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<document id="7E90E602CD06C5D8D878049B6925C51B" ID-CLB-Dataset="21114" ID-DOI="10.11646/zootaxa.4665.1.1" ID-GBIF-Dataset="8053cc5c-900d-490b-84e9-1ca29a46aa3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4000553" ID-ZooBank="78F3F422-DE06-4C61-AC47-69AA684EB5AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598382139973" checkinUser="felipe" docAuthor="Vilkamaa, Pekka &amp; Menzel, Frank" docDate="2019" docId="A703878CA953FFA6FF26FC4198100B96" docLanguage="en" docName="zootaxa.4665.1.1.pdf.imf" docOrigin="Zootaxa 4665 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trichocoelina ithyspina Vilkamaa &amp; Menzel 2019, sp. n." docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="5B3AFFF4A94FFFBAFFB1FF949A6C0C01" masterDocTitle="Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="29" updateTime="1698845439498" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="53980935FEE9BC663490EBAC1ED8BE67">Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species</mods:title>
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<mods:namePart id="574119F4C6C63059F509C0BC2C47C5EC">Vilkamaa, Pekka</mods:namePart>
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<mods:namePart id="5753642A8F3BAE2C121BC5306E6F220F">Menzel, Frank</mods:namePart>
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<mods:title id="8E95B92BC6B6721F42FEF7EAB3A473D0">Zootaxa</mods:title>
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<treatment id="A703878CA953FFA6FF26FC4198100B96" ID-DOI="http://doi.org/10.5281/zenodo.4340430" ID-GBIF-Taxon="167366628" ID-Zenodo-Dep="4340430" LSID="urn:lsid:plazi:treatment:A703878CA953FFA6FF26FC4198100B96" httpUri="http://treatment.plazi.org/id/A703878CA953FFA6FF26FC4198100B96" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="67B06511A953FFA6FF26FC419B930FEE" box="[151,511,981,1008]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="2F15369AA953FFA6FF26FC419B930FEE" blockId="28.[151,511,981,1043]" box="[151,511,981,1008]" pageId="28" pageNumber="29">
<heading id="745D81F6A953FFA6FF26FC419B930FEE" bold="true" box="[151,511,981,1008]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="1DDEEA88A953FFA6FF26FC419B930FEE" bold="true" box="[151,511,981,1008]" pageId="28" pageNumber="29">
<taxonomicName id="E8AA4D19A953FFA6FF26FC419BDA0FF1" ID-CoL="8FHFF" authority="Vilkamaa &amp; Menzel, 2019" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" box="[151,438,981,1008]" class="Insecta" family="Sciaridae" genus="Trichocoelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ithyspina" status="sp. nov.">
<emphasis id="1DDEEA88A953FFA6FF26FC419BDA0FF1" bold="true" box="[151,438,981,1008]" italics="true" pageId="28" pageNumber="29">Trichocoelina ithyspina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED57F3A953FFA6FE0CFC419B930FEE" box="[445,511,981,1007]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="67B06511A953FFA6FF26FC6D9B1C0812" box="[151,368,1017,1043]" pageId="28" pageNumber="29" type="description">
<paragraph id="2F15369AA953FFA6FF26FC6D9B1C0812" blockId="28.[151,511,981,1043]" box="[151,368,1017,1043]" pageId="28" pageNumber="29">
<figureCitation id="B7912A1FA953FFA6FF26FC6D9A950812" box="[151,249,1017,1043]" captionStart="FIGURE 8" captionStartId="30.[151,250,1980,2005]" captionTargetBox="[333,1292,212,1929]" captionTargetId="figure@30.[280,1308,166,1960]" captionTargetPageId="30" captionText="FIGURE 8. Hypopygium, ventral. A. Trichocoelina ithyspina sp. n. (holotype). B. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000569" httpUri="https://zenodo.org/record/4000569/files/figure.png" pageId="28" pageNumber="29">Figs 8 A</figureCitation>
,
<figureCitation id="B7912A1FA953FFA6FEB5FC6D9B420812" box="[260,302,1017,1043]" captionStart="FIGURE 9" captionStartId="31.[151,250,1974,1999]" captionTargetBox="[174,1431,220,1882]" captionTargetId="figure@31.[151,1436,166,1954]" captionTargetPageId="31" captionText="FIGURE 9. Gonostylus, ventral. A. Trichocoelina incrassata sp. n. (holotype). B. T. ithyspina sp. n. (holotype). C. T. magnifica sp. n. (holotype). D. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000571" httpUri="https://zenodo.org/record/4000571/files/figure.png" pageId="28" pageNumber="29">9 B</figureCitation>
,
<figureCitation id="B7912A1FA953FFA6FE8BFC6D9B1C0812" box="[314,368,1017,1043]" captionStart="FIGURE 18" captionStartId="48.[151,250,1741,1766]" captionTargetBox="[160,1376,185,1720]" captionTargetId="figure@48.[151,1436,166,1722]" captionTargetPageId="48" captionText="FIGURE 18. Intergonocoxal lobes of hypopygium, ventral. A. Trichocoelina aemula (holotype). B. T. dispansa sp. n. (paratype). C. T. ithyspina sp. n. (holotype). D. T. jukkai sp. n. (paratype). E. T. planilobata sp. n. (holotype). F. T. semusta sp. n. (holotype). Scale 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4000591" httpUri="https://zenodo.org/record/4000591/files/figure.png" pageId="28" pageNumber="29">18 C</figureCitation>
</paragraph>
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<subSubSection id="67B06511A953FFA6FF26FBD49AB908A2" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="2F15369AA953FFA6FF26FBD49AB908A2" blockId="28.[151,1437,1088,1943]" pageId="28" pageNumber="29">
<emphasis id="1DDEEA88A953FFA6FF26FBD49B33085A" bold="true" box="[151,351,1088,1115]" pageId="28" pageNumber="29">Material studied</emphasis>
.
<materialsCitation id="9FC23CC7A953FFA6FEDEFBD59AB908A2" ID-GBIF-Occurrence="2849081331" collectingDate="2008-06-30" collectingDateMax="2008-07-07" collectingDateMin="2008-06-30" collectorName="E. Stur &amp; Sample ID" country="Norway" elevation="380" latitude="61.746" location="river Atna Solbakken" longLatPrecision="61" longitude="10.746" municipality="Stor-Elvdal" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Hedmark" typeStatus="holotype">
<emphasis id="1DDEEA88A953FFA6FEDEFBD59878085A" box="[367,532,1089,1115]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="F0118838A953FFA6FEDEFBD59BBA085A" box="[367,470,1089,1115]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<collectingCountry id="57BD760AA953FFA6FD95FBD598F6085A" box="[548,666,1089,1115]" name="Norway" pageId="28" pageNumber="29">NORWAY</collectingCountry>
,
<collectingRegion id="ED6EF878A953FFA6FD16FBD5997E085A" box="[679,786,1089,1115]" country="Norway" name="Hedmark" pageId="28" pageNumber="29">Hedmark</collectingRegion>
,
<collectingMunicipality id="CF71ACE0A953FFA6FCAEFBD599CE085A" box="[799,930,1089,1115]" pageId="28" pageNumber="29">Stor-Elvdal</collectingMunicipality>
,
<location id="2A756041A953FFA6FC1EFBD59ECF085A" LSID="urn:lsid:plazi:treatment:A703878CA953FFA6FF26FC4198100B96:2A756041A953FFA6FC1EFBD59ECF085A" box="[943,1187,1089,1115]" country="Norway" latitude="61.746" longLatPrecision="61" longitude="10.746" municipality="Stor-Elvdal" name="river Atna Solbakken" pageId="28" pageNumber="29" stateProvince="Hedmark">river Atna Solbakken</location>
,
<geoCoordinate id="4A9E505DA953FFA6FB01FBD59F73085A" box="[1200,1311,1089,1115]" degrees="61.746" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="55" value="61.746">61.746°N</geoCoordinate>
,
<geoCoordinate id="4A9E505DA953FFA6FA9DFBD59FF4085A" box="[1324,1432,1089,1115]" degrees="10.746" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="55" value="10.746">10.746°E</geoCoordinate>
,
<quantity id="E8529B7FA953FFA6FF26FBF19A8D087E" box="[151,225,1125,1151]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="28" pageNumber="29" unit="m" value="380.0">
<elevation id="A487D1A9A953FFA6FF26FBF19A8D087E" box="[151,225,1125,1151]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="28" pageNumber="29" unit="m" value="380.0">380 m</elevation>
</quantity>
,
<collectingMethod id="F6EB4E8DA953FFA6FF5FFBF19B11087E" box="[238,381,1125,1151]" pageId="28" pageNumber="29">Malaise trap</collectingMethod>
,
<date id="5B14105AA953FFA6FE3BFBF1983A087E" box="[394,598,1125,1151]" pageId="28" pageNumber="29" value="2008-06-30" valueMax="2008-07-07" valueMin="2008-06-30">
<collectingDate id="4B50E9B2A953FFA6FE3BFBF1983A087E" box="[394,598,1125,1151]" pageId="28" pageNumber="29" value="2008-06-30" valueMax="2008-07-07" valueMin="2008-06-30">30.VI7.VII.2008</collectingDate>
</date>
,
<collectorName id="825F534CA953FFA6FDD2FBF198DE087E" box="[611,690,1125,1151]" pageId="28" pageNumber="29">E. Stur</collectorName>
(NTNU-VM 201775, BOLD
<collectorName id="825F534CA953FFA6FBB6FBF19EEE087E" box="[1031,1154,1125,1151]" pageId="28" pageNumber="29">Sample ID</collectorName>
TRD-Sci029, in NTNU- VM).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="67B06511A953FFA6FF76FB389BBA0B06" pageId="28" pageNumber="29" type="description">
<paragraph id="2F15369AA953FFA6FF76FB389EF10AE2" blockId="28.[151,1437,1088,1943]" pageId="28" pageNumber="29">
<emphasis id="1DDEEA88A953FFA6FF76FB389B3D08C6" bold="true" box="[199,337,1196,1223]" pageId="28" pageNumber="29">Description</emphasis>
. Male.
<emphasis id="1DDEEA88A953FFA6FE1BFB399B8608C6" bold="true" box="[426,490,1197,1223]" pageId="28" pageNumber="29">Head</emphasis>
. Face and antenna concolorous brown but pedicellus yellow, maxillary palpus pale yellowish. Eye bridge, face and clypeus poorly visible in the specimen studied. Maxillary palpus with 3 segments, 1
<superScript id="D8DF9BD2A953FFA6FF14FB609ADC0903" attach="left" box="[165,176,1268,1282]" fontSize="6" pageId="28" pageNumber="29">st</superScript>
segment longer than 3
<superScript id="D8DF9BD2A953FFA6FE0BFB609BA40903" attach="left" box="[442,456,1268,1282]" fontSize="6" pageId="28" pageNumber="29">rd</superScript>
segment, 2
<superScript id="D8DF9BD2A953FFA6FDFCFB6098310903" attach="left" box="[589,605,1268,1282]" fontSize="6" pageId="28" pageNumber="29">nd</superScript>
segment shortest; 1
<superScript id="D8DF9BD2A953FFA6FCF4FB60993C0903" attach="left" box="[837,848,1268,1282]" fontSize="6" pageId="28" pageNumber="29">st</superScript>
segment with 1 seta, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4
<superScript id="D8DF9BD2A953FFA6FD52FA8C989C0927" attach="left" box="[739,752,1304,1318]" fontSize="6" pageId="28" pageNumber="29">th</superScript>
antennal flagellomere 2.9x as long as wide, the neck shorter than broad, the longest setae longer than the width of flagellomere.
<emphasis id="1DDEEA88A953FFA6FC3FFAA9998B0956" bold="true" box="[910,999,1341,1367]" pageId="28" pageNumber="29">Thorax</emphasis>
. Brown, setae pale. Anterior pronotum with 9 setae. Proepisternum with 11 setae. Scutellum with 4 longer and some short and fine setae.
<emphasis id="1DDEEA88A953FFA6FB5CFAF49F42097A" bold="true" box="[1261,1326,1376,1403]" pageId="28" pageNumber="29">Wing</emphasis>
. Fumose. Length
<quantity id="E8529B7FA953FFA6FF5CFA119B29099E" box="[237,325,1413,1439]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="29" unit="mm" value="2.3">2.3 mm</quantity>
. Width/length 0.45. Anal lobe weak. Veins distinct. c/w 0.65. R
<subScript id="B32E34DFA953FFA6FBB4FA029E6109A5" attach="left" box="[1029,1037,1430,1444]" fontSize="6" pageId="28" pageNumber="29">1</subScript>
/R 0.80. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 1 seta. Halter yellow.
<emphasis id="1DDEEA88A953FFA6FC36FA3D99D309C2" bold="true" box="[903,959,1449,1475]" pageId="28" pageNumber="29">Legs</emphasis>
. Yellow, coxal setae pale. Fore tibial organ forming a large indistinct patch in depression. Fore tibial spur longer than the tibial width.
<emphasis id="1DDEEA88A953FFA6FB22FA589F6409E6" bold="true" box="[1171,1288,1484,1511]" pageId="28" pageNumber="29">Abdomen</emphasis>
. Pale brown, setae pale, moderately long and fine.
<emphasis id="1DDEEA88A953FFA6FDF6FA64988F0A0A" bold="true" box="[583,739,1520,1547]" pageId="28" pageNumber="29">Hypopygium</emphasis>
(
<figureCitation id="B7912A1FA953FFA6FD45FA6599340A0A" box="[756,856,1521,1547]" captionStart="FIGURE 8" captionStartId="30.[151,250,1980,2005]" captionTargetBox="[333,1292,212,1929]" captionTargetId="figure@30.[280,1308,166,1960]" captionTargetPageId="30" captionText="FIGURE 8. Hypopygium, ventral. A. Trichocoelina ithyspina sp. n. (holotype). B. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000569" httpUri="https://zenodo.org/record/4000569/files/figure.png" pageId="28" pageNumber="29">Fig. 8 A</figureCitation>
). Brown, as abdomen. Intergonocoxal area moderately long, with long, apically bifid setose lobe (
<figureCitation id="B7912A1FA953FFA6FD0FF98199420A2E" box="[702,814,1557,1583]" captionStart="FIGURE 18" captionStartId="48.[151,250,1741,1766]" captionTargetBox="[160,1376,185,1720]" captionTargetId="figure@48.[151,1436,166,1722]" captionTargetPageId="48" captionText="FIGURE 18. Intergonocoxal lobes of hypopygium, ventral. A. Trichocoelina aemula (holotype). B. T. dispansa sp. n. (paratype). C. T. ithyspina sp. n. (holotype). D. T. jukkai sp. n. (paratype). E. T. planilobata sp. n. (holotype). F. T. semusta sp. n. (holotype). Scale 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4000591" httpUri="https://zenodo.org/record/4000591/files/figure.png" pageId="28" pageNumber="29">Fig. 18 C</figureCitation>
). Gonocoxa moderately broad, longer than gonostylus, medial margin basally smoothly curved, with rather long but sparse setosity. Gonostylus (
<figureCitation id="B7912A1FA953FFA6FB23F9AD9E9D0A52" box="[1170,1265,1593,1619]" captionStart="FIGURE 9" captionStartId="31.[151,250,1974,1999]" captionTargetBox="[174,1431,220,1882]" captionTargetId="figure@31.[151,1436,166,1954]" captionTargetPageId="31" captionText="FIGURE 9. Gonostylus, ventral. A. Trichocoelina incrassata sp. n. (holotype). B. T. ithyspina sp. n. (holotype). C. T. magnifica sp. n. (holotype). D. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000571" httpUri="https://zenodo.org/record/4000571/files/figure.png" pageId="28" pageNumber="29">Fig. 9 B</figureCitation>
) long, apically truncate, strongly impressed; with normal setosity, with a rather long apical tooth, with three megasetae subapically near the tooth and two more basally, megasetae long and slender, with distinct basal bodies; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen broader than long, laterally roundish, apically straight, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long.
</paragraph>
<paragraph id="2F15369AA953FFA6FF76F9799BBA0B06" blockId="28.[151,1437,1088,1943]" box="[199,470,1772,1799]" pageId="28" pageNumber="29">
<emphasis id="1DDEEA88A953FFA6FF76F9799A950B06" bold="true" box="[199,249,1773,1799]" pageId="28" pageNumber="29">BIN</emphasis>
. BOLD:ACX5966.
</paragraph>
</subSubSection>
<subSubSection id="67B06511A953FFA6FF76F8849EE60B4E" pageId="28" pageNumber="29" type="discussion">
<paragraph id="2F15369AA953FFA6FF76F8849EE60B4E" blockId="28.[151,1437,1088,1943]" pageId="28" pageNumber="29">
<emphasis id="1DDEEA88A953FFA6FF76F8849B290B2A" bold="true" box="[199,325,1808,1835]" pageId="28" pageNumber="29">Discussion</emphasis>
. The gonostylus shown in
<figureCitation id="B7912A1FA953FFA6FD31F885988F0B2A" box="[640,739,1809,1835]" captionStart="FIGURE 8" captionStartId="30.[151,250,1980,2005]" captionTargetBox="[333,1292,212,1929]" captionTargetId="figure@30.[280,1308,166,1960]" captionTargetPageId="30" captionText="FIGURE 8. Hypopygium, ventral. A. Trichocoelina ithyspina sp. n. (holotype). B. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000569" httpUri="https://zenodo.org/record/4000569/files/figure.png" pageId="28" pageNumber="29">Figs 8 A</figureCitation>
and
<figureCitation id="B7912A1FA953FFA6FCAFF88599250B2A" box="[798,841,1809,1835]" captionStart="FIGURE 9" captionStartId="31.[151,250,1974,1999]" captionTargetBox="[174,1431,220,1882]" captionTargetId="figure@31.[151,1436,166,1954]" captionTargetPageId="31" captionText="FIGURE 9. Gonostylus, ventral. A. Trichocoelina incrassata sp. n. (holotype). B. T. ithyspina sp. n. (holotype). C. T. magnifica sp. n. (holotype). D. T. jukkai sp. n. (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4000571" httpUri="https://zenodo.org/record/4000571/files/figure.png" pageId="28" pageNumber="29">9 B</figureCitation>
was slightly distorted in preparation.
<taxonomicName id="E8AA4D19A953FFA6FAB0F8859A920B4E" authority="Vilkamaa &amp; Menzel, 2019" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" class="Insecta" family="Sciaridae" genus="Trichocoelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ithyspina" status="sp. nov.">
<emphasis id="1DDEEA88A953FFA6FAB0F8859A920B4E" italics="true" pageId="28" pageNumber="29">Trichocoelina ithyspina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED57F3A953FFA6FEB4F8A19B2D0B4E" box="[261,321,1845,1871]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
resembles most
<taxonomicName id="E8AA4D19A953FFA6FE4DF8A298300B4E" authority="Vilkamaa &amp; Menzel, 2019" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" box="[508,604,1845,1871]" class="Insecta" family="Sciaridae" genus="Trichocoelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="jukkai" status="sp. nov.">
<emphasis id="1DDEEA88A953FFA6FE4DF8A298300B4E" box="[508,604,1845,1871]" italics="true" pageId="28" pageNumber="29">T. jukkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED57F3A953FFA6FDD2F8A198F30B4E" box="[611,671,1845,1871]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
For discussion, see under
<taxonomicName id="E8AA4D19A953FFA6FC79F8A29E240B4E" authority="Vilkamaa &amp; Menzel, 2019" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" box="[968,1096,1845,1871]" class="Insecta" family="Sciaridae" genus="Trichocoelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dispansa" status="sp. nov.">
<emphasis id="1DDEEA88A953FFA6FC79F8A29E240B4E" box="[968,1096,1845,1871]" italics="true" pageId="28" pageNumber="29">T. dispansa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06ED57F3A953FFA6FBFFF8A19EE60B4E" box="[1102,1162,1845,1871]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="67B06511A953FFA6FF76F8CD98100B96" pageId="28" pageNumber="29" type="etymology">
<paragraph id="2F15369AA953FFA6FF76F8CD98100B96" blockId="28.[151,1437,1088,1943]" pageId="28" pageNumber="29">
<emphasis id="1DDEEA88A953FFA6FF76F8CD9B240B72" bold="true" box="[199,328,1881,1907]" pageId="28" pageNumber="29">Etymology</emphasis>
. The name is derived from the Greek word
<emphasis id="1DDEEA88A953FFA6FC9DF8CD99040B72" box="[812,872,1881,1907]" italics="true" pageId="28" pageNumber="29">ithys,</emphasis>
straight, and the Latin word
<emphasis id="1DDEEA88A953FFA6FB19F8CE9E870B72" box="[1192,1259,1882,1907]" italics="true" pageId="28" pageNumber="29">spina,</emphasis>
spine, referring to the straight megasetae of the gonostylus.
</paragraph>
</subSubSection>
</treatment>
</document>