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<document id="720DA39B5678279EF4D93A5CA1A1C0E2" ID-DOI="10.5281/zenodo.195835" ID-GBIF-Dataset="5fe3da15-3dd3-495f-9c15-7f9d606315aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="195835" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460234920266" checkinUser="plazi" docAuthor="Emberson, Rowan M." docDate="2010" docId="03997670FFB8E6648ECBFA0A765F6C74" docLanguage="en" docName="zt02501p053.pdf" docOrigin="Zootaxa 2501" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Geholaspis Berlese 1918" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FFA00E08FFBBE6608E5CFFF873136F26" masterDocTitle="A reappraisal of some basal lineages of the family Macrochelidae, with the description of a new genus (Acarina: Mesostigmata)" masterLastPageNumber="53" masterPageNumber="37" pageNumber="40" updateTime="1698239325266" updateUser="plazi">
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<mods:title id="607C64637992BC3ABA309A7A457CBDDC">A reappraisal of some basal lineages of the family Macrochelidae, with the description of a new genus (Acarina: Mesostigmata)</mods:title>
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<mods:namePart id="2A7BA270A156F1DA0DC9F9758EAE7EA5">Emberson, Rowan M.</mods:namePart>
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<mods:date id="9D0B43709D0F7E2CB0B3160A55BA2D60">2010</mods:date>
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<mods:classification id="27285EC1AB3CB39F051897FBF3A29313">journal article</mods:classification>
<mods:identifier id="557188C1B1236A756F2384B37DD0610E" type="DOI">10.5281/zenodo.195835</mods:identifier>
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<treatment id="03997670FFB8E6648ECBFA0A765F6C74" ID-DOI="http://doi.org/10.5281/zenodo.6212244" ID-GBIF-Taxon="119387876" ID-Zenodo-Dep="6212244" LSID="urn:lsid:plazi:treatment:03997670FFB8E6648ECBFA0A765F6C74" httpUri="http://treatment.plazi.org/id/03997670FFB8E6648ECBFA0A765F6C74" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
<subSubSection id="C32A94EDFFB8E6638ECBFA0A71856862" pageId="3" pageNumber="40" type="nomenclature">
<paragraph id="8B8FC766FFB8E6638ECBFA0A72DB692A" blockId="3.[151,456,1522,1548]" box="[151,456,1522,1548]" pageId="3" pageNumber="40">
<heading id="D0C7700AFFB8E6638ECBFA0A72DB692A" bold="true" box="[151,456,1522,1548]" fontSize="11" level="1" pageId="3" pageNumber="40" reason="1">
<taxonomicName id="4C30BCE5FFB8E6638ECBFA0A72DB692A" authority="Berlese, 1918" authorityName="Berlese" authorityYear="1918" box="[151,456,1522,1548]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB8E6638ECBFA0A72DB692A" bold="true" box="[151,456,1522,1548]" pageId="3" pageNumber="40">
<emphasis id="B9441B74FFB8E6638ECBFA0A720E692A" bold="true" box="[151,285,1522,1548]" italics="true" pageId="3" pageNumber="40">Geholaspis</emphasis>
<bibRefCitation id="EFA1BA97FFB8E6638F78FA0A72DB692A" author="Berlese" box="[292,456,1522,1548]" pageId="3" pageNumber="51" refString="Berlese, A. (1918) Centuria quarta di Acari nuovi. Redia, 13, 115 - 192." type="journal article" year="1918">Berlese, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8FC766FFB8E6638ECBF9C271526997" pageId="3" pageNumber="40">
<table id="F93035C6FFB8199F8ECBF9C2768F6997" box="[151,1436,1594,1713]" gridcols="4" gridrows="4" pageId="3" pageNumber="40">
<tr id="3500C524FFB8199F8ECBF9C2768F6977" box="[151,1436,1594,1617]" gridrow="0" pageId="3" pageNumber="40">
<th id="76D1AC58FFB8199F8ECBF9C2717F6977" box="[151,620,1594,1617]" gridcol="0" gridrow="0" pageId="3" pageNumber="40">
<taxonomicName id="4C30BCE5FFB8E6638ECBF9C271766977" authority="Berlese, 1918" authorityName="Berlese" authorityYear="1918" box="[151,613,1594,1617]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="subGenus" subGenus="Geholaspis">
<emphasis id="B9441B74FFB8E6638ECBF9C272336977" box="[151,288,1594,1617]" italics="true" pageId="3" pageNumber="40">Macrocheles</emphasis>
(
<emphasis id="B9441B74FFB8E6638F6BF9C272BC6977" box="[311,431,1594,1617]" italics="true" pageId="3" pageNumber="40">Geholaspis</emphasis>
) Berlese, 1918
</taxonomicName>
:
</th>
<th id="76D1AC58FFB8199F8C28F9C270D66977" box="[628,965,1594,1617]" gridcol="1" gridrow="0" pageId="3" pageNumber="40">
145. Type species:
<taxonomicName id="4C30BCE5FFB8E6638D00F9C270AC6977" box="[860,959,1594,1617]" class="Arachnida" family="Ascidae" genus="Gamasus" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB8E6638D00F9C270AC6977" box="[860,959,1594,1617]" italics="true" pageId="3" pageNumber="40">Gamasus</emphasis>
</taxonomicName>
</th>
<taxonomicName id="4C30BCE5FFB8E6638D92F9C2761E6977" authority="Kramer, 1876" authorityName="Kramer" authorityYear="1876" box="[974,1293,1594,1617]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="longispinosus">
<th id="76D1AC58FFB8199F8D91F9C277726977" box="[973,1121,1594,1617]" gridcol="2" gridrow="0" pageId="3" pageNumber="40">
<emphasis id="B9441B74FFB8E6638D92F9C277726977" box="[974,1121,1594,1617]" italics="true" pageId="3" pageNumber="40">longispinosus</emphasis>
</th>
<th id="76D1AC58FFB8199F8A35F9C2768F6977" box="[1129,1436,1594,1617]" gridcol="3" gridrow="0" pageId="3" pageNumber="40">Kramer, 1876, by original</th>
</taxonomicName>
</tr>
<tr id="3500C524FFB8199F8ECBF9A2768F6957" box="[151,1436,1626,1649]" gridrow="1" pageId="3" pageNumber="40" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76D1AC58FFB8199F8ECBF9A2717F6957" box="[151,620,1626,1649]" gridcol="0" gridrow="1" pageId="3" pageNumber="40">designation.</th>
</tr>
<tr id="3500C524FFB8199F8ECBF982768F69B7" box="[151,1436,1658,1681]" gridrow="2" pageId="3" pageNumber="40">
<th id="76D1AC58FFB8199F8ECBF982717F69B7" box="[151,620,1658,1681]" gridcol="0" gridrow="2" pageId="3" pageNumber="40">
<taxonomicName id="4C30BCE5FFB8E6638ECBF982717469B7" authority="Valle, 1953: 323" authorityName="Valle" authorityPageNumber="323" authorityYear="1953" box="[151,615,1658,1681]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="subGenus" subGenus="Cyrtocheles">
<emphasis id="B9441B74FFB8E6638ECBF982721C69B7" box="[151,271,1658,1681]" italics="true" pageId="3" pageNumber="40">Geholaspis</emphasis>
(
<emphasis id="B9441B74FFB8E6638F7CF982728C69B7" box="[288,415,1658,1681]" italics="true" pageId="3" pageNumber="40">Cyrtocheles</emphasis>
) Valle, 1953: 323
</taxonomicName>
,
</th>
<td id="76D1AC58FFB8199F8C28F98270D669B7" box="[628,965,1658,1681]" gridcol="1" gridrow="2" pageId="3" pageNumber="40">
340. Type species:
<taxonomicName id="4C30BCE5FFB8E6638D1BF98270D669B7" box="[839,965,1658,1681]" class="Arachnida" family="Macrochelidae" genus="Holostaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB8E6638D1BF98270D669B7" box="[839,965,1658,1681]" italics="true" pageId="3" pageNumber="40">Holostaspis</emphasis>
</taxonomicName>
</td>
<td id="76D1AC58FFB8199F8D91F982777269B7" box="[973,1121,1658,1681]" gridcol="2" gridrow="2" pageId="3" pageNumber="40">
<taxonomicName id="4C30BCE5FFB8E6638D91F982777369B7" box="[973,1120,1658,1681]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="B9441B74FFB8E6638D91F982777369B7" box="[973,1120,1658,1681]" italics="true" pageId="3" pageNumber="40">longispinosus</emphasis>
</taxonomicName>
</td>
<td id="76D1AC58FFB8199F8A35F982768F69B7" box="[1129,1436,1658,1681]" gridcol="3" gridrow="2" pageId="3" pageNumber="40">
<taxonomicName id="4C30BCE5FFB8E6638A35F982766669B7" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[1129,1397,1658,1681]" pageId="3" pageNumber="40" rank="variety" variety="asper">
var.
<emphasis id="B9441B74FFB8E6638AC5F98277C769B7" box="[1177,1236,1658,1681]" italics="true" pageId="3" pageNumber="40">asper</emphasis>
Berlese, 1904
</taxonomicName>
, by
</td>
</tr>
<tr id="3500C524FFB8199F8ECBF962768F6997" box="[151,1436,1690,1713]" gridrow="3" pageId="3" pageNumber="40" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76D1AC58FFB8199F8ECBF962717F6997" box="[151,620,1690,1713]" gridcol="0" gridrow="3" pageId="3" pageNumber="40">original designation and monotypy.</th>
</tr>
</table>
</paragraph>
<paragraph id="8B8FC766FFB8E6638ECBF91871856862" blockId="3.[151,1437,1760,2010]" pageId="3" pageNumber="40">
Geholaspini with normally developed, robust chelicerae, fixed chelae with fewer than six teeth (
<figureCitation id="130BDBE3FFB8E6638B1CF918769F69DC" box="[1344,1420,1760,1786]" captionStart="FIGURES 1 – 7. 1 – 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 – 7. 1 – 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 – 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="3" pageNumber="40">Fig. 4</figureCitation>
); epistome triangular basally, with elongate median process (
<figureCitation id="130BDBE3FFB8E6638D02F8FD70B56839" box="[862,934,1797,1823]" captionStart="FIGURES 1 – 7. 1 – 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 – 7. 1 – 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 – 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="3" pageNumber="40">Fig. 5</figureCitation>
), or narrow, bifid median process; ventrianal shield with five pairs of pre-anal setae.
</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFB8E6648E98F8A8765F6C74" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="description">
<paragraph id="8B8FC766FFB8E6648E98F8A871B76D01" blockId="3.[151,1437,1760,2010]" lastBlockId="4.[151,1437,152,850]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
The dorsal shield has 28 pairs of setae, which may be either aciculate, lightly plumose, or distinctly plumose. A group of aciculate setae is present in the centre of the shield including at least j5, j6, z5, z6, and J2. Setae j1 are relatively short and plumose, while setae z1 are long and usually aciculate, but may be lightly plumose (
<taxonomicName id="4C30BCE5FFB8E6638F50F838711768FC" authority="Berlese" authorityName="Berlese" box="[268,516,1984,2010]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="pauperior">
<emphasis id="B9441B74FFB8E6638F50F838720D68FF" box="[268,286,1984,2009]" italics="true" pageId="3" pageNumber="40">G</emphasis>
.
<emphasis id="B9441B74FFB8E6638F70F83872B268FF" box="[300,417,1984,2009]" italics="true" pageId="3" pageNumber="40">pauperior</emphasis>
Berlese
</taxonomicName>
,
<taxonomicName id="4C30BCE5FFB8E6638C4CF83870BF68FC" authority="Bregetova &amp; Koroleva" authorityName="Bregetova &amp; Koroleva" box="[528,940,1984,2010]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9441B74FFB8E6638C4CF838713168FF" box="[528,546,1984,2009]" italics="true" pageId="3" pageNumber="40">G</emphasis>
.
<emphasis id="B9441B74FFB8E6638C6DF838718668FF" box="[561,661,1984,2009]" italics="true" pageId="3" pageNumber="40">ponticus</emphasis>
Bregetova &amp; Koroleva
</taxonomicName>
). Setae j5 are displaced posteriorly so that they lie behind setae z5; they may be either anterior or just posterior to setae j6. The ventral sclerotisation is well developed; the metasternal platelets may either be free with the usual seta and pore (subgenus
<emphasis id="B9441B74FFBFE6648B44FF4573B16FDD" italics="true" pageId="4" pageNumber="41">
<taxonomicName id="4C30BCE5FFBFE6648B44FF45768F6FF0" box="[1304,1436,189,214]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus" sensu="stricto" status="s">Geholaspis</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBFE6648ECBFF1A73B16FDD" box="[151,162,226,251]" pageId="4" pageNumber="41" sensu="stricto">s</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B9441B74FFBFE6648EEFFF1A73C16FDD" box="[179,210,226,251]" italics="true" pageId="4" pageNumber="41">str</emphasis>
.), or fused to the endopodal shields (subgenus
<taxonomicName id="4C30BCE5FFBFE6648D48FF1A70B66FDD" box="[788,933,226,251]" class="Arachnida" family="Macrochelidae" genus="Cyrtocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648D48FF1A70B66FDD" box="[788,933,226,251]" italics="true" pageId="4" pageNumber="41">Cyrtocheles</emphasis>
</taxonomicName>
). The epigynal shield is well sclerotised, ornamented with a punctate lineal pattern and has a truncate posterior margin. The ventri-anal shield is usually broader than long, and bears five pairs of pre-anal setae and the inguinal pores. The chelicerae are robust, with fewer than six teeth on the fixed chela (
<figureCitation id="130BDBE3FFBFE6648C3EFEAA71B96E4A" box="[610,682,338,364]" captionStart="FIGURES 1 – 7. 1 – 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 – 7. 1 – 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 – 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
). The epistome is basically triangular and may have an elongate, tapering median process with distinct shoulders (i.e.
<taxonomicName id="4C30BCE5FFBFE6648D49FE8077406EB4" authority="Kramer" authorityName="Kramer" box="[789,1107,376,402]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="B9441B74FFBFE6648D49FE8070346EB7" box="[789,807,376,401]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648D6BFE8070C86EB7" box="[823,987,376,401]" italics="true" pageId="4" pageNumber="41">longispinosus</emphasis>
(Kramer)
</taxonomicName>
(
<figureCitation id="130BDBE3FFBFE6648A39FE8077BE6EB4" box="[1125,1197,376,402]" captionStart="FIGURES 1 – 7. 1 – 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 – 7. 1 – 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 – 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 5</figureCitation>
)), or dentate lateral margins and a narrow, distally bifid median process. Spermathecal structures are usually not apparent, but the elongate tubuli and rami of one species have been illustrated by
<bibRefCitation id="EFA1BA97FFBFE6648DCCFE3A778A6EFA" author="Athias-Henriot" box="[912,1177,450,476]" pageId="4" pageNumber="51" refString="Athias-Henriot, C. (1968) L'appareil d'insemination laelapoide (Acariens anactinotriches: Laelapoidea, P). Premieres observations. Possibilite d'emploi a des fins taxonomiques. Bulletin scientifique de Bourgogne, 25, 229 - 274." type="journal article" year="1968">Athias-Henriot (1968)</bibRefCitation>
. Males, where known (
<taxonomicName id="4C30BCE5FFBFE6648EFDFE10729F6D24" authority="Berlese" authorityName="Berlese" box="[161,396,488,514]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B9441B74FFBFE6648EFDFE1073A06D27" box="[161,179,488,513]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648E9DFE1072046D27" box="[193,279,488,513]" italics="true" pageId="4" pageNumber="41">alpinus</emphasis>
(Berlese)
</taxonomicName>
,
<taxonomicName id="4C30BCE5FFBFE6648FC5FE1071B56D24" authority="Krauss" authorityName="Krauss" box="[409,678,488,514]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="biperforatus">
<emphasis id="B9441B74FFBFE6648FC5FE1072B86D27" box="[409,427,488,513]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648FE5FE1071596D27" box="[441,586,488,513]" italics="true" pageId="4" pageNumber="41">biperforatus</emphasis>
Krauss
</taxonomicName>
,
<taxonomicName id="4C30BCE5FFBFE6648CEEFE1070266D27" box="[690,821,488,514]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9441B74FFBFE6648CEEFE1071D76D27" box="[690,708,488,513]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648C8EFE1070266D27" box="[722,821,488,513]" italics="true" pageId="4" pageNumber="41">ponticus</emphasis>
</taxonomicName>
), have holoventral shields, a short, dorsally directed spermatodactyl and a small spur on femur II.
</paragraph>
<paragraph id="8B8FC766FFBFE6648E9AFDCA776F6D9A" blockId="4.[151,1437,152,850]" pageId="4" pageNumber="41">
<emphasis id="B9441B74FFBFE6648E9AFDCA72006D6A" bold="true" box="[198,275,562,588]" pageId="4" pageNumber="41">Notes.</emphasis>
<taxonomicName id="4C30BCE5FFBFE6648F46FDCA728D6D6D" box="[282,414,562,587]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648F46FDCA728D6D6D" box="[282,414,562,587]" italics="true" pageId="4" pageNumber="41">Geholaspis</emphasis>
</taxonomicName>
is a mainly Western Palaearctic genus, known from
<collectingCountry id="F32787F6FFBFE6648A5EFDCA77466D6A" box="[1026,1109,562,588]" name="Ireland" pageId="4" pageNumber="41">Ireland</collectingCountry>
to the Caucasus (
<bibRefCitation id="EFA1BA97FFBFE6648B7EFDCA72686D54" author="Bregetova" pageId="4" pageNumber="51" refString="Bregetova, N. G. &amp; Koroleva, E. V. (1960) The macrochelid mites (Gamasoidea, Macrochelidae) in the USSR. Akademiia Nauk SSSR Zoologicheskii institut Parazitologicheskii Sbornik, 29, 32 - 154. [in Russian]." type="journal article" year="1960">Bregetova &amp; Koroleva, 1960</bibRefCitation>
;
<bibRefCitation id="EFA1BA97FFBFE6648FD5FDA071D66D54" author="Hyatt" box="[393,709,600,626]" pageId="4" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson, 1988</bibRefCitation>
), but it is apparently most diverse in Central and Southern Europe (
<bibRefCitation id="EFA1BA97FFBFE6648EA0FD85728E6DB1" author="Krauss" box="[252,413,637,663]" pageId="4" pageNumber="52" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918. (Eine systematische Studie aus dem Jahr 1960). Acarologie, Schriftenreihe fur Vergleichende Milbenkunde, 14, 1 - 43 + 20 plates." type="book chapter" year="1970">Krauss, 1970</bibRefCitation>
;
<bibRefCitation id="EFA1BA97FFBFE6648FF7FD85715A6DB1" author="Masan" box="[427,585,637,663]" pageId="4" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
). The genus is also recorded from North
<collectingCountry id="F32787F6FFBFE6648A6DFD8577896DB1" box="[1073,1178,637,663]" name="United States of America" pageId="4" pageNumber="41">America</collectingCountry>
, and
<taxonomicName id="4C30BCE5FFBFE6648A84FD85768F6DB0" box="[1240,1436,637,663]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="B9441B74FFBFE6648A84FD8577F96DB0" box="[1240,1258,637,662]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648AA6FD85768F6DB0" box="[1274,1436,637,662]" italics="true" pageId="4" pageNumber="41">longispinosus</emphasis>
</taxonomicName>
has established in
<collectingCountry id="F32787F6FFBFE6648F31FD5A711F6D9A" box="[365,524,674,700]" name="New Zealand" pageId="4" pageNumber="41">New Zealand</collectingCountry>
, evidently through human agency (
<bibRefCitation id="EFA1BA97FFBFE6648DF4FD5A777C6D9A" author="Emberson" box="[936,1135,674,700]" pageId="4" pageNumber="51" refString="Emberson, R. M. (1973) Additions to the macrochelid mites in N. Z. (Acarina: Mesostigmata: Macrochelidae). New Zealand Entomologist, 5, 294 - 302." type="journal article" year="1973">Emberson, 1973</bibRefCitation>
).
</paragraph>
<paragraph id="8B8FC766FFBFE6648E9AFD3F765F6C74" blockId="4.[151,1437,152,850]" pageId="4" pageNumber="41">
<emphasis id="B9441B74FFBFE6648E9AFD3F728C6DC7" bold="true" box="[198,415,711,737]" pageId="4" pageNumber="41">Included species:</emphasis>
Subgenus
<emphasis id="B9441B74FFBFE6648C79FD3071AD6DC7" box="[549,702,712,737]" italics="true" pageId="4" pageNumber="41">
<taxonomicName id="4C30BCE5FFBFE6648C79FD3071BA6DC7" box="[549,681,712,737]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus" sensu="stricto" status="s">Geholaspis</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBFE6648CEFFD3071AD6DC7" box="[691,702,712,737]" pageId="4" pageNumber="41" sensu="stricto">s</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B9441B74FFBFE6648C8CFD3071FE6DC7" box="[720,749,712,737]" italics="true" pageId="4" pageNumber="41">str</emphasis>
.:
<taxonomicName id="4C30BCE5FFBFE6648D5BFD3077356DC4" authority="Krauss, 1970" authorityName="Krauss" authorityYear="1970" box="[775,1062,712,738]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="aeneus">
<emphasis id="B9441B74FFBFE6648D5BFD30700A6DC7" box="[775,793,712,737]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648D76FD30706F6DC7" box="[810,892,712,737]" italics="true" pageId="4" pageNumber="41">aeneus</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648DD9FD3077356DC4" author="Krauss" box="[901,1062,712,738]" pageId="4" pageNumber="52" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918. (Eine systematische Studie aus dem Jahr 1960). Acarologie, Schriftenreihe fur Vergleichende Milbenkunde, 14, 1 - 43 + 20 plates." type="book chapter" year="1970">Krauss, 1970</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648A69FD3076676DC4" authority="Berlese, 1887" authorityName="Berlese" authorityYear="1887" box="[1077,1396,712,738]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B9441B74FFBFE6648A69FD3077546DC7" box="[1077,1095,712,737]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648A05FD3077BC6DC7" box="[1113,1199,712,737]" italics="true" pageId="4" pageNumber="41">alpinus</emphasis>
(
<bibRefCitation id="EFA1BA97FFBFE6648A9DFD3076786DC4" author="Berlese" box="[1217,1387,712,738]" pageId="4" pageNumber="51" refString="Berlese, A. (1887) Acari, Myriopoda et Scorpiones hucusque in Italiana reperta, 43, 9, 10." type="journal article" year="1887">Berlese, 1887</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648BDFFD3072846C21" authority="Valle, 1953" authorityName="Valle" authorityYear="1953" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="berlesei">
<emphasis id="B9441B74FFBFE6648BDFFD3076866DC7" box="[1411,1429,712,737]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648ECBFD1573E86C20" box="[151,251,749,774]" italics="true" pageId="4" pageNumber="41">berlesei</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648F58FD1572846C21" author="Valle" box="[260,407,749,775]" pageId="4" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle, 1953</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648FF9FD1570046C21" authority="Krauss, 1970" authorityName="Krauss" authorityYear="1970" box="[421,791,749,775]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="biperforatus">
<emphasis id="B9441B74FFBFE6648FF9FD1572AB6C20" box="[421,440,749,774]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648F95FD1571766C20" box="[457,613,749,774]" italics="true" pageId="4" pageNumber="41">biperforatus</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648C31FD1570046C21" author="Krauss" box="[621,791,749,775]" pageId="4" pageNumber="52" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918. (Eine systematische Studie aus dem Jahr 1960). Acarologie, Schriftenreihe fur Vergleichende Milbenkunde, 14, 1 - 43 + 20 plates." type="book chapter" year="1970">Krauss, 1970</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648D79FD1577CB6C21" authority="Lombardini, 1962" authorityName="Lombardini" authorityYear="1962" box="[805,1240,749,775]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="comelicensis">
<emphasis id="B9441B74FFBFE6648D79FD15702B6C20" box="[805,824,749,774]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648D15FD1570FA6C20" box="[841,1001,749,774]" italics="true" pageId="4" pageNumber="41">comelicensis</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648DAEFD1577CB6C21" author="Lombardini" box="[1010,1240,749,775]" pageId="4" pageNumber="53" refString="Lombardini, G. (1962) Acari nuovi del Comelico (Alpi orientali). Annali Centro di Economia Montana delle Ven ezie, Padova, 2, 189 - 216." type="journal article" year="1962">Lombardini, 1962</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648ABBFD15727E6C0A" authority="Lombardini, 1943" authorityName="Lombardini" authorityYear="1943" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="forolivensis">
<emphasis id="B9441B74FFBFE6648ABBFD1577E96C20" box="[1255,1274,749,774]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648B57FD15768E6C20" box="[1291,1437,749,774]" italics="true" pageId="4" pageNumber="41">forolivensis</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648ECBFCEA727E6C0A" author="Lombardini" box="[151,365,786,812]" pageId="4" pageNumber="53" refString="Lombardini, G. (1943) Acari della Collezione Zangheri, Fauna di Romagnola. Bolletino della Societa Entomologica Italiana, 75, 17 - 22." type="journal article" year="1943">Lombardini, 1943</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648F25FCEA71EA6C0A" authority="Kramer, 1876" authorityName="Kramer" authorityYear="1876" box="[377,761,786,812]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="B9441B74FFBFE6648F25FCEA72986C0D" box="[377,395,786,811]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648FC5FCEA71296C0D" box="[409,570,786,811]" italics="true" pageId="4" pageNumber="41">longispinosus</emphasis>
(
<bibRefCitation id="EFA1BA97FFBFE6648C14FCEA71E36C0A" author="Kramer" box="[584,752,786,812]" pageId="4" pageNumber="52" refString="Kramer, P. (1876) Zur Naturgeschichte einiger Gattungen aus der Familie der Gamasiden. Archiv fur Naturgeschichte, 42, 46 - 105." type="journal article" year="1876">Kramer, 1876</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648D58FCEA77576C0A" authority="Berlese, 1918" authorityName="Berlese" authorityYear="1918" box="[772,1092,786,812]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pauperior">
<emphasis id="B9441B74FFBFE6648D58FCEA70056C0D" box="[772,790,786,811]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648D78FCEA708A6C0D" box="[804,921,786,811]" italics="true" pageId="4" pageNumber="41">pauperior</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648DFCFCEA77576C0A" author="Berlese" box="[928,1092,786,812]" pageId="4" pageNumber="51" refString="Berlese, A. (1918) Centuria quarta di Acari nuovi. Redia, 13, 115 - 192." type="journal article" year="1918">Berlese, 1918</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648A13FCEA768B6C0A" authority="Berlese, 1889" authorityName="Berlese" authorityYear="1889" box="[1103,1432,786,812]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="terrestris">
<emphasis id="B9441B74FFBFE6648A13FCEA77726C0D" box="[1103,1121,786,811]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648A33FCEA77CA6C0D" box="[1135,1241,786,811]" italics="true" pageId="4" pageNumber="41">terrestris</emphasis>
(
<bibRefCitation id="EFA1BA97FFBFE6648AB4FCEA769C6C0A" author="Berlese" box="[1256,1423,786,812]" pageId="4" pageNumber="51" refString="Berlese, A. (1889) Acari, Myriopoda et Scorpiones hucusque in Italiana reperta, 52, 7." type="journal article" year="1889">Berlese, 1889</bibRefCitation>
)
</taxonomicName>
. Subgenus
<taxonomicName id="4C30BCE5FFBFE6648F4CFCC0713E6C74" authority="Valle, 1953" authorityName="Valle" authorityYear="1953" box="[272,557,824,850]" class="Arachnida" family="Macrochelidae" genus="Cyrtocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648F4CFCC0728F6C77" box="[272,412,824,849]" italics="true" pageId="4" pageNumber="41">Cyrtocheles</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648FFFFCC0713E6C74" author="Valle" box="[419,557,824,850]" pageId="4" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle, 1953</bibRefCitation>
</taxonomicName>
:
<taxonomicName id="4C30BCE5FFBFE6648C64FCC0704A6C74" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[568,857,824,850]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B9441B74FFBFE6648C64FCC071596C77" box="[568,586,824,849]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648C05FCC071896C77" box="[601,666,824,849]" italics="true" pageId="4" pageNumber="41">asper</emphasis>
(
<bibRefCitation id="EFA1BA97FFBFE6648CF5FCC070436C74" author="Berlese" box="[681,848,824,850]" pageId="4" pageNumber="51" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFBFE6648D39FCC0765B6C74" authority="Bregetova &amp; Koroleva, 1960" authorityName="Bregetova &amp; Koroleva" authorityYear="1960" box="[869,1352,824,850]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="B9441B74FFBFE6648D39FCC070646C77" box="[869,887,824,849]" italics="true" pageId="4" pageNumber="41">G</emphasis>
.
<emphasis id="B9441B74FFBFE6648DD9FCC070FB6C77" box="[901,1000,824,849]" italics="true" pageId="4" pageNumber="41">ponticus</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648DACFCC0765B6C74" author="Bregetova" box="[1008,1352,824,850]" pageId="4" pageNumber="51" refString="Bregetova, N. G. &amp; Koroleva, E. V. (1960) The macrochelid mites (Gamasoidea, Macrochelidae) in the USSR. Akademiia Nauk SSSR Zoologicheskii institut Parazitologicheskii Sbornik, 29, 32 - 154. [in Russian]." type="journal article" year="1960">Bregetova &amp; Koroleva, 1960</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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