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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.617.9970" ID-GBIF-Dataset="009e9922-b41f-4d4c-beac-e07a023454b4" ID-PMC="PMC5027771" ID-Pensoft-Pub="1313-2970-617-65" ID-PubMed="27667955" ID-ZBK="A22B0E2CF24E4C80B0E505794FC95DA3" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-617-65" ModsDocOrigin="ZooKeys 617" ModsDocTitle="Taxonomic revision of the New World genus Callotillus Wolcott (Cleridae, Tillinae), with the description of the new genus Neocallotillus, and an illustrated key of identification to species" checkinTime="1474013904775" checkinUser="pensoft" docAuthor="Burke, Alan F. &amp; Zolnerowich, Gregory" docDate="2016" docId="B9195D3A476292C5C251D8F5DB830374" docLanguage="en" docName="ZooKeys 617: 65-89" docOrigin="ZooKeys 617" docSource="http://dx.doi.org/10.3897/zookeys.617.9970" docTitle="Neocallotillus Burke, gen. n." docType="treatment" docUuid="08D462B4-C0D4-4BE0-82BC-A604244AAAE6" docUuidSource="ZooBank" docVersion="6" lastPageNumber="68" masterDocId="FFE5FF88FF9EFFE7331FFFB4FFA0FF8D" masterDocTitle="Taxonomic revision of the New World genus Callotillus Wolcott (Cleridae, Tillinae), with the description of the new genus Neocallotillus, and an illustrated key of identification to species" masterLastPageNumber="89" masterPageNumber="65" pageNumber="67" updateTime="1668163519545" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the New World genus Callotillus Wolcott (Cleridae, Tillinae), with the description of the new genus Neocallotillus, and an illustrated key of identification to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burke, Alan F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zolnerowich, Gregory</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>617</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>89</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.617.9970</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.617.9970</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-617-65</mods:identifier>
<mods:identifier type="ZBK">A22B0E2CF24E4C80B0E505794FC95DA3</mods:identifier>
<mods:identifier type="ZooBank">A22B0E2CF24E4C80B0E505794FC95DA3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127864979" LSID="urn:lsid:zoobank.org:act:08D462B4-C0D4-4BE0-82BC-A604244AAAE6" httpUri="http://treatment.plazi.org/id/B9195D3A476292C5C251D8F5DB830374" lastPageId="4" lastPageNumber="68" pageId="2" pageNumber="67">
<subSubSection pageId="2" pageNumber="67" type="multiple">
<paragraph pageId="2" pageNumber="67">Taxon classification Animalia Coleoptera Cleridae</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="67" type="nomenclature">
<paragraph pageId="2" pageNumber="67">
<taxonomicName LSID="http://zoobank.org/08D462B4-C0D4-4BE0-82BC-A604244AAAE6" authority="Burke" class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus Burke</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="67">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="67" type="type species">
<paragraph pageId="2" pageNumber="67">Type species.</paragraph>
<paragraph pageId="2" pageNumber="67">
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus elegans" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="elegans">Neocallotillus elegans</taxonomicName>
(Erichson, 1847), here designated.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="67" type="differential diagnosis">
<paragraph pageId="2" pageNumber="67">Differential diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="67">
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
is most closely related to
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
. The new genus can be differentiated from
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
based on the following combination of characters: in
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
species the male has antennomeres 1-2 filiform; the third antennomere is moderately serrate; antennomeres 4-9 are strongly pectinate; and the tenth antennomere is ovoid in shape and laterally depressed (Fig. 3
<normalizedToken originalValue="A–C">A-C</normalizedToken>
); the length of the tenth antennomere may vary by species. Females have antennomeres 1-3 filiform; the fourth antennomere is moderately serrate; and antennomeres 4-9 are robust, moderately, and gradually increase is size toward the distal end (Fig. 3
<normalizedToken originalValue="D–F">D-F</normalizedToken>
); the tenth antennomere of females is similar to that of the males. Males of
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
have antennomeres 1-2 filiform; the third antennomere is moderately serrate; antennomeres 4-9 are strongly serrate and approximately equal in length; and the tenth antennomere is broadly ovoid and about the same length as antennomeres 8-9 combined (Fig. 3G, H). The antennal structure of females is similar to that of males, except antennomeres 4-9 are moderately serrate and the tenth antennomere is cylindrical to moderately ovoid (Fig. 3I, J). Additionally,
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
species are relatively slender and elongate (Figs 1
<normalizedToken originalValue="A–E">A-E</normalizedToken>
, 4A), while
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
species are conspicuously more robust (Figs 1
<normalizedToken originalValue="F–G">F-G</normalizedToken>
, 4B).
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
species lack an elytral swelling present on the anterior third of the elytral disc of
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
(Fig. 4
<normalizedToken originalValue="A–B">A-B</normalizedToken>
).
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
also resembles
<taxonomicName class="Insecta" family="Cleridae" genus="Barrotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barrotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Barrotillus</taxonomicName>
Rifkind (Fig. 1I), however, the antenna of
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
is composed of 10 antennomeres (Fig. 3
<normalizedToken originalValue="A–F">A-F</normalizedToken>
), while the antenna of
<taxonomicName class="Insecta" family="Cleridae" genus="Barrotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barrotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Barrotillus</taxonomicName>
has 11 antennomeres (Fig. 3K). The restricted distribution of
<taxonomicName class="Insecta" family="Cleridae" genus="Barrotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barrotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Barrotillus</taxonomicName>
, recorded only from a confined locality in Honduras, will also serve to separate it from the widely distributed
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="68" pageId="2" pageNumber="67" type="description">
<paragraph pageId="2" pageNumber="67">Description.</paragraph>
<paragraph pageId="2" pageNumber="67">
Size: 3-7 mm. Color: Light testaceous to dark brown (Fig. 1
<normalizedToken originalValue="A–F">A-F</normalizedToken>
); costae on elytral disc variously adorned, ranging from light testaceous to brown. Form: small to medium sized individuals; body elongate; elytra subparallel to moderately expanded posteriorly.
</paragraph>
<paragraph pageId="3" pageNumber="68">
<pageBreakToken pageId="3" pageNumber="68" start="start">Head</pageBreakToken>
: Eyes medium sized, moderately taller than wide, conspicuously bulging laterally, strongly emarginate at antennal insertion; diameter of ommatidia small (Fig. 4A); clypeus approximately 3
<normalizedToken originalValue="×">x</normalizedToken>
the width of eye emargination and moderately emarginate medially. Antennae composed of 10 antennomeres; sexual dimorphism observable in antennal shape, where the antennae are moderately pectinate and strongly depressed dorsoventrally in males (Fig. 3
<normalizedToken originalValue="A–C">A-C</normalizedToken>
), but serrate and somewhat depressed dorsoventrally in females (Fig. 3
<normalizedToken originalValue="D–F">D-F</normalizedToken>
); tenth antennomere ovoid in both sexes. Labrum elongate, subquadrate; terminal maxillary palp conical, acuminate posteriorly; terminal labial palp securiform.
</paragraph>
<paragraph pageId="3" pageNumber="68">Thorax: Shape of pronotum scutiform, rounded laterally, narrower than anterior margin of elytra; disc feebly to moderately convex; inconspicuously broader at middle, feebly sinuate, conspicuously constricted on last fourth; anterior depression and antescutellar impression absent. Tibial spur formula 2-2-2, pulvillar formula 4-4-4. Prosternum: Smooth to feebly puncticulate; conspicuously wider than long. Mesoventrite: smooth, puncticulate. Metepisternum visible throughout its length in lateral view, not concealed by elytron. Metaventrite: moderately to strongly convex; variously punctate.</paragraph>
<paragraph pageId="3" pageNumber="68">Elytra: Slender to feebly expanded posteriorly, elongate; median region of elytral disc feebly depressed in lateral view; sides subparallel to moderately expanded posteriorly in dorsal view; elytral declivity feebly to moderately gradual; elytral markings always present in various shapes, may be protruding or not.</paragraph>
<paragraph pageId="3" pageNumber="68">Legs: Femora smooth, variably vested. Tibiae feebly to moderately rugulose, weakly expanded posteriorly, variously vested. Two tarsal denticles conspicuously separated from each other, inner tarsal denticles trigonal, outer tarsal denticles digitiform.</paragraph>
<paragraph pageId="3" pageNumber="68">
Abdomen: Smooth to glossy, moderately vested, feebly to moderately convex, with six visible ventrites; lateral margins of ventrites 1-5 parallel, posterior margins truncate; sixth ventrite triangular to subquadrate in shape; male pygidia moderately differentiated from female pygidia (Fig. 5
<normalizedToken originalValue="A–F">A-F</normalizedToken>
).
</paragraph>
<caption pageId="3" pageNumber="68">
<paragraph pageId="3" pageNumber="68">
Figure 5. Pygidium of: A
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Neocallotillus elegans</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="elegans" subspecies="elegans">Callotillus elegans elegans</taxonomicName>
) male B
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Neocallotillus elegans</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="elegans" subspecies="vafer">Callotillus elegans vafer</taxonomicName>
) male C
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Neocallotillus elegans</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="elegans" subspecies="elegans">Callotillus elegans elegans</taxonomicName>
) female D
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Neocallotillus elegans</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="subspecies" species="elegans" subspecies="vafer">Callotillus elegans vafer</taxonomicName>
) female E
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus intricatus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="intricatus">Neocallotillus intricatus</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus intricatus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="intricatus">Callotillus intricatus</taxonomicName>
) male F
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus intricatus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="intricatus">Neocallotillus intricatus</taxonomicName>
(
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus intricatus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="intricatus">Callotillus intricatus</taxonomicName>
) female G
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus bahamensis" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="bahamensis">Callotillus bahamensis</taxonomicName>
male H
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus bahamensis" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="bahamensis">Callotillus bahamensis</taxonomicName>
female I
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus eburneocinctus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="eburneocinctus">Callotillus eburneocinctus</taxonomicName>
male J
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus eburneocinctus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="eburneocinctus">Callotillus eburneocinctus</taxonomicName>
female.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="68">Aedeagus: Moderately robust; phallobasic apodeme short, slender distally; endophallic struts elongate, slender throughout their length.</paragraph>
<paragraph pageId="3" pageNumber="68">
Etymology: This generic name, which is preceded by the Latin prefix neo (new), refers to the superficial similarity to the genus
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="69" pageId="3" pageNumber="68" type="remarks">
<paragraph pageId="3" pageNumber="68">Remarks.</paragraph>
<paragraph lastPageId="4" lastPageNumber="69" pageId="3" pageNumber="68">
Expressing tentative assignment of some of his species to
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
,
<bibRefCitation author="Wolcott, AB" journalOrPublisher="American Museum Novitates" pageId="22" pageNumber="87" pagination="1 - 3" title="Two new species of West Indian Cleridae (Coleoptera)." volume="59" year="1923">Wolcott (1923)</bibRefCitation>
wrote: &quot;
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus crusoe" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="crusoe">Callotillus crusoe</taxonomicName>
, as well as
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Callotillus elegans</taxonomicName>
and
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus vafer" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="vafer">Callotillus vafer</taxonomicName>
, are placed in
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
provisionally only, as it differs from the other members of the genus in several important characters. No doubt, eventually, the creation of a new genus will be necessary for the reception of this new species and
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Callotillus elegans</taxonomicName>
and
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus vafer" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="vafer">Callotillus vafer</taxonomicName>
. In
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus eburneocinctus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="eburneocinctus">Callotillus eburneocinctus</taxonomicName>
, the terminal segment of the maxillary palps is sub-cylindrical, the eyes are emarginate internally and the abdomen has but five segments. In
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus elegans" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="elegans">Callotillus elegans</taxonomicName>
,
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus vafer" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="vafer">Callotillus vafer</taxonomicName>
, and
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus crusoe" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="crusoe">Callotillus crusoe</taxonomicName>
the maxillary palpi have the terminal segment conical, the eyes are deeply emarginate anteriorly, and the abdomen has six distinct well developed segments&quot;. The morphological differences listed by Wolcott, together with the presence of pectinate antennae on males of
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
versus serrate antennae on males of
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
(Fig. 3
<normalizedToken originalValue="A–C">A-C</normalizedToken>
,
<normalizedToken originalValue="G–H">G-H</normalizedToken>
), and an elytral swelling present in
<taxonomicName class="Insecta" family="Cleridae" genus="Callotillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Callotillus</taxonomicName>
but absent in
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
(Fig. 4
<normalizedToken originalValue="A–B">A-B</normalizedToken>
), sup
<pageBreakToken pageId="4" pageNumber="69" start="start">port</pageBreakToken>
the recognition of two genera within the group. The monotypic
<taxonomicName class="Insecta" family="Cleridae" genus="Barrotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barrotillus" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="genus">Barrotillus</taxonomicName>
was also examined in this study in order to assess possible congenericity with
<taxonomicName class="Insecta" family="Cleridae" genus="Neocallotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocallotillus" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="genus">Neocallotillus</taxonomicName>
. The structure of the antennae and number of antennomeres serve as evidence to conclude that these closely related genera should be considered as separate taxa (Fig. 3
<normalizedToken originalValue="A–K">A-K</normalizedToken>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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