pageNumber="19">
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||
<emphasis id="B967EA9E5D72FFD7FEFCD1CD7F36FA76" bold="true" box="[312,621,1504,1530]" pageId="18" pageNumber="19">Geographic distribution.</emphasis>
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(
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<figureCitation id="13282A095D72FFD7FDB9D1CC7F8EFA77" box="[637,725,1505,1531]" captionStart="FIGURE 33" captionStartId="28.[268,372,525,549]" captionTargetBox="[556,1316,326,1010]" captionTargetId="figure@28.[556,1316,326,1010]" captionTargetPageId="28" captionText="FIGURE 33. Geographical distribution map of the species of Aspasiola in Costa Rica: A. bonita Erwin n. sp. (solid circle," httpUri="https://zenodo.org/record/169370/files/figure.png" pageId="18" pageNumber="19">Fig. 33</figureCitation>
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||
). Known only from
|
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<collectingCountry id="F304761C5D72FFD7FC07D1CC7913FA77" box="[963,1096,1505,1531]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |