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<document id="B3CBEB4420B914C67E46A97792B548C0" ID-DOI="10.5281/zenodo.193344" ID-GBIF-Dataset="83079833-f316-4407-bcc3-79bea9db1097" ID-ISSN="1175-5326" ID-Zenodo-Dep="193344" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460135803912" checkinUser="plazi" docAuthor="Chen, Xiang-Sheng &amp; Yang, Lin" docDate="2010" docId="947C87F5CE09E04710D668B9314B0B76" docLanguage="en" docName="zt02344p038.pdf" docOrigin="Zootaxa 2344" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kakuna zhongtuana Chen &amp; Yang, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="6845FF8DCE0FE04F1041682E32510951" masterDocTitle="Oriental bamboo delphacid planthoppers: three new species of genus Kakuna Matsumura (Hemiptera: Fulgoromorpha: Delphacidae) from Guizhou Province, China" masterLastPageNumber="38" masterPageNumber="29" pageNumber="35" updateTime="1698235212942" updateUser="plazi">
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<mods:title id="2D664AD9F08BB0992917F74CF68F23F5">Oriental bamboo delphacid planthoppers: three new species of genus Kakuna Matsumura (Hemiptera: Fulgoromorpha: Delphacidae) from Guizhou Province, China</mods:title>
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<mods:name id="E96F1F9937AE642419CCC223D2BA2F73" type="personal">
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<mods:namePart id="DB120A16629011E42DB05E95C33688A7">Chen, Xiang-Sheng</mods:namePart>
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<mods:namePart id="582C0C2A1D48A8157E05927286E0E50F">Yang, Lin</mods:namePart>
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<mods:date id="7FF5D0A14DBA93586F55F6B2F8B278BD">2010</mods:date>
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<treatment id="947C87F5CE09E04710D668B9314B0B76" ID-DOI="http://doi.org/10.5281/zenodo.6197648" ID-GBIF-Taxon="119383054" ID-Zenodo-Dep="6197648" LSID="urn:lsid:plazi:treatment:947C87F5CE09E04710D668B9314B0B76" httpUri="http://treatment.plazi.org/id/947C87F5CE09E04710D668B9314B0B76" lastPageId="8" lastPageNumber="37" pageId="6" pageNumber="35">
<subSubSection id="54CF6568CE09E04910D668B9337C0985" pageId="6" pageNumber="35" type="nomenclature">
<paragraph id="1C6A36E3CE09E04910D668B933A309E0" blockId="6.[151,498,151,212]" box="[151,498,151,177]" pageId="6" pageNumber="35">
<heading id="4722818FCE09E04910D668B933A309E0" bold="true" box="[151,498,151,177]" fontSize="11" level="1" pageId="6" pageNumber="35" reason="1">
<emphasis id="2EA1EAF1CE09E04910D668B933A309E0" bold="true" box="[151,498,151,177]" pageId="6" pageNumber="35">
<taxonomicName id="DBD54D60CE09E04910D668B933DC09E0" ID-CoL="3QYF5" box="[151,397,151,177]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="zhongtuana" status="sp. nov.">
<emphasis id="2EA1EAF1CE09E04910D668B933DC09E0" bold="true" box="[151,397,151,177]" italics="true" pageId="6" pageNumber="35">Kakuna zhongtuana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3592578ACE09E04911D468B933A309E0" box="[405,498,151,177]" pageId="6" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1C6A36E3CE09E04910D66894337C0985" blockId="6.[151,498,151,212]" box="[151,301,186,212]" pageId="6" pageNumber="35">
(
<figureCitation id="84EE2A66CE09E04910DE689433740985" box="[159,293,186,212]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="6" pageNumber="35">Figs 2334</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="54CF6568CE09E04810D6692B333D0A03" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35" type="description">
<paragraph id="1C6A36E3CE09E04910D6692B33F70815" blockId="6.[151,1436,261,324]" pageId="6" pageNumber="35">
<emphasis id="2EA1EAF1CE09E04910D6692B3365084E" bold="true" box="[151,308,261,287]" pageId="6" pageNumber="35">Description.</emphasis>
Body length (from apex of vertex to tip of forewings): male 5.556.00 mm (N = 4); female
<quantity id="DB2D9B06CE09E04910D6690433100815" box="[151,321,298,324]" metricMagnitude="-3" metricUnit="m" metricValue="6.425" metricValueMax="6.6" metricValueMin="6.25" pageId="6" pageNumber="35" unit="mm" value="6.425" valueMax="6.6" valueMin="6.25">6.256.60 mm</quantity>
(N = 6).
</paragraph>
<caption id="48AA666BCE09E04910D66EA130500E76" httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="6" pageNumber="35" targetBox="[161,1426,389,1648]" targetPageId="6">
<paragraph id="1C6A36E3CE09E04910D66EA130500E76" blockId="6.[151,1436,1679,1831]" pageId="6" pageNumber="35">
<emphasis id="2EA1EAF1CE09E04910D66EA133390FF6" bold="true" box="[151,360,1679,1703]" pageId="6" pageNumber="35">FIGURES 2334.</emphasis>
<taxonomicName id="DBD54D60CE09E04911316EBE30030FF6" box="[368,594,1680,1703]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="zhongtuana" status="sp. nov.">
<emphasis id="2EA1EAF1CE09E04911316EBE30030FF6" box="[368,594,1680,1703]" italics="true" pageId="6" pageNumber="35">Kakuna zhongtuana</emphasis>
</taxonomicName>
<emphasis id="2EA1EAF1CE09E049121A6EA130980FF6" bold="true" box="[603,713,1679,1703]" pageId="6" pageNumber="35">
<taxonomicNameLabel id="3592578ACE09E049121A6EA130E60FF6" box="[603,695,1679,1703]" pageId="6" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
(
</emphasis>
23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 2324); 2.0 mm (Figs 2527); 0.3 mm (Figs 2930); 0.2 mm (Figs 28, 3134).
</paragraph>
</caption>
<paragraph id="1C6A36E3CE09E04810856F7B30DD09AD" blockId="6.[151,1437,1877,2019]" lastBlockId="7.[151,1437,152,850]" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35">
<emphasis id="2EA1EAF1CE09E04910856F7B330D0E3E" bold="true" box="[196,348,1877,1903]" pageId="6" pageNumber="35">Coloration.</emphasis>
General color yellowish brown to brown. Ocelli reddish brown, eyes blackish brown. Rostrum with apex fuscous. Antennae yellowish brown, except apex of basal segment and base of second segment, fuscous. Forewing (
<figureCitation id="84EE2A66CE09E04911AF6F8E30120EEB" box="[494,579,1952,1978]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="6" pageNumber="35">Fig. 25</figureCitation>
) pale yellowish brown, with a fuscous large, longitudinal band from base of costal area to apex in male, apex of I A with a small blackish brown marking. Thorax with lateral and ventral areas pale brown. Legs yellowish brown, except apex of digitus fuscous. Abdomen brown to fuscous. Color of female paler than male, forewing (
<figureCitation id="84EE2A66CE08E04812696893302C0986" box="[552,637,189,215]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 26</figureCitation>
) pale yellowish brown, apical part with a brown, broad, longitudinal band from front of transverse vein to apex.
</paragraph>
<paragraph id="1C6A36E3CE08E0481085692933B60B53" blockId="7.[151,1437,152,850]" pageId="7" pageNumber="36">
<emphasis id="2EA1EAF1CE08E0481085692933F00870" bold="true" box="[196,417,263,289]" pageId="7" pageNumber="36">Head and thorax.</emphasis>
Vertex including eyes narrower than pronotum (0.79:1). Vertex shorter in middle line than wide at base (0.65:1), broad at apex narrower than at base (0.78:1). Frons longer in middle line than wide at widest part (about 2.30: 1). Antennae with basal segment 1.85 times as long as broad at apex, shorter than second segment (0.56:1). Pronotum as long as vertex in middle line. Mesonotum 1.78 times as long as vertex and pronotum together in middle line. Forewings longer in middle line than broad at widest part (3.37 3.41:1). Post-tibial spur with about 27 teeth along hind margin. Basal segment of digitus longer than second and third combined (1.70:1).
</paragraph>
<paragraph id="1C6A36E3CE08E04810856A23333D0A03" blockId="7.[151,1437,152,850]" pageId="7" pageNumber="36">
<emphasis id="2EA1EAF1CE08E04810856A2333D90B76" bold="true" box="[196,392,525,551]" pageId="7" pageNumber="36">Male genitalia.</emphasis>
Anal segment of male (
<figureCitation id="84EE2A66CE08E04812F56A23315E0B76" box="[692,783,525,551]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 28</figureCitation>
) short, ring-like, almost sunk in the emargination of pygofer, ventral margin without process. Pygofer in profile (
<figureCitation id="84EE2A66CE08E04813246A1C31ED0B1D" box="[869,956,562,588]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 29</figureCitation>
) with dorsal margin shorter than ventral margin, laterodorsal angle strongly produced caudad, posterior margin slightly sinuate; in posterior view (
<figureCitation id="84EE2A66CE08E048152D6A7632E90BC6" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 30</figureCitation>
), with opening longer than wide, ventral margin broadly concave. Suspensorium ring-like. Aedeagus in profile (
<figureCitation id="84EE2A66CE08E04810B96A8C331E0BED" box="[248,335,674,700]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 33</figureCitation>
) tubular, broadly curving medially, apex rounded; in ventral view (
<figureCitation id="84EE2A66CE08E04814396A8C36810BED" box="[1144,1232,674,700]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 32</figureCitation>
) with apical half subtriangular, having two rows teeth along lateral margins. Diaphragm (
<figureCitation id="84EE2A66CE08E04813B06AE636160BB3" box="[1009,1095,712,738]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 31</figureCitation>
) with dorsal margin broadly produced dorsad, median part with two spine-like processes, which having a long common stalk basally. Genital style in profile (
<figureCitation id="84EE2A66CE08E04811EE6B3C30550A7D" box="[431,516,786,812]" captionStart="FIGURES 23 34" captionStartId="6.[151,275,1679,1703]" captionTargetBox="[161,1426,389,1648]" captionTargetId="figure@6.[153,1433,378,1658]" captionTargetPageId="6" captionText="FIGURES 23 34. Kakuna zhongtuana sp. nov. (23) Head and thorax, dorsal view; (24) Frons and clypeus; (25) Forewing (3); (26) Forewing (Ƥ); (27) Hindwing; (28) Male anal segment, lateral view; (29) Male genital segment, lateral view; (30) Male genital segment, posterior view; (31) Diaphragm; (32) Aedeagus, ventral view; (33) Aedeagus, lateral view; (34) Left genital style, lateral view. Scale bars = 1.0 mm (Figs 23 24); 2.0 mm (Figs 25 27); 0.3 mm (Figs 29 30); 0.2 mm (Figs 28, 31 34)." httpUri="https://zenodo.org/record/193347/files/figure.png" pageId="7" pageNumber="36">Fig. 34</figureCitation>
) broad at base, narrowing apically, apex acute, inner margin with a nipple-like process at middle.
</paragraph>
</subSubSection>
<caption id="48AA666BCE08E04810D66F2931E30E6E" httpUri="https://zenodo.org/record/193348/files/figure.png" pageId="7" pageNumber="36" targetBox="[167,1421,893,1769]" targetPageId="7">
<paragraph id="1C6A36E3CE08E04810D66F2931E30E6E" blockId="7.[151,1436,1798,1855]" pageId="7" pageNumber="36">
<emphasis id="2EA1EAF1CE08E04810D66F2933760E4E" bold="true" box="[151,295,1799,1823]" pageId="7" pageNumber="36">FIGURE 35.</emphasis>
Geographic distribution of
<taxonomicName id="DBD54D60CE08E048120E6F2630F30E4E" box="[591,674,1800,1823]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="2EA1EAF1CE08E048120E6F2630F30E4E" box="[591,674,1800,1823]" italics="true" pageId="7" pageNumber="36">Kakuna</emphasis>
</taxonomicName>
species:
<taxonomicName id="DBD54D60CE08E04813456F26364C0E4E" authority="Matsumura" authorityName="Matsumura" box="[772,1053,1800,1823]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="2EA1EAF1CE08E04813456F2631CC0E4E" box="[772,925,1800,1823]" italics="true" pageId="7" pageNumber="36">K. kuwayamai</emphasis>
Matsumura
</taxonomicName>
(♦);
<taxonomicName id="DBD54D60CE08E04814136F2636D90E4E" box="[1106,1160,1800,1823]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lii" status="sp. nov.">
<emphasis id="2EA1EAF1CE08E04814136F2636D90E4E" box="[1106,1160,1800,1823]" italics="true" pageId="7" pageNumber="36">K. lii</emphasis>
</taxonomicName>
<emphasis id="2EA1EAF1CE08E04814CF6F2936B50E4E" bold="true" box="[1166,1252,1799,1823]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="3592578ACE08E04814CF6F2936B50E4E" box="[1166,1252,1799,1823]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(●);
<taxonomicName id="DBD54D60CE08E048155A6F2637CD0E4E" box="[1307,1436,1800,1823]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="nonspina" status="sp. nov.">
<emphasis id="2EA1EAF1CE08E048155A6F2637CD0E4E" box="[1307,1436,1800,1823]" italics="true" pageId="7" pageNumber="36">K. nonspina</emphasis>
</taxonomicName>
<emphasis id="2EA1EAF1CE08E04810D66F0932BC0E6E" bold="true" box="[151,237,1831,1855]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="3592578ACE08E04810D66F0932BC0E6E" box="[151,237,1831,1855]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
();
<taxonomicName id="DBD54D60CE08E04811626F0630110E6E" authority="Yang, 1989" authorityName="Yang" authorityYear="1989" box="[291,576,1832,1855]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="yushaniae">
<emphasis id="2EA1EAF1CE08E04811626F06336D0E6E" box="[291,316,1832,1855]" italics="true" pageId="7" pageNumber="36">K.</emphasis>
<emphasis id="2EA1EAF1CE08E04811036F0633FF0E6E" box="[322,430,1832,1855]" italics="true" pageId="7" pageNumber="36">yushaniae</emphasis>
(Yang, 1989)
</taxonomicName>
(ˑ);
<taxonomicName id="DBD54D60CE08E04812C06F06314D0E6E" box="[641,796,1832,1855]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="zhongtuana" status="sp. nov.">
<emphasis id="2EA1EAF1CE08E04812C06F06314D0E6E" box="[641,796,1832,1855]" italics="true" pageId="7" pageNumber="36">K. zhongtuana</emphasis>
</taxonomicName>
<emphasis id="2EA1EAF1CE08E04813626F0931290E6E" bold="true" box="[803,888,1831,1855]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="3592578ACE08E04813626F0931290E6E" box="[803,888,1831,1855]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(˔).
</paragraph>
</caption>
<subSubSection id="54CF6568CE08E04710856F5E372A0873" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36" type="materials_examined">
<paragraph id="1C6A36E3CE08E04710856F5E329309AD" blockId="7.[151,1436,1901,2011]" lastBlockId="8.[151,1437,152,551]" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
<emphasis id="2EA1EAF1CE08E04810856F5E332B0EDB" bold="true" box="[196,378,1904,1930]" pageId="7" pageNumber="36">
<typeStatus id="C36E8841CE08E04810856F5E33500EDB" box="[196,257,1904,1930]" pageId="7" pageNumber="36">Type</typeStatus>
material.
</emphasis>
<typeStatus id="C36E8841CE08E04811C36F5E33A50EDB" box="[386,500,1904,1930]" pageId="7" pageNumber="36" type="holotype">Holotype</typeStatus>
: 3,
<collectingCountry id="64C27673CE08E04812656F5E303E0EDB" box="[548,623,1904,1930]" name="China" pageId="7" pageNumber="36">China</collectingCountry>
: Guizhou, Leishan, Leigongshan National Natural Reserve (
<geoCoordinate id="79E15024CE08E04815756F5E37C60EDB" box="[1332,1431,1901,1930]" direction="north" orientation="latitude" pageId="7" pageNumber="36" precision="925" value="26.466667">26o28N</geoCoordinate>
,
<geoCoordinate id="79E15024CE08E04810D66FB633580EE3" box="[151,265,1941,1970]" direction="east" orientation="longitude" pageId="7" pageNumber="36" precision="925" value="108.28333">108 o17E</geoCoordinate>
),
<quantity id="DB2D9B06CE08E048115E6FB6332D0EE3" box="[287,380,1944,1970]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="36" unit="m" value="1900.0">1900 m</quantity>
,
<date id="686B1023CE08E04811C96FB6304E0EE3" box="[392,543,1944,1970]" pageId="7" pageNumber="36" value="2004-08-02">2 Aug. 2004</date>
, D.-Y. Ge and P. Xu;
<typeStatus id="C36E8841CE08E048135E6FB631C50EE3" box="[799,916,1944,1970]" pageId="7" pageNumber="36" type="paratype">paratypes</typeStatus>
: 1 3, 5 Ƥ, same data as
<typeStatus id="C36E8841CE08E04814F46FB637710EE3" box="[1205,1312,1944,1970]" pageId="7" pageNumber="36" type="holotype">holotype</typeStatus>
; 1 3, 1 Ƥ, Guizhou, Suiyang, Kuankuoshui National Natural Reserve (
<geoCoordinate id="79E15024CE08E04813C46FEF31BC0E8A" box="[901,1005,1982,2011]" direction="north" orientation="latitude" pageId="7" pageNumber="36" precision="925" value="27.966667">27o58N</geoCoordinate>
,
<geoCoordinate id="79E15024CE08E04813BA6FEF36220E8A" box="[1019,1139,1982,2011]" direction="east" orientation="longitude" pageId="7" pageNumber="36" precision="925" value="107.183334">107 o11E</geoCoordinate>
),
<date id="686B1023CE08E04814CA6FEF377B0E8A" box="[1163,1322,1985,2011]" pageId="7" pageNumber="36" value="1984-08-04">4 Aug. 1984</date>
, Z.-Z Li; Guizhou, Daozhen, Dashahe Provincial Natural Reserve (
<geoCoordinate id="79E15024CE07E047130368B631FA09E3" box="[834,939,152,178]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="925" value="28.883333">28°53´N</geoCoordinate>
,
<geoCoordinate id="79E15024CE07E04713F968B6367D09E3" box="[952,1068,152,178]" direction="east" orientation="longitude" pageId="8" pageNumber="37" precision="925" value="107.6">107°36´E</geoCoordinate>
),
<quantity id="DB2D9B06CE07E047140368B636CE09E3" box="[1090,1183,152,178]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="37" unit="m" value="1400.0">1400 m</quantity>
,
<date id="686B1023CE07E04714ED68B6370009E3" box="[1196,1361,152,178]" pageId="8" pageNumber="37" value="2004-08-17">17 Aug. 2004</date>
, X.-S. Chen; 1 3, Guizhou, Libo, Maolan National Natural Reserve (
<geoCoordinate id="79E15024CE07E047132E689331890986" box="[879,984,189,215]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="925" value="25.666666">25°40´N</geoCoordinate>
,
<geoCoordinate id="79E15024CE07E04713A5689336060986" box="[996,1111,189,215]" direction="east" orientation="longitude" pageId="8" pageNumber="37" precision="925" value="108.083336">108°05´E</geoCoordinate>
),
<quantity id="DB2D9B06CE07E047142D689336E80986" box="[1132,1209,189,215]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="37" unit="m" value="600.0">600 m</quantity>
,
<date id="686B1023CE07E0471485689337080986" box="[1220,1369,189,215]" pageId="8" pageNumber="37" value="2006-08-04">4 Aug. 2006</date>
, F.-L. Xu.
</paragraph>
<paragraph id="1C6A36E3CE07E04710856929372A0873" blockId="8.[151,1437,152,551]" box="[196,1403,263,290]" pageId="8" pageNumber="37">
<emphasis id="2EA1EAF1CE07E04710856929331F0870" bold="true" box="[196,334,263,289]" pageId="8" pageNumber="37">Host plant.</emphasis>
Bamboo [
<taxonomicName id="DBD54D60CE07E047118B692631380873" authority="Hsueh &amp; Yi" authorityName="Hsueh &amp; Yi" box="[458,873,264,290]" class="Liliopsida" family="Poaceae" genus="Qiongzhuea" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="8" pageNumber="37" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="2EA1EAF1CE07E047118B692630870870" box="[458,726,264,289]" italics="true" pageId="8" pageNumber="37">Qiongzhuea communis</emphasis>
Hsueh &amp; Yi
</taxonomicName>
and
<taxonomicName id="DBD54D60CE07E04713E3692637210873" authority="(Hayata) Naki" authorityName="(Hayata) Naki" box="[930,1392,264,290]" class="Liliopsida" family="Poaceae" genus="Sinobambusa" kingdom="Plantae" order="Poales" pageId="8" pageNumber="37" phylum="Tracheophyta" rank="species" species="kunishii">
<emphasis id="2EA1EAF1CE07E04713E3692636E00870" box="[930,1201,264,289]" italics="true" pageId="8" pageNumber="37">Sinobambusa kunishii</emphasis>
(Hayata) Naki
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="54CF6568CE07E0471085690336EE0816" box="[196,1215,301,327]" pageId="8" pageNumber="37" type="distribution">
<paragraph id="1C6A36E3CE07E0471085690336EE0816" blockId="8.[151,1437,152,551]" box="[196,1215,301,327]" pageId="8" pageNumber="37">
<emphasis id="2EA1EAF1CE07E0471085690333350816" bold="true" box="[196,356,301,327]" pageId="8" pageNumber="37">Distribution.</emphasis>
Southwest
<collectingCountry id="64C27673CE07E04711AC690330620816" box="[493,563,301,327]" name="China" pageId="8" pageNumber="37">China</collectingCountry>
(Guizhou: Leishan, Suiyang, Daozhen, Libo) (
<figureCitation id="84EE2A66CE07E047141B690336E30816" box="[1114,1202,301,327]" captionStart="FIGURE 35" captionStartId="7.[151,255,1799,1823]" captionTargetBox="[167,1421,893,1769]" captionTargetId="figure@7.[151,1436,878,1778]" captionTargetPageId="7" captionText="FIGURE 35. Geographic distribution of Kakuna species: K. kuwayamai Matsumura (♦); K. lii sp. nov. (●); K. nonspina sp. nov. (); K. yushaniae (Yang, 1989) (ˑ); K. zhongtuana sp. nov. (˔)." httpUri="https://zenodo.org/record/193348/files/figure.png" pageId="8" pageNumber="37">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="54CF6568CE07E0471085697C3636088D" pageId="8" pageNumber="37" type="discussion">
<paragraph id="1C6A36E3CE07E0471085697C3636088D" blockId="8.[151,1437,152,551]" pageId="8" pageNumber="37">
<emphasis id="2EA1EAF1CE07E0471085697C336F083D" bold="true" box="[196,318,338,364]" pageId="8" pageNumber="37">Remarks.</emphasis>
This new species is similar to
<taxonomicName id="DBD54D60CE07E04712EE697C3108083A" box="[687,857,338,363]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="2EA1EAF1CE07E04712EE697C3108083A" box="[687,857,338,363]" italics="true" pageId="8" pageNumber="37">K. kuwayamai</emphasis>
</taxonomicName>
and
<taxonomicName id="DBD54D60CE07E04713D5697C3672083A" box="[916,1059,338,363]" class="Insecta" family="Delphacidae" genus="Kakuna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nonspina">
<emphasis id="2EA1EAF1CE07E04713D5697C3672083A" box="[916,1059,338,363]" italics="true" pageId="8" pageNumber="37">K. nonspina</emphasis>
</taxonomicName>
, but can be distinguished by:
<emphasis id="2EA1EAF1CE07E04715CB697C37C3083A" box="[1418,1426,338,363]" italics="true" pageId="8" pageNumber="37">i</emphasis>
) body color paler, forewing with the marking of apex of IA unclear;
<emphasis id="2EA1EAF1CE07E0471398695631BB08C0" box="[985,1002,376,401]" italics="true" pageId="8" pageNumber="37">ii</emphasis>
) anal segment without process;
<emphasis id="2EA1EAF1CE07E0471539695637C308C0" box="[1400,1426,376,401]" italics="true" pageId="8" pageNumber="37">iii</emphasis>
) aedeagus with two rows teeth along lateral margin of apical half;
<emphasis id="2EA1EAF1CE07E04713DC69B331E308E7" box="[925,946,413,438]" italics="true" pageId="8" pageNumber="37">iv</emphasis>
) two processes of diaphragm with a long common stalk;
<emphasis id="2EA1EAF1CE07E047110D69EC3308088A" box="[332,345,450,475]" italics="true" pageId="8" pageNumber="37">v</emphasis>
) genital style with a nipple-like process at middle of inner margin.
</paragraph>
</subSubSection>
<subSubSection id="54CF6568CE07E047108569C9314B0B76" pageId="8" pageNumber="37" type="etymology">
<paragraph id="1C6A36E3CE07E047108569C9314B0B76" blockId="8.[151,1437,152,551]" pageId="8" pageNumber="37">
<emphasis id="2EA1EAF1CE07E047108569C933050B50" bold="true" box="[196,340,487,513]" pageId="8" pageNumber="37">Etymology.</emphasis>
The name is derived from transliteration of the Chinese “zhongtu”, meaning genital style having a nipple-like process at middle of inner margin.
</paragraph>
</subSubSection>
</treatment>
</document>