552 lines
74 KiB
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552 lines
74 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1196.112146" ID-Pensoft-Pub="1313-2970-1196-15" ID-Pensoft-UUID="E53B8BDDAE9B58BDBE4B7058562B2B14" ID-ZooBank="7954DFBF803A48F5B79142DD09FE5D01" ModsDocID="1313-2970-1196-15" checkinTime="1711274297545" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Huang, Chih-Wei, Sutcharit, Chirasak & Lee, Chi-Tse" docDate="2024" docId="9306C6D6F206550BA92197BD95C518F4" docLanguage="en" docName="ZooKeys 1196: 15-78" docOrigin="ZooKeys 1196" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/zookeys.1196.112146" docTitle="Amphidromus ingens Moellendorff 1900" docType="treatment" docVersion="2" id="E53B8BDDAE9B58BDBE4B7058562B2B14" lastPageNumber="15" masterDocId="E53B8BDDAE9B58BDBE4B7058562B2B14" masterDocTitle="The arboreal snail genus Amphidromus Albers, 1850 (Eupulmonata, Camaenidae) of Southeast Asia: 1. Molecular systematics of some Vietnamese species and related species from Cambodia, Indonesia, and Laos" masterLastPageNumber="78" masterPageNumber="15" pageNumber="15" updateTime="1711274602338" updateUser="ExternalLinkService">
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<mods:title id="47A7C465FCE12C55ECB732975399F5EB">The arboreal snail genus Amphidromus Albers, 1850 (Eupulmonata, Camaenidae) of Southeast Asia: 1. Molecular systematics of some Vietnamese species and related species from Cambodia, Indonesia, and Laos</mods:title>
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<mods:namePart id="6F2E62F7569A8A8FB99B3A607E047B2C">Jirapatrasilp, Parin</mods:namePart>
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<mods:affiliation id="859D9962CE0024A1F30219ED04831423">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand & Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, Hamburg, Germany</mods:affiliation>
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<mods:namePart id="EF9B022F50762EFFF3F4E50992D102D5">Huang, Chih-Wei</mods:namePart>
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<mods:nameIdentifier id="4ECB86A8D232F4D7E9E5B9609437A764" type="ORCID">https://orcid.org/0000-0002-2921-4294</mods:nameIdentifier>
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<mods:affiliation id="848CF24F84005D04A58875BC98C8B0F7">School of Life Science, National Taiwan Normal University, Taipei, Taiwan</mods:affiliation>
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<mods:namePart id="4E39AC75D6E9F3B5370460D753159AC5">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="7CC164B97B2E4168251D5A63FFE5FDED">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
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<mods:nameIdentifier id="B55A4B7DFF8EA0E310E253D1118EBC81" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="5E793E4C7B2B80780F1336B3CA85509B">Lee, Chi-Tse</mods:namePart>
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<mods:affiliation id="FA9A7734F329638AB19BC2CC2BD68A91">Department of Life Sciences, National Chung Hsing University, Taichung, Taiwan</mods:affiliation>
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<treatment id="9306C6D6F206550BA92197BD95C518F4" LSID="urn:lsid:plazi:treatment:9306C6D6F206550BA92197BD95C518F4" httpUri="http://treatment.plazi.org/id/9306C6D6F206550BA92197BD95C518F4" lastPageNumber="15" pageId="0" pageNumber="15">
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<subSubSection id="7D1CAC9C255D8D89855F34941C8C6DB4" pageId="0" pageNumber="15" type="nomenclature">
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<paragraph id="B27FE8352ECABA2BF9FB85D2FA16EF98" pageId="0" pageNumber="15">
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<taxonomicName id="98ED9F7C14AAF19F2A43EB1EEB7BF0ED" ID-CoL="66G2B" LSID="9306C6D6-F206-550B-A921-97BD95C518F4" authority="Moellendorff, 1900" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
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Amphidromus ingens
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<normalizedToken id="90C2D28CFFBAF7B17D53174AC027DE65" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1900
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="10817C6BA56ACFEAF4C90CF09B1DC5B4" pageId="0" pageNumber="15" type="description">
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<paragraph id="79BD7A4B23E730F21FC0E1F4DE9EA6F5" pageId="0" pageNumber="15">
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<figureCitation id="FEED230CD5AAC4C4925CCD45F86D4457" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">Figs 3</figureCitation>
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<figureCitation id="5C0DDC4FD53E85C8216E9C40AF4163ED" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the radula of Amphidromus spp A Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100) B Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001) C Amphidromus bozhii Wang, 2019 from Tuy Hoa, Phu Yen, Vietnam (NMNS- 8764 - 016). Central teeth are marked in yellow. The left and right images show the outer and inner sections of each radula, respectively." figureDoi="10.3897/zookeys.1196.112146.figure4" httpUri="https://binary.pensoft.net/fig/1007822" pageId="0" pageNumber="15">, 4A</figureCitation>
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<figureCitation id="78DDD1B4F8B55BD76CA014798D3E1646" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber and gametolytic duct D-F Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15">, 5A-C</figureCitation>
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<figureCitation id="8790B67AAEEC1EA0EE62F0A3C1CDC28B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Living Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Dak Lak, Vietnam D Amphidromus placostylus Moellendorff, 1900 from Hoai An, An Lao, Binh Dinh, Vietnam E, F Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam G Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam H, I Amphidromus buelowi Fruhstorfer, 1905 from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam J Amphidromus thachi Huber, 2015 from Krong Bong, Dak Lak, Vietnam." figureDoi="10.3897/zookeys.1196.112146.figure6" httpUri="https://binary.pensoft.net/fig/1007824" pageId="0" pageNumber="15">, 6A-C</figureCitation>
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<figureCitation id="818BFBB592A04A3BE69567ADEF4465DA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mitochondrial haplotype minimum spanning networks of Amphidromus ingens Moellendorff, 1900 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure7" httpUri="https://binary.pensoft.net/fig/1007825" pageId="0" pageNumber="15">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1CC484D02D59359583ED0CA8832E3142" pageId="0" pageNumber="15" type="reference_group">
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<paragraph id="1E3CAAF58AC3371B39F53D6E04FE9035" pageId="0" pageNumber="15">
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<taxonomicName id="7FC400EFBDEAFA77C8FAC264F0940BDC" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">Amphidromus ingens</taxonomicName>
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<normalizedToken id="DA20E8871F2C7DA7EAEB95F6FFBA7863" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1900b: 23-24. Type locality: Berg "Mutter und Kind", Annam [Vietnam].
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<bibRefCitation id="9752A51FB7563CE2E22755B34965662D" author="Pilsbry, HA" journalOrPublisher="Academy of Natural Sciences of Philadelphia, Philadelphia" pageId="0" pageNumber="15" publicationUrl="https://www.biodiversitylibrary.org/item/16708" refId="B48" refString="Pilsbry, HA, 1900. Manual of Conchology, Second Series, Vol. 13. Academy of Natural Sciences of Philadelphia, Philadelphia, https://www.biodiversitylibrary.org/item/16708" title="Manual of Conchology, Second Series, Vol. 13." url="https://www.biodiversitylibrary.org/item/16708" year="1900">Pilsbry 1900</bibRefCitation>
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||
: 175-176.
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<bibRefCitation id="3C8E76CEDBD6B7BF2DBC537EEB5D70D0" DOI="https://doi.org/10.5962/bhl.title.50990" author="Fischer, H" editor="Leroux, E" journalOrPublisher="Leroux, E., Paris" pageId="0" pageNumber="15" pagination="390 - 450" refId="B14" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. Leroux, E., Paris: 390 - 450, DOI: https://doi.org/10.5962/bhl.title.50990" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." url="https://doi.org/10.5962/bhl.title.50990" volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 406.
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<bibRefCitation id="1706904292D6815A747992929CFD650A" DOI="https://doi.org/10.5962/bhl.title.2832" author="Laidlaw, FF" journalOrPublisher="Zoology" pageId="0" pageNumber="15" pagination="507 - 677" refId="B34" refString="Laidlaw, FF, Solem, A, 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana. Zoology 41: 507 - 677, DOI: https://doi.org/10.5962/bhl.title.2832" title="The land snail genus Amphidromus: A synoptic catalogue. Fieldiana." url="https://doi.org/10.5962/bhl.title.2832" volume="41" year="1961">Laidlaw and Solem 1961</bibRefCitation>
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: 529, 629.
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<bibRefCitation id="5684E20A0611F785DE9E24B2C8FD8221" author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="15" pagination="1 - 479" publicationUrl="https://www.biodiversitylibrary.org/item/262223" refId="B51" refString="Richardson, L, 1985. Camaenidae: Catalogue of Species. Tryonia 12: 1 - 479, https://www.biodiversitylibrary.org/item/262223" title="Camaenidae: Catalogue of Species." url="https://www.biodiversitylibrary.org/item/262223" volume="12" year="1985">Richardson 1985</bibRefCitation>
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: 21.
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<bibRefCitation id="34A16695BBA144FBFD67C5347458F7D2" author="Thach, NN" journalOrPublisher="Conchbooks, Hackenheim" pageId="0" pageNumber="15" refId="B71" refString="Thach, NN, 2005. Shells of Vietnam. Conchbooks, Hackenheim" title="Shells of Vietnam." year="2005">Thach 2005</bibRefCitation>
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||
: 235, pl. 73, fig. 22.
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<bibRefCitation id="FCE1FC35B9CAA759AAD349BFBC3672FC" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="15" pagination="1 - 68" publicationUrl="https://www.biotaxa.org/Ruthenica/article/view/3603" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68, https://www.biotaxa.org/Ruthenica/article/view/3603" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." url="https://www.biotaxa.org/Ruthenica/article/view/3603" volume="21" year="2011">Schileyko 2011</bibRefCitation>
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: 50.
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<bibRefCitation id="09F078DA5653E2CFB952D6F4428A4696" DOI="https://doi.org/10.1080/00222933.2021.1933230" author="Sutcharit, C" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="15" pagination="1059 - 1069" refId="B69" refString="Sutcharit, C, Naggs, F, Ablett, JD, Sang, PV, Hao, LV, Panha, S, 2021. Anatomical note on a tree snail Amphidromus (Amphidromus) cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata: Camaenidae). Journal of Natural History 55 (17-18): 1059 - 1069, DOI: https://doi.org/10.1080/00222933.2021.1933230" title="Anatomical note on a tree snail Amphidromus (Amphidromus) cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata: Camaenidae)." url="https://doi.org/10.1080/00222933.2021.1933230" volume="55" year="2021">Sutcharit et al. 2021</bibRefCitation>
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: fig. 1g.
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||
</paragraph>
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||
<paragraph id="B88661BAA14B12EAAB007BD05568D9BB" pageId="0" pageNumber="15">
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<taxonomicName id="10D40C4DB562F4CD2C62A5C95B5584F2" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
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<taxonomicName id="C78DA578CEFD45A0F32F26F8138762CB" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus (Amphidromus) ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens" subGenus="Amphidromus">Amphidromus (Amphidromus) ingens</taxonomicName>
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.
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<bibRefCitation id="7B91FDBBBBEC43E1A4A0065BB81E8A0B" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="15" pagination="131 - 140" refId="B83" refString="Zilch, A, 1953. Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1). Archiv fuer Molluskenkunde 82: 131 - 140" title="Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1)." volume="82" year="1953">Zilch 1953</bibRefCitation>
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||
: 135, pl. 23, fig. 25.
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||
</paragraph>
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||
<paragraph id="B1AB322745F9F997D35ACA0E9241C504" pageId="0" pageNumber="15">
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<taxonomicName id="12B7022C0E411C7DACC2E809F16FC2B5" authorityName="Thach & F.Huber" authorityYear="2014" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus naggsi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">Amphidromus naggsi</taxonomicName>
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Thach & Huber, 2014: 35-37, figs 1-13, 15. Type locality: Don Duong district, Lam Dong Province, South Vietnam.
|
||
<bibRefCitation id="5F5F7D2D0E5B6D099B83B7E9C5D1CE42" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="15" pagination="35 - 76" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
|
||
<normalizedToken id="BD5EA1442B28DC5A28AA843CE251975B" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
et al. 2020
|
||
</bibRefCitation>
|
||
: 53.
|
||
<bibRefCitation id="7318580C040517A1C4A2E7B4C31A06E1" DOI="https://doi.org/10.54173/F522184" author="Thach, NN" journalOrPublisher="The Festivus" pageId="0" pageNumber="15" pagination="184 - 190" refId="B79" refString="Thach, NN, Simone, LRL, Parsons, J, Abbas, J, Huber, F, 2020. Comments on "Vandalism" in Malacology. The Festivus 52 (2): 184 - 190, DOI: https://doi.org/10.54173/F522184" title="Comments on " Vandalism " in Malacology." url="https://doi.org/10.54173/F522184" volume="52" year="2020">Thach et al. 2020</bibRefCitation>
|
||
: 185, 187, pl. 1, fig. 6a.
|
||
<bibRefCitation id="BCE2CD40CB9A1160392CAA1919FCF9E7" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B75" refString="Thach, NN, 2020a. New shells of South Asia. Vol. 2. Seashells*Freshwater*Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia. Vol. 2. Seashells * Freshwater * Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error." year="2020 a">Thach 2020a</bibRefCitation>
|
||
: 70, fig. 881 left.
|
||
<bibRefCitation id="3AAB3FD1D3B1BFA3676059AD7E6ED893" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B77" refString="Thach, NN, 2021. New shells of South Asia and Taiwan, China, Tanzania. Seashells*Freshwater*Land snails. With 116 new species and subspecies and rejected synonyms, accepted species. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia and Taiwan, China, Tanzania. Seashells * Freshwater * Land snails. With 116 new species and subspecies and rejected synonyms, accepted species." year="2021">Thach 2021</bibRefCitation>
|
||
: 70. syn. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9A5A2BA3F51CDC6D25C42B621E3B838C" pageId="0" pageNumber="15" type="materials_examined">
|
||
<paragraph id="A6E337F94A4D557CEA94000A5BEB554F" pageId="0" pageNumber="15">Material examined.</paragraph>
|
||
<paragraph id="27FEC09C10BEFC40D404667B79488EBF" pageId="0" pageNumber="15">
|
||
<materialsCitation id="8911A283BC6E59E801668FD639B62361" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Vietnam" location="Dextral" specimenCount="1" typeStatus="lectotype">
|
||
<collectingCountry id="F1637F0C58F3B33CD629526EEEA92966" name="Vietnam">Vietnam</collectingCountry>
|
||
:
|
||
<location id="364EA5D6BCB984C02C73A55BDC1D1C11" LSID="urn:lsid:plazi:treatment:9306C6D6F206550BA92197BD95C518F4:364EA5D6BCB984C02C73A55BDC1D1C11" country="Vietnam" name="Dextral">Dextral</location>
|
||
,
|
||
<emphasis id="11EDBD8C8F1C4999A1C90AEFF4379E8E" bold="true" italics="true" pageId="0" pageNumber="15">
|
||
<typeStatus id="070B6D8B93B33174D4AD23A56D03AAE7" type="lectotype">lectotype</typeStatus>
|
||
</emphasis>
|
||
of "
|
||
<taxonomicName id="C9BF6C842EE363794BCC8D9683FE34D7" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="2B8D59B2E34D714B8C93F7ED5AFFB8D5" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
", SMF 7565/1 (Fig.
|
||
<figureCitation id="A61BBA873B11913C693F8AEC6D16D390" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3A</figureCitation>
|
||
); 2D + 2S,
|
||
<typeStatus id="40338945CF3217C9DCDC8AFBA20C941F" type="paralectotype">paralectotypes</typeStatus>
|
||
of "
|
||
<taxonomicName id="7A02C798AC5741AD7CD2348CC4F0BA6A" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="32911E62727E3F48FA0B7FD4D840AB91" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
", SMF 7566/4 (Fig.
|
||
<figureCitation id="6CEC300A58BF404EF4D59A7364630D88" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3B</figureCitation>
|
||
).
|
||
<collectingCountry id="4A8C31E3B987B5CB0CE5AD16EE7C4893" name="Vietnam">Vietnam</collectingCountry>
|
||
: Dextral,
|
||
<emphasis id="FB2A58E00E86B3AE0D2FB20546908C54" bold="true" italics="true" pageId="0" pageNumber="15">
|
||
<typeStatus id="631924D0275E5AAA28DDD371FBA4AAF7" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
of "
|
||
<taxonomicName id="F9AE22A44130FD883E3CD70950ADA431" authorityName="Thach & F.Huber" authorityYear="2014" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus naggsi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="EA37D77C178C984FBE88D8921993F085" italics="true" pageId="0" pageNumber="15">Amphidromus naggsi</emphasis>
|
||
</taxonomicName>
|
||
", RMNH.5003908 (Fig.
|
||
<figureCitation id="2C4A02F5C3FF851458644051C5478C73" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="D013D382CD31D3C9F479236D07F82E16" doi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15" start="Figure 3" startId="F3">
|
||
<paragraph id="5D667696985BA74A94C3F815F218906F" pageId="0" pageNumber="15">
|
||
<emphasis id="841CA5108948D60256EFB251BD1571B1" bold="true" pageId="0" pageNumber="15">Figure 3.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="BEABC06DFB69BF3964CDC2814612C830" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="C06C4290D86FD1B2E8D024EB589C76B2" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="1C363BE83D379BC82726424EC9F1A5DB" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900
|
||
<emphasis id="BB86EB541DD94EF73E970EA44E2DD1F6" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
<typeStatus id="3BFF508D30DD8F464DC0A4EC7B28E20D" type="lectotype">lectotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="9DFE3D2A4DBB8CB4882DA40679C9EA76" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="156B7AC8D4C255455B14C4307742E904" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
" (SMF 7565)
|
||
<emphasis id="6B3F1541F1B449A21817278CEB4C5FBB" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
<typeStatus id="1CB1E13ABDFB055270F34868C05B7338" type="paralectotype">paralectotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="431E2B25FD0817A0FC599A74CE1F0D33" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="240417C7F8F65AACE82A5D201C79DACC" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
" (SMF 7566)
|
||
<emphasis id="2F615C5796489C8FBDC68D0BFFDEA41F" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
<typeStatus id="56A1A6FA88A2BBE197743153E37FD2E0" type="holotype">holotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="061D6FDA9C29FAC7866F5D0879AA01A0" authorityName="Thach & F.Huber" authorityYear="2014" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus naggsi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="26DFD2AA0A4F5B3644F512D9A916188A" italics="true" pageId="0" pageNumber="15">Amphidromus naggsi</emphasis>
|
||
</taxonomicName>
|
||
" (RMNH.5003908)
|
||
<emphasis id="D7F8D2EBFF62926534EFA0AFDCFDC8DA" bold="true" pageId="0" pageNumber="15">D, E</emphasis>
|
||
specimens from
|
||
<normalizedToken id="F0609A21378ECD5CAD9DC31B54F6CD5A" originalValue="M’drak">M'drak</normalizedToken>
|
||
,
|
||
<collectingRegion id="14560D772E75C47C3E350B230F490791" country="Vietnam" name="Dak Lak">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingCountry id="F421A121BB87C22064161ED8AE3535A6" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-082, NMNS-8764-084)
|
||
<emphasis id="5823523105D488CE0EE0735EC19B73BF" bold="true" pageId="0" pageNumber="15">F</emphasis>
|
||
specimen from Ea
|
||
<normalizedToken id="38103E6B419BF7833D434E0861930E85" originalValue="M’doal">M'doal</normalizedToken>
|
||
,
|
||
<normalizedToken id="D24E99CE0351B0FAD9D07592C13B9DFF" originalValue="M’drak">M'drak</normalizedToken>
|
||
,
|
||
<collectingRegion id="8588C136DB73B7833D012B545B058D6B" country="Vietnam" name="Dak Lak">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingCountry id="9D81361BF947AA56E4769756C3184479" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-087)
|
||
<emphasis id="F2D1C16130CF483AD97335AE6254B793" bold="true" pageId="0" pageNumber="15">G</emphasis>
|
||
specimen from Krong A,
|
||
<normalizedToken id="782A8ECB34E8ADECADCA993FBBA5F981" originalValue="M’drak">M'drak</normalizedToken>
|
||
,
|
||
<collectingRegion id="3F7992575027F7E2AC0ADAD26D1C1CD3" country="Vietnam" name="Dak Lak">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingCountry id="F64843D09DDF8E75895B08B8018587C8" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-088)
|
||
<emphasis id="A63CA607E65B75CCEA8D832E4357DA25" bold="true" pageId="0" pageNumber="15">H, I</emphasis>
|
||
specimens from Ea Sup,
|
||
<collectingRegion id="7E48F062E19E2837D692BEDBA856E4A5" country="Vietnam" name="Dak Lak">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingCountry id="06628BA59E3489F2874A001F95DFEB3E" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-093, NMNS-8764-101). Credit: J. Goud, RMNH (
|
||
<emphasis id="50CA721683F8A513568FC6B9BF27825B" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="D4F96722171D51A989D5E00AB9526609" pageId="0" pageNumber="15" type="materials_examined">
|
||
<paragraph id="8963BE080820DA79C26EC5DB4F17F8DA" pageId="0" pageNumber="15">Other material examined.</paragraph>
|
||
<paragraph id="9E23D7025E35FDCA1674FD1CB4D450DB" pageId="0" pageNumber="15">
|
||
<materialsCitation id="2CE75F877F015C651AD27F0DABE1A802" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Krong A" country="Vietnam" location="Ea Sup District" specimenCount="1" stateProvince="Dak Lak Province">
|
||
<collectingCountry id="AE2BD83BC10E0A5D3AC310C15C2E2102" name="Vietnam">Vietnam</collectingCountry>
|
||
: 4D + 1S specimens,
|
||
<normalizedToken id="CDACE04AC9E763E3D8C7C9B3A162FA07" originalValue="M’drak">M'drak</normalizedToken>
|
||
District,
|
||
<collectingRegion id="A308226D2B061A7F4508E2DDB420B4F8" country="Vietnam" name="Dak Lak">Dak Lak Province</collectingRegion>
|
||
, NMNS-8764-082- NMNS-8764-086 (Fig.
|
||
<figureCitation id="39310953B5647A300FC28507C52CAE94" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3D, E</figureCitation>
|
||
); 1D specimen, Ea
|
||
<normalizedToken id="4E13723F45BA0996FEC82B42C423B35F" originalValue="M’doal">M'doal</normalizedToken>
|
||
ward,
|
||
<normalizedToken id="373405D66DB327729AC1FC078801D3C8" originalValue="M’drak">M'drak</normalizedToken>
|
||
District,
|
||
<collectingRegion id="FC501A2E14FCF7C9D36732C6E38973BE" country="Vietnam" name="Dak Lak">Dak Lak Province</collectingRegion>
|
||
, NMNS-8764-087 (Fig.
|
||
<figureCitation id="EB8AA4E4799305E589F24718A52E37B6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3F</figureCitation>
|
||
); 4D + 1S specimens,
|
||
<collectorName id="F45AD5D65FF75460B0DCF85143FE11DF">Krong A</collectorName>
|
||
ward,
|
||
<normalizedToken id="7D3514F75F2117D620D2F60E1C7C9838" originalValue="M’drak">M'drak</normalizedToken>
|
||
District,
|
||
<collectingRegion id="D58A3ED267DE513923052E602C12CCFB" country="Vietnam" name="Dak Lak">Dak Lak Province</collectingRegion>
|
||
, NMNS-8764-088- NMNS-8764-092 (Fig.
|
||
<figureCitation id="C92FDB907C0BC3476BD4314347037CC2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3G</figureCitation>
|
||
); 7D + 5S specimens,
|
||
<location id="FDF3D6212E32D2E7F0788D3556AF9F13" LSID="urn:lsid:plazi:treatment:9306C6D6F206550BA92197BD95C518F4:FDF3D6212E32D2E7F0788D3556AF9F13" country="Vietnam" name="Ea Sup District" stateProvince="Dak Lak Province">Ea Sup District</location>
|
||
,
|
||
<collectingRegion id="E76730973A5C7B97BC2D2701634987AF" country="Vietnam" name="Dak Lak">Dak Lak Province</collectingRegion>
|
||
, NMNS-8764-093- NMNS-8764-104 (Fig.
|
||
<figureCitation id="DE5692FD831CB67209D7DFDA82DC5345" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3H, I</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F457932B05B885307B112402F426677B" pageId="0" pageNumber="15" type="diagnosis">
|
||
<paragraph id="9928212EDF32F7B7EED9C2B9C74CA977" pageId="0" pageNumber="15">Diagnosis.</paragraph>
|
||
<paragraph id="2481D8AD697F45F71D237FB12A7F6332" pageId="0" pageNumber="15">Shell large conical and chirally dimorphic (sinistral and dextral coiling). Shell surface with coarse growth lines; last whorl with subsutural depression area and more or less prominent keel on periphery. Genitalia with appendix.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F3822216983A634018E69422495607" pageId="0" pageNumber="15" type="differential diagnosis">
|
||
<paragraph id="FF83B6F553E49884A18C4C7918F7B78C" pageId="0" pageNumber="15">Differential diagnosis.</paragraph>
|
||
<paragraph id="10A25FE83CC602BBAB5A328B9E0946D7" pageId="0" pageNumber="15">
|
||
<taxonomicName id="0F7E3500C394DF6043445A6F22DA0925" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="BD927D7263B9CFF795BF8A397138FC84" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
is unique among all reported Vietnamese species (
|
||
<bibRefCitation id="2ED0924111EE62CCC86136C17FE89921" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="15" pagination="1 - 68" publicationUrl="https://www.biotaxa.org/Ruthenica/article/view/3603" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68, https://www.biotaxa.org/Ruthenica/article/view/3603" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." url="https://www.biotaxa.org/Ruthenica/article/view/3603" volume="21" year="2011">Schileyko 2011</bibRefCitation>
|
||
) in having a last whorl with subsutural depression area and more or less prominent keel on periphery.
|
||
<taxonomicName id="ED967935A5C1CEC2EFBA6A452F1A8D63" authorityName="Wang" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus bozhii" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bozhii">
|
||
<emphasis id="11704EB6CE40A585A3C27547BB2DC0E2" italics="true" pageId="0" pageNumber="15">Amphidromus bozhii</emphasis>
|
||
</taxonomicName>
|
||
is similar in most of the shell form and sculpture, but the shell sculpture of
|
||
<taxonomicName id="110A03117EEA9DCE29CCA184996341AE" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. bozhii" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bozhii">
|
||
<emphasis id="AFCC0C195CEF4B87B103F1857DC65395" italics="true" pageId="0" pageNumber="15">A. bozhii</emphasis>
|
||
</taxonomicName>
|
||
has a very weak spiral depression area and sometimes with or without keel, and the shell colour is generally rose-pink to dark colour, with last whorl stained with dark brown colour below periphery and ~ 1/2 of upper periphery. On the other hand,
|
||
<taxonomicName id="FA447F7FC02455675EAE3128DAFA7D66" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="BEAEA7E3DA0107E6733C46E1B70EEC88" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
has a monochrome (whitish, yellowish, tinted pink) shell, often stained with dark brown to blackish below periphery.
|
||
<taxonomicName id="4A74B80D9DC1671CE85893886E821E31" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="95E01BD6EEBB672F660D142DF45953CD" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
is also recognised by a distinct clade in the molecular phylogeny (Fig.
|
||
<figureCitation id="9569BEEE9732CF31743E362433E22B1D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian phylogeny of Amphidromus spp. based on mitochondrial COI and 16 S genes. Nodal support values are given as SH-aLRT / aBayes / ultra-fast bootstrap (IQ-TREE, ML) / posterior probability (MrBayes, BI). An asterisk on each branch indicates a clade with all well-supported values (SH-aLRT ≥ 80 %, aBayes ≥ 0.95, BS ≥ 95 %, PP ≥ 0.95). Species numbers correspond to those in Fig. 1 and Tables 1, 2." figureDoi="10.3897/zookeys.1196.112146.figure2" httpUri="https://binary.pensoft.net/fig/1007820" pageId="0" pageNumber="15">2</figureCitation>
|
||
), with the closest
|
||
<emphasis id="9AD7A88EF6C13B515FFC849E64F0ABD8" italics="true" pageId="0" pageNumber="15">p</emphasis>
|
||
-distance to
|
||
<taxonomicName id="975695E0ACE7BB0A38E2FBA7AE2FC368" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingensoides" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingensoides">
|
||
<emphasis id="404FA6EA4BD66004BC86B378C8D140A5" italics="true" pageId="0" pageNumber="15">A. ingensoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in COI (10.2%) and
|
||
<taxonomicName id="65F05E89D92B1C34A9D138389B6CE92C" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. bozhii" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bozhii">
|
||
<emphasis id="52B7A911A6A691BA1BD1CEF5EB6EF230" italics="true" pageId="0" pageNumber="15">A. bozhii</emphasis>
|
||
</taxonomicName>
|
||
in 16S (2.76%) (Table
|
||
<tableCitation id="45404D3F1D32DCBF4D674BD2D56565D7" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Percentage of uncorrected pairwise interspecific distances for the partial COI (above the diagonal) and 16 S rRNA (below the diagonal) gene fragments among the Amphidromus species in this study. Intraspecific distances for COI / 16 S rRNA are shown on the diagonal (bold). Species numbers correspond to those in Figs 1, 2, and Table 1. The numbers of sequences used to calculate the distances of each respective gene fragment are given as n = COI, 16 S in the first column." httpUri="http://table.plazi.org/id/A1B4F960F999761C3840BD02D5BE2608" pageId="0" pageNumber="15" tableUuid="A1B4F960F999761C3840BD02D5BE2608">2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="190B28EC7599A317A8ADFA7584671F25" pageId="0" pageNumber="15" type="description">
|
||
<paragraph id="FE02C47B1746C04EB366A243F83C2D07" pageId="0" pageNumber="15">Description.</paragraph>
|
||
<paragraph id="1368340B60FB17C0A5B6AAFC197E8C02" pageId="0" pageNumber="15">
|
||
<emphasis id="DF39FBE61DDCC5E66C922B93CD287676" bold="true" italics="true" pageId="0" pageNumber="15">Shell</emphasis>
|
||
large (height 62.3-74.6 mm, width 38.5-42.5 mm), chirally dimorphic, solid, and ovate conical shape. Spire long conical to elongate conical, apex acute without black spot on tip. Whorls 5-7 convex; suture wide and depressed; last whorl rounded to slightly angulated. Periostracum brownish to thin corneous; varix usually absent. Shell surface generally with irregular and coarse growth lines; below sutural with broad subsutural depression area, and with blunt or low to prominent keel on periphery. Shell colour highly variable: monochrome (whitish, yellowish, tinted pink) to stained with dark brown to blackish below periphery. Parietal callus thickened and white, dilated at umbilical area. Aperture broadly ovate; inner side of outer wall with white, yellow or dark brown to blackish colour. Peristome thickened, expanded, and reflexed but not attached to last whorl; lip whitish. Columella white, straight, or little twisted. Umbilicus imperforate.
|
||
</paragraph>
|
||
<paragraph id="DC0FC9E98F4B4775AAD981A989B5005D" pageId="0" pageNumber="15">
|
||
<emphasis id="6282B9D6B9AA4BB0F7871A720E9D7B41" bold="true" pageId="0" pageNumber="15">
|
||
<emphasis id="EB23D94CC0AA9BA91CBD390CD57C4A45" italics="true" pageId="0" pageNumber="15">Radula</emphasis>
|
||
.
|
||
</emphasis>
|
||
Teeth arranged in anteriorly pointed V-shaped rows. Central tooth monocuspid and spatulate with truncated cusp. Lateral teeth bicuspid; endocone small, slightly curved, with wide notch and dull cusp; ectocone large with truncated to slightly curved cusp. Lateral teeth gradually transformed to asymmetric tricuspid marginal teeth (Fig.
|
||
<figureCitation id="2C04190248FE562FED530D3C20B1E333" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of the radula of Amphidromus spp A Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100) B Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001) C Amphidromus bozhii Wang, 2019 from Tuy Hoa, Phu Yen, Vietnam (NMNS- 8764 - 016). Central teeth are marked in yellow. The left and right images show the outer and inner sections of each radula, respectively." figureDoi="10.3897/zookeys.1196.112146.figure4" httpUri="https://binary.pensoft.net/fig/1007822" pageId="0" pageNumber="15">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="47579C7D0D8081936D949E32935031E7" doi="10.3897/zookeys.1196.112146.figure4" httpUri="https://binary.pensoft.net/fig/1007822" pageId="0" pageNumber="15" start="Figure 4" startId="F4">
|
||
<paragraph id="C43B354DEF4C9E3E05E61165789F7CC9" pageId="0" pageNumber="15">
|
||
<emphasis id="B8BF5335D4AA34E8DAAD86B0EEF9032C" bold="true" pageId="0" pageNumber="15">Figure 4.</emphasis>
|
||
SEM images of the radula of
|
||
<taxonomicName id="DFFDDE573DAD1DCE27BD582550DFF3C0" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7E18057B55B76D9FC17EEC24BDF4004B" italics="true" pageId="0" pageNumber="15">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis id="1022CB8B4655949CCEAAD9D64292A424" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
<taxonomicName id="5D0B79161AD09B23E09125FFDB510652" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="0EC1F01126653E30EC3E7588936463D4" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="E840446C825EFB3E58D08D62C23DA32B" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS-8764-100)
|
||
<emphasis id="D0221BFD6503A08C5111493E8F49FBCB" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
<taxonomicName id="952A6C387068D983E89EA6182AF04400" authorityName="Jirapatrasilp & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus asperoides" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="79EA1C1D9CF1124E7144934095CE8A05" italics="true" pageId="0" pageNumber="15">Amphidromus asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS-8764-001)
|
||
<emphasis id="437E23B2C50B76AF501FAC35542AE2AA" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
<taxonomicName id="7B267887E52B8827E8EDDE608930E145" authorityName="Wang" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus bozhii" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bozhii">
|
||
<emphasis id="FF123080D6E6579B7606DA8FAAAF53A5" italics="true" pageId="0" pageNumber="15">Amphidromus bozhii</emphasis>
|
||
</taxonomicName>
|
||
Wang, 2019 from Tuy Hoa, Phu Yen, Vietnam (NMNS-8764-016). Central teeth are marked in yellow. The left and right images show the outer and inner sections of each radula, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0F5DC6C1F21514DC611FCE61A673B0E4" pageId="0" pageNumber="15">
|
||
<emphasis id="24727E0CCDC30C1AA37C7A234D38C887" bold="true" pageId="0" pageNumber="15">
|
||
<emphasis id="4BCC2C2461B9DD81D79BE269EB97BCE3" italics="true" pageId="0" pageNumber="15">Genital organs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Atrium relatively short. Penis slender, conical, and short ~ 1/3 of vaginal length. Penial retractor muscle thickened and inserting on epiphallus close to penis. Epiphallus long, slender tube, slightly narrower than penis. Flagellum short ~ 1/2 of epiphallus and terminating in slightly enlarged folded coil. Appendix short, slender tube, approximately as long as epiphallus, and ~ 2
|
||
<normalizedToken id="5E07DEFB4DF131D1FE18F90160D6250C" originalValue="×">x</normalizedToken>
|
||
longer than flagellum. Vas deferens slender tube passing from free oviduct and terminating at epiphallus-flagellum junction (Fig.
|
||
<figureCitation id="60A30C028D198D69BF931822BEC9B7C1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber and gametolytic duct D-F Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15">5A</figureCitation>
|
||
). Internal wall of penis corrugated, exhibiting series of thickened and swollen longitudinal penial pilasters forming fringe around penial wall, and with nearly smooth to weak folds around base of penial verge. Penial verge short conical with nearly smooth surface, and with opening on the tip (Fig.
|
||
<figureCitation id="C3E4C7448925AA7CD4DFE3F91FC72BBC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber and gametolytic duct D-F Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0E1B64E773EC2AD3D9F87784087279FB" doi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15" start="Figure 5" startId="F5">
|
||
<paragraph id="6E48FD1EC0E7FB7B6D57EAC1FAF5AFC9" pageId="0" pageNumber="15">
|
||
<emphasis id="2E0214323EAB530DBFF7C7CAA7B6520C" bold="true" pageId="0" pageNumber="15">Figure 5.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="9B2D68843991E832EA11CC0B220F48B6" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="FA64C3939589F444C8C8FDDA179E8D87" italics="true" pageId="0" pageNumber="15">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis id="EDD5212AD237CFEEBE7D6F186ED6E3B1" bold="true" pageId="0" pageNumber="15">A-C</emphasis>
|
||
<taxonomicName id="6775FDD458CF1656442347BE2012377B" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="D24D98E0961B02096DDF4DA9B19AE815" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="011AEF224F29611F9DE70D049602CE8E" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS-8764-100), showing
|
||
<emphasis id="5F9A3F741B3F4E24FC9C7039204A79C3" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
general view of genitalia
|
||
<emphasis id="845A581B142A47ED071DE8CF92F1A027" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
interior structures of penis
|
||
<emphasis id="2CB72B46329A89F8E1E8F52D7C147E02" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
interior structures of vagina chamber and gametolytic duct
|
||
<emphasis id="96F77FCF4004FA2F073870E1403BE7FB" bold="true" pageId="0" pageNumber="15">D-F</emphasis>
|
||
<taxonomicName id="040B9F853D06EDA1FADBDBB1348C68FB" authorityName="Jirapatrasilp & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus asperoides" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="2E9A97180C02B8F063DB568FDA70A699" italics="true" pageId="0" pageNumber="15">Amphidromus asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS-8764-001), showing
|
||
<emphasis id="18CAB07A243EC94158D27BF599408662" bold="true" pageId="0" pageNumber="15">D</emphasis>
|
||
general view of genitalia
|
||
<emphasis id="36BCD60BE1023C7E7FB01BA49F2AB1CF" bold="true" pageId="0" pageNumber="15">E</emphasis>
|
||
interior structures of penis
|
||
<emphasis id="4A9BA63DB384596223ED5A1CFE6DEF9D" bold="true" pageId="0" pageNumber="15">F</emphasis>
|
||
interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl,
|
||
<normalizedToken id="406BD86C77DC69EAE192AB7F562FF6A8" originalValue="flagellum">flagellum</normalizedToken>
|
||
; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DCB1C75D9AC30AB4C52B00FEE76A14CC" pageId="0" pageNumber="15">
|
||
Vagina slender, long cylindrical, and ~ 3
|
||
<normalizedToken id="299AC47A43187753EE6CF8F4AC4D36E7" originalValue="×">x</normalizedToken>
|
||
longer than penis. Gametolytic duct enlarged cylindrical tube then abruptly tapering to slender tube terminally and connected to gametolytic sac (missing during dissecting). Free oviduct short; oviduct compact and enlarged to form lobule alveoli (Fig.
|
||
<figureCitation id="EAA5139D2BBDB1DF47FEE8FA9FA287C5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber and gametolytic duct D-F Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15">5A</figureCitation>
|
||
). Internal wall of vagina possessing corrugated ridges near genital orifice; ridges becoming thinner and smooth surfaced longitudinal vaginal pilasters, swollen with irregularly shaped and deep crenelations close to free oviduct opening. Spermatophore (in part) dark brown stuck inside gametolytic duct (Fig.
|
||
<figureCitation id="78BEC11C835FA8AAEBA20CFA7B020061" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 100), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber and gametolytic duct D-F Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam (NMNS- 8764 - 001), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure5" httpUri="https://binary.pensoft.net/fig/1007823" pageId="0" pageNumber="15">5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="538EE470EFC37147F94A00F6150B2E95" pageId="0" pageNumber="15">
|
||
<emphasis id="E54AC3B8C1421597B34E76E062779E4C" bold="true" italics="true" pageId="0" pageNumber="15">Living specimens</emphasis>
|
||
generally with pale brown to yellowish body covered with reticulated skin. Foot broad and long with uniform pale brownish to yellowish colour to posterior tail. Dorsal side of anterior body usually with stripe of darkly reticulated skin; head area at base and just behind upper tentacle with orange patch. Upper tentacles drumstick shaped, orange to paler and with dark eyespots on tentacular tips; lower tentacles short and pale orange in colour (Fig.
|
||
<figureCitation id="718CF044AF5E451F6895046694BAA1BA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Living Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Dak Lak, Vietnam D Amphidromus placostylus Moellendorff, 1900 from Hoai An, An Lao, Binh Dinh, Vietnam E, F Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam G Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam H, I Amphidromus buelowi Fruhstorfer, 1905 from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam J Amphidromus thachi Huber, 2015 from Krong Bong, Dak Lak, Vietnam." figureDoi="10.3897/zookeys.1196.112146.figure6" httpUri="https://binary.pensoft.net/fig/1007824" pageId="0" pageNumber="15">6A-C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B6B7AF3FCB99F9D81B0CCFE7819E743A" doi="10.3897/zookeys.1196.112146.figure6" httpUri="https://binary.pensoft.net/fig/1007824" pageId="0" pageNumber="15" start="Figure 6" startId="F6">
|
||
<paragraph id="3A0C9149D9B27B56EA89DED99FD2C2F8" pageId="0" pageNumber="15">
|
||
<emphasis id="D01BBD4CBD17B6BD28042BD6E9D5C1E5" bold="true" pageId="0" pageNumber="15">Figure 6.</emphasis>
|
||
Living
|
||
<taxonomicName id="460F5E7A993153FF8C11DB2923C08EF8" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="43DF60D963A38865D37C3B415CE9C07F" italics="true" pageId="0" pageNumber="15">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis id="7DFCB83520608B9B4CCA269BE3526C2C" bold="true" pageId="0" pageNumber="15">A-C</emphasis>
|
||
<taxonomicName id="DCCC0BD92BE99E2E58C3BB9ADFCF23DE" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="2EBE3EA6D696F64D1C91B0D95099B04F" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="50C4D32FDCFE7DCE4934904525228258" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900 from Dak Lak, Vietnam
|
||
<emphasis id="1F94419113C92EAAA88478B6CACAEE1E" bold="true" pageId="0" pageNumber="15">D</emphasis>
|
||
<taxonomicName id="8D7AA201965FE6983F542B8C2C026E19" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus placostylus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="placostylus">
|
||
<emphasis id="8305104D8CEA2F4C476931B68CB6B3A8" italics="true" pageId="0" pageNumber="15">Amphidromus placostylus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="6901C0AF1C1315A8B74DA992F8AE6445" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900 from Hoai An, An Lao, Binh Dinh, Vietnam
|
||
<emphasis id="705DFFB6554C00310128756725ACD79E" bold="true" pageId="0" pageNumber="15">E, F</emphasis>
|
||
<taxonomicName id="9288D4F0AFF48BA3971BB536F1DDBACB" authorityName="Jirapatrasilp & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingensoides" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingensoides">
|
||
<emphasis id="E71AF54A1602421CD362A4513151D84C" italics="true" pageId="0" pageNumber="15">Amphidromus ingensoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam
|
||
<emphasis id="ABDECB4C9FC117B217E4158FB38CD4B8" bold="true" pageId="0" pageNumber="15">G</emphasis>
|
||
<taxonomicName id="EB2A68AF1ED7BA24B46008E39B6D9C4D" authorityName="Jirapatrasilp & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus asperoides" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="069B9CBF1E685C461404D7F4A035F397" italics="true" pageId="0" pageNumber="15">Amphidromus asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam
|
||
<emphasis id="1629D9B6B9E34CEFB0CD13F18BC58081" bold="true" pageId="0" pageNumber="15">H, I</emphasis>
|
||
<taxonomicName id="7C58D148C1910E4A13423A24A27F5D6C" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="8A0D479B9C20A631D5A5FE07B194D26E" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
Fruhstorfer, 1905 from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam
|
||
<emphasis id="E823D51A752AC16493ED58C67161BD81" bold="true" pageId="0" pageNumber="15">J</emphasis>
|
||
<taxonomicName id="07D7D51ADB8F0E06455154B192149696" authorityName="Huber" authorityYear="2015" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus thachi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="thachi">
|
||
<emphasis id="883FBDC9E95A65F42DB45BC658619FE1" italics="true" pageId="0" pageNumber="15">Amphidromus thachi</emphasis>
|
||
</taxonomicName>
|
||
Huber, 2015 from Krong Bong, Dak Lak, Vietnam.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="36CAC840C4B93D8D4BFDF11FB06DE76E" pageId="0" pageNumber="15" type="haplotype network">
|
||
<paragraph id="59A1F181FDEEBBB6BBAE8BA14FE93D7B" pageId="0" pageNumber="15">Haplotype network.</paragraph>
|
||
<paragraph id="264E04C8CB74320EA1176617F92DB068" pageId="0" pageNumber="15">
|
||
There was a total of 12 COI haplotypes (Fig.
|
||
<figureCitation id="C77A61B6C77068DF628BA770DC1FFB73" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mitochondrial haplotype minimum spanning networks of Amphidromus ingens Moellendorff, 1900 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure7" httpUri="https://binary.pensoft.net/fig/1007825" pageId="0" pageNumber="15">7A</figureCitation>
|
||
) and nine 16S haplotypes (Fig.
|
||
<figureCitation id="BE96E69A464B3F23F4C7DDB2A4C21BC5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mitochondrial haplotype minimum spanning networks of Amphidromus ingens Moellendorff, 1900 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure7" httpUri="https://binary.pensoft.net/fig/1007825" pageId="0" pageNumber="15">7B</figureCitation>
|
||
) of
|
||
<taxonomicName id="D9A03A9668D4B952BF8AB4510D77780D" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="6E6D3600C861A759A04E81EDE644C8B2" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
in this study, and the highest numbers of mutational steps in the COI and 16S minimum spanning networks are 13 and three, respectively.
|
||
</paragraph>
|
||
<caption id="CAF5DF1391EB148A5BBDC59A64C10F9B" doi="10.3897/zookeys.1196.112146.figure7" httpUri="https://binary.pensoft.net/fig/1007825" pageId="0" pageNumber="15" start="Figure 7" startId="F7">
|
||
<paragraph id="CAC810376869E8FB725B9A1A227CC336" pageId="0" pageNumber="15">
|
||
<emphasis id="BA7C66E49D1C3C2D46AE62E8072EBD09" bold="true" pageId="0" pageNumber="15">Figure 7.</emphasis>
|
||
Mitochondrial haplotype minimum spanning networks of
|
||
<taxonomicName id="F92CA20F36B305F48AB1670D830EA00C" authorityName="Moellendorff" authorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus ingens" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="3E4A6071A772E0DB385E29D90695FFC3" italics="true" pageId="0" pageNumber="15">Amphidromus ingens</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="38D39F10C4C29A0CEC2986701254A570" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, 1900
|
||
<emphasis id="104BF84D401F9D6CF4A9445EDF6135C1" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
COI and
|
||
<emphasis id="2E90FFEBEAF5141B378F62CB87A59720" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
16S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table
|
||
<tableCitation id="2FBE23B27548C18E31DECB149E647D77" captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of specimens used in this study with species name, locality details, voucher and GenBank accession numbers. Species numbers correspond to those in Figs 1, 2, and Table 2." httpUri="http://table.plazi.org/id/536020F98423605B1D469F287C9C42E1" pageId="0" pageNumber="15" tableUuid="536020F98423605B1D469F287C9C42E1">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5AAD267C6F3307AB32473E9F6E89E8FF" pageId="0" pageNumber="15" type="distribution">
|
||
<paragraph id="5960EF9F2D727DC6DE58D80F6799C16E" pageId="0" pageNumber="15">Distribution.</paragraph>
|
||
<paragraph id="9FFECBCB68D702079946D24C2A17F71A" pageId="0" pageNumber="15">The distribution range of the species covers Dak Lak and Lam Dong provinces, Vietnam.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D2C728DC6B06FD6F40A5EE5B8D7687D9" pageId="0" pageNumber="15" type="remarks">
|
||
<paragraph id="C5DD64117C8B18E974E9DC0633D3D394" pageId="0" pageNumber="15">Remarks.</paragraph>
|
||
<paragraph id="79F10556EAB43729AFECC096C5F75609" pageId="0" pageNumber="15">
|
||
<bibRefCitation id="0921AD7BA5CF8447BFF1564100666C3D" author="Thach, NN" journalOrPublisher="Basteria" pageId="0" pageNumber="15" pagination="35 - 37" publicationUrl="https://natuurtijdschriften.nl/pub/643896/BAST2014078001004.pdf" refId="B78" refString="Thach, NN, Huber, F, 2014. A new Amphidromus (Gastropoda: Camaenidae) from Vietnam. Basteria 78: 35 - 37, https://natuurtijdschriften.nl/pub/643896/BAST2014078001004.pdf" title="A new Amphidromus (Gastropoda: Camaenidae) from Vietnam." url="https://natuurtijdschriften.nl/pub/643896/BAST2014078001004.pdf" volume="78" year="2014">Thach and Huber (2014)</bibRefCitation>
|
||
introduced
|
||
<taxonomicName id="3C089103248E3FBE59030CABDEE5812E" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. naggsi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="CFA5109C2AAED164D9C625F4507E4581" italics="true" pageId="0" pageNumber="15">A. naggsi</emphasis>
|
||
</taxonomicName>
|
||
, which is described to differ from
|
||
<taxonomicName id="4D50E98C58229E68AF388BBF8C7DC63A" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="FFE2F3947A86F3D8B289D55106323700" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
in its wrinkled outer surface, the presence of 2-3 broad spiral channels on the body whorl, the more prominent sculpture on the penultimate whorl, and a more elongate aperture. However, upon examining the type specimens of both
|
||
<taxonomicName id="BF444AF8CE15A80DE68BD708B5EA7DC4" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="9C9961959249653817784CEBCEE5E4B2" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2EF7615030777F757D9A4D249294D9C6" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. naggsi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="384E6C0DC649D96B714FD150C7C625D7" italics="true" pageId="0" pageNumber="15">A. naggsi</emphasis>
|
||
</taxonomicName>
|
||
, these diagnostic characters were also present in the lectotype and paralectotypes of
|
||
<taxonomicName id="240DF207A3053AC5F4C538DC6278E88B" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="09B92D6C6B9066502AEBEF27CF21FCB1" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
, and the holotype of
|
||
<taxonomicName id="14D2C39F22868C38C16B496521ADCF05" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. naggsi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="079C4D947503F73578409C3E91A48798" italics="true" pageId="0" pageNumber="15">A. naggsi</emphasis>
|
||
</taxonomicName>
|
||
agrees well with all the type specimens of
|
||
<taxonomicName id="DB47A2543D2D4FD9C213F7B224E53827" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="5593033C498305517E4AA59460A28719" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
in terms of shell shape, shell surface sculpture, peristome, and apertural shape. Thus,
|
||
<taxonomicName id="1B3A01CBA3FE7E3EB3A358B91EF0793F" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. naggsi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="naggsi">
|
||
<emphasis id="81E91261531115415BFBEA615151F90D" italics="true" pageId="0" pageNumber="15">A. naggsi</emphasis>
|
||
</taxonomicName>
|
||
is regarded herein as a junior subjective synonym of
|
||
<taxonomicName id="1951BBB4152E04412792DCBE0F3CEB52" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="146647AAC6ED797E9433E421250D48B5" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EE24F0D726BBD9D858AEFB9D7A9C93D2" pageId="0" pageNumber="15">
|
||
The shell colour generally varies from whitish (typical) to yellowish to rose-pink colour (Fig.
|
||
<figureCitation id="83226B7C1F83FFBB7B177B9F995BF974" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Amphidromus ingens Moellendorff, 1900 A lectotype of " Amphidromus ingens " (SMF 7565) B paralectotype of " Amphidromus ingens " (SMF 7566) C holotype of " Amphidromus naggsi " (RMNH. 5003908) D, E specimens from M'drak, Dak Lak, Vietnam (NMNS- 8764 - 082, NMNS- 8764 - 084) F specimen from Ea M'doal, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 087) G specimen from Krong A, M'drak, Dak Lak, Vietnam (NMNS- 8764 - 088) H, I specimens from Ea Sup, Dak Lak, Vietnam (NMNS- 8764 - 093, NMNS- 8764 - 101). Credit: J. Goud, RMNH (C)." figureDoi="10.3897/zookeys.1196.112146.figure3" httpUri="https://binary.pensoft.net/fig/1007821" pageId="0" pageNumber="15">3</figureCitation>
|
||
). In our examined specimens, many are stained with dark brown colour below periphery and some are stained nearly entirely on the last and penultimate whorl. The shell sculpture generally has two depression areas, one upper periphery and one below suture, and the conspicuous keel to weak keel is generally present on periphery.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |