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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.66.8457" ID-PMC="PMC4957028" ID-Pensoft-Pub="1314-2003-66-1" ID-Pensoft-UUID="704E9F435B62FFD93345FFD55178FFE6" ID-PubMed="27489494" ID-Zenodo-Dep="128816" ModsDocID="1314-2003-66-1" checkinTime="1468449185517" checkinUser="pensoft" docAuthor="Knapp, Sandra &amp; Vorontsova, Maria S." docDate="2016" docId="41E31C342BCB555EB49B60D72E6A18CE" docLanguage="en" docName="PhytoKeys 66: 1-142" docOrigin="PhytoKeys 66" docPubDate="2016-07-13" docSource="http://dx.doi.org/10.3897/phytokeys.66.8457" docTitle="Solanum runsoriense C. H. Wright, Uganda Prot. 1: 362. 1902." docType="treatment" docVersion="3" id="704E9F435B62FFD93345FFD55178FFE6" lastPageNumber="67" masterDocId="704E9F435B62FFD93345FFD55178FFE6" masterDocTitle="A revision of the &quot; African Non-Spiny &quot; Clade of Solanum L. (Solanum sections Afrosolanum Bitter, Benderianum Bitter, Lemurisolanum Bitter, Lyciosolanum Bitter, Macronesiotes Bitter, and Quadrangulare Bitter: Solanaceae)" masterLastPageNumber="142" masterPageNumber="1" pageNumber="60" updateTime="1668141734746" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the &quot; African Non-Spiny &quot; Clade of Solanum L. (Solanum sections Afrosolanum Bitter, Benderianum Bitter, Lemurisolanum Bitter, Lyciosolanum Bitter, Macronesiotes Bitter, and Quadrangulare Bitter: Solanaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:affiliation>Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vorontsova, Maria S.</mods:namePart>
<mods:affiliation>Comparative Plant and Fungal Biology, Royal Botanic Gardens, Kew, Richmond, Surrey TW 7 3 AB, United Kingdom</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-07-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>142</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.66.8457</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.66.8457</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-66-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">704E9F435B62FFD93345FFD55178FFE6</mods:identifier>
<mods:identifier type="Zenodo-Dep">128816</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226982" LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE" httpUri="http://treatment.plazi.org/id/41E31C342BCB555EB49B60D72E6A18CE" lastPageId="66" lastPageNumber="67" pageId="59" pageNumber="60">
<subSubSection pageId="59" pageNumber="60" type="nomenclature">
<paragraph pageId="59" pageNumber="60">
<taxonomicName LSID="41E31C34-2BCB-555E-B49B-60D72E6A18CE" authority="C. H. Wright, Uganda Prot. 1: 362. 1902." authorityName="C. H. Wright, Uganda Prot. 1: 362. 1902." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="runsoriense">Solanum runsoriense C.H.Wright, Uganda Prot. 1: 362. 1902.</taxonomicName>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Solanum runsoriense C. H. Wright. A Flowering branch from short-styled plant B Bud from short-styled flower C Open short-styled flower D Flowering branch of long-styled plant E Open long-styled flower with long-exserted style F Infructescence G Detail of dense dendritic pubescence of leaf undersides H Elongate dendritic trichome. (Based on: A-C Napier 5130; D, E, G, H Friis et al. 3610; F Mooney 7016). Scale bar: A, D, F = 4 cm; B, C = 7 mm; E = 1.5 cm; G = 5 mm; H = 0.4 mm. Drawn by Lucy T. Smith." figureDoi="10.3897/phytokeys.66.8457.figure21" httpUri="https://binary.pensoft.net/fig/95594" pageId="59" pageNumber="60">Figure 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="reference_group">
<paragraph pageId="60" pageNumber="61">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="benderianum">
<pageBreakToken pageId="60" pageNumber="61" start="start">Solanum</pageBreakToken>
benderianum
</taxonomicName>
C.H.Wright, Fl. Trop. Afr. [Oliver et al.] 4(2): 212. 1906, as &quot;
<emphasis italics="true" pageId="60" pageNumber="61">bendirianum</emphasis>
&quot;. Type. Ethiopia. Oromia: Harwash and Maki Rivers [Shewa Region], Jan 1899,
<emphasis italics="true" pageId="60" pageNumber="61">M.S. Wellby s.n.</emphasis>
(lecctotype, designated here: K [K000788686]).
</paragraph>
<paragraph lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="benderianum">Solanum benderianum</taxonomicName>
Dammer, Bot. Jahrb. Syst. 38: 184. 1906 [
<normalizedToken originalValue="“1907”">&quot;1907&quot;</normalizedToken>
], nom. illeg., isonym, not
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="benderianum">Solanum benderianum</taxonomicName>
C.H.Wright, 1906.
<pageBreakToken pageId="61" pageNumber="62" start="start">Type</pageBreakToken>
. Ethiopia. Amhara:
<normalizedToken originalValue="“Kirchengehölzes">&quot;Kirchengehoelzes</normalizedToken>
Herroe Gottes Georgis bei Gaffat&quot; [South Gonder, near Debre Tabor], 8400 ft., 1 Oct 1863,
<emphasis italics="true" pageId="61" pageNumber="62">G.H.W. Schimper 1227</emphasis>
(lectotype, designated here: BM [BM000847516]; isolectotypes: BM [E [E00193238], K [K000414030, K000414031], US [US0027474= US-806522], W [W18990146870], WU [WU0033424, WU0033425]).
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum subsp. var. var. lanceolatum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="variety" species="benderianum" subSpecies="var." variety="lanceolatum">Solanum benderianum C.H.Wright var. lanceolatum</taxonomicName>
Bitter, Bot. Jahrb. Syst. 54: 488. 1917. Type. Ethiopia. Oromia: &quot;Gallahochland, Kiritscha, Utadara&quot;, Dec 1900,
<emphasis italics="true" pageId="61" pageNumber="62">O. Neumann 62</emphasis>
(B, destroyed, no duplicates found); Sidamo, Njam-Njam [Jem-Jem], Dec 1900,
<emphasis italics="true" pageId="61" pageNumber="62">H. Ellenbeck 1761</emphasis>
(no herbarium cited).
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum subsp. var. var. ruwenzoriense" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="variety" species="benderianum" subSpecies="var." variety="ruwenzoriense">Solanum benderianum C.H.Wright var. ruwenzoriense</taxonomicName>
Bitter, Bot. Jahrb. Syst. 54: 489. 1917. Type. Uganda. Western: Ruwenzori, Kivatu, 2500-2800 m, 1893-1894,
<emphasis italics="true" pageId="61" pageNumber="62">G.F. Scott-Elliot 7733</emphasis>
(lectotype, designated here: BM [BM000847515]; isolectotypes: K [K000414032]).
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum keniense" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="keniense">Solanum keniense</taxonomicName>
Standl., Smithson. Misc. Coll. 68, no. 5: 16. 1917, nom. illeg., non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum keniense" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="keniense">Solanum keniense</taxonomicName>
Turrill, 1915. Type. Kenya. Central: western slopes of Mount Kenya, along the trail from West Kenya Forest station to the summit, ca. 3630 m, 21-27 Sep 1917,
<emphasis italics="true" pageId="61" pageNumber="62">E.A. Mearns 1416</emphasis>
(holotype: US [US00027640]).
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="benderianum">Solanum benderianum</taxonomicName>
A.Schimp. ex Engl., Abh. Naturwiss. Vereine Bremen 25: 246. 1922. Type. Probably not intended as a new name; orthographic correction to
<bibRefCitation author="Wright, CH" journalOrPublisher="Flora of Tropical Africa" pageId="112" pageNumber="113" pagination="207 - 261" refId="B120" refString="Wright, CH, 1906. Solanaceae. Flora of Tropical Africa 4 (2): 207 - 261" title="Solanaceae." volume="4" year="1906">Wright (1906)</bibRefCitation>
.
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum longipedicellatum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="longipedicellatum">Solanum longipedicellatum</taxonomicName>
De Wild., Pl. Bequaert. 1: 428. 1922, nom. illeg., non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum longipedicellatum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="longipedicellatum">Solanum longipedicellatum</taxonomicName>
Bittter, 1912. Type. Democratic Republic of the Congo. Nord-Kivu: Ruwenzori, Lanuri [Valley], ca. 3000 m, 3 Jun 1914,
<emphasis italics="true" pageId="61" pageNumber="62">J. Bequaert 4676</emphasis>
(lectotype, designated here: BR [BR0000008993151]; isolectotypes: BR [BR0000008993120], LWI [LWI486608136]).
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Robyns" authorityYear="1947" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum dewildemanianum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="dewildemanianum">Solanum dewildemanianum</taxonomicName>
Robyns, Fl. Spermatophyt. Parc. Nat. Albert 2: 209. 1947. Type. Based on
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum longipedicellatum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="longipedicellatum">Solanum longipedicellatum</taxonomicName>
De Wild.
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Edmonds" authorityYear="2012" baseAuthorityName="C. H. Wright" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense subsp. benderianum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="subSpecies" species="runsoriense" subSpecies="benderianum">Solanum runsoriense C.H.Wright subsp. benderianum</taxonomicName>
(C.H.Wright) Edmonds, Fl. Trop. East Africa, Solan. 120. 2012, as &quot;
<emphasis italics="true" pageId="61" pageNumber="62">bendirianum</emphasis>
&quot;. Type. Based on
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="benderianum">Solanum benderianum</taxonomicName>
C.H.Wright
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="materials_examined">
<paragraph pageId="61" pageNumber="62">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="61" pageNumber="62">
<materialsCitation collectingDate="1901-03-13" collectorName="W. G. Doggett" country="Uganda" location="Western" municipality="Ruwenzori" specimenCount="1" typeStatus="holotype">
<collectingCountry name="Uganda">Uganda</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:E92A5F56003AEEA4605FA03012ACB2EC" country="Uganda" municipality="Ruwenzori" name="Western">Western</location>
:
<collectingMunicipality>Ruwenzori</collectingMunicipality>
, [rec. at Kew
<collectingDate value="1901-03-13">13 Mar 1901</collectingDate>
],
<emphasis italics="true" pageId="61" pageNumber="62">
<collectorName>W.G. Doggett</collectorName>
s.n.
</emphasis>
(
<typeStatus>holotype</typeStatus>
: K [K000413968])
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
<paragraph pageId="61" pageNumber="62">Description.</paragraph>
<paragraph lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62">
Vine or woody liana, sometimes semi-herbaceous, to 10 m. Stems flexuous, terete, glabrous to densely pubescent with mixed simple 4-6-celled and short-branched dendritic uniseriate trichomes 0.5-1(1.5) mm long, glabrescent when older; new growth almost glabrous to densely pubescent with dendritic and simple uniseriate trichomes, when these dense the new growth appearing golden in dry material. Bark of older stems reddish brown, smooth. Sympodial units plurifoliate, the leaves not geminate, evenly distributed along branches. Leaves simple, 4-12 cm long, 2-5.6
<pageBreakToken pageId="62" pageNumber="63" start="start">cm</pageBreakToken>
wide, elliptic or less commonly ovate, membraneous, the adaxial surfaces glabrous to moderately pubescent with mostly simple uniseriate trichomes 0.5-1 mm long, in more pubescent individuals some trichomes dendritic, the abaxial surfaces glabrous or with a few simple uniseriate trichomes along the veins and margins to densely pubescent with short-branched dendritic trichomes 0.5-1.5 mm long, these often drying golden; major veins 5-10 pairs, usually yellowish beneath, especially on glabrous individuals; base acute; margins entire, if the leaves otherwise glabrous the margins have a few simple trichomes to 1 mm long near the base; apex acute to acuminate; petioles 0.7-3.2 cm long, very variable in length along the stem, nearly glabrous to densely dendritic-pubescent, sometimes twining. Inflorescences terminal, 5-15(+) cm long, lax and open, many times branched, with 20-100 flowers many open at the same time, pubescence in parallel to that of stems and leaf undersides; peduncle).5-)2-4.5 cm long, sometimes the inflorescence branches starting very near the last stem leaves; pedicels 0.8-1.2 cm long, ca. 0.5 mm in diameter at the base, ca. 1 mm in diameter at the apex, spreading at anthesis, pubescent like the rest of the inflorescence axis, but slightly less densely, articulated at the base leaving a very minute raised portion of the axis; pedicel scars irregularly spaced 1-6 mm apart. Buds ellipsoid, the corolla completely enclosed in the calyx tube until shortly before anthesis, when young the elongate sepals spreading at the bud tips. Flowers 5-merous, heterostylous, short-styled and long-styled flowers apparently borne on different plants and the plants possibly dioecious. Calyx tube 1-1.5 mm long, open-conical, the lobes 1-2 mm long, narrowly triangular with a long-acuminate tip, usually thickened and keeled abaxially (this easier to see on glabrous specimens), glabrous to densely pubescent with mixed dendritic and simple uniseriate trichomes, even if glabrous the tips with a tuft of simple uniseriate trichomes. Corolla (1.5)2-3.5 cm in diameter, pale violet or white with a darker purple centre, stellate, lobed ca. halfway to the base, the lobes 7-9 mm long, 6-8 mm wide, spreading at anthesis, variably pubescent, from almost glabrous with dense papillae on the tips and margins to densely pubescent with minute dendritic trichomes all usually &lt;0.5 mm long (to 1 mm long in
<emphasis italics="true" pageId="62" pageNumber="63">Taylor 1408</emphasis>
). Stamens equal; filament tube absent; free portion of the filaments 1.5-2.5 mm long, glabrous or sparsely pubsvent with dendritic trichomes (on pubescent plants); anthers 2.5-3 mm long, 1.1.5 mm wide, ellipsoid, bright yellow, connivent to somewhat spreading, smooth abaxially, somewhat sagittate at the base, poricidal at the tips, the pores lengthening to slits with age, in dry material the pores thickened and paler at the distal tips. Ovary globose, glabrous; style of different lengths on individual plants, the short-styled plants with styles 2.5-3 mm long, held within the anther cone or exceptionally to 5 mm long and at the level of the anther tips, the long-styled plants with styles 7-9 mm long, exserted 4-6 mm beyond the anther cone, glabrous (on
<emphasis italics="true" pageId="62" pageNumber="63">Wollaston s.n.</emphasis>
[SS plant from Ruwenzori] densely dendritic-pubescent in the distal half); stigma of short styles minutely capitate, of long styles strongly bilobed or clavate, the surfaces minutely papillate. Fruit a globose berry, 0.8-0.9 cm in diameter, purple when ripe, the pericarp thin and brittle in dry material, probably shiny in fresh material; fruiting pedicels 2-2.5 cm long, tapering markedly from a base ca. 1 mm in diameter to the apex 4-5 mm in diameter, the distal half usually darker in dry material, spreading, possibly
<pageBreakToken pageId="63" pageNumber="64" start="start">somewhat</pageBreakToken>
woody; fruiting calyx lobes to 0.5 cm long, spreading. Seeds 10-12(16) per berry, 2.5-3.5 mm long, 2-2.5 mm wide, ovoid reniform, reddish brown, the surfaces shallowly pitted, the testal cells very small, pentagonal in outline.
</paragraph>
<caption ID-Zenodo-Dep="980371" doi="10.3897/phytokeys.66.8457.figure21" httpUri="https://binary.pensoft.net/fig/95594" pageId="63" pageNumber="64" start="Figure 21" startId="F21">
<paragraph pageId="63" pageNumber="64">
<emphasis bold="true" pageId="63" pageNumber="64">Figure 21.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum runsoriense</emphasis>
</taxonomicName>
C.H.Wright.
<emphasis bold="true" pageId="63" pageNumber="64">A</emphasis>
Flowering branch from short-styled plant
<emphasis bold="true" pageId="63" pageNumber="64">B</emphasis>
Bud from short-styled flower
<emphasis bold="true" pageId="63" pageNumber="64">C</emphasis>
Open short-styled flower
<emphasis bold="true" pageId="63" pageNumber="64">D</emphasis>
Flowering branch of long-styled plant
<emphasis bold="true" pageId="63" pageNumber="64">E</emphasis>
Open long-styled flower with long-exserted style
<emphasis bold="true" pageId="63" pageNumber="64">F</emphasis>
Infructescence
<emphasis bold="true" pageId="63" pageNumber="64">G</emphasis>
Detail of dense dendritic pubescence of leaf undersides
<emphasis bold="true" pageId="63" pageNumber="64">H</emphasis>
Elongate dendritic trichome. (Based on:
<emphasis bold="true" pageId="63" pageNumber="64">A-C</emphasis>
<emphasis italics="true" pageId="63" pageNumber="64">Napier 5130</emphasis>
;
<emphasis bold="true" pageId="63" pageNumber="64">D, E, G, H</emphasis>
<emphasis italics="true" pageId="63" pageNumber="64">Friis et al. 3610</emphasis>
;
<emphasis bold="true" pageId="63" pageNumber="64">F</emphasis>
<emphasis italics="true" pageId="63" pageNumber="64">Mooney 7016</emphasis>
). Scale bar:
<emphasis bold="true" pageId="63" pageNumber="64">A, D, F</emphasis>
= 4 cm;
<emphasis bold="true" pageId="63" pageNumber="64">B, C</emphasis>
= 7 mm;
<emphasis bold="true" pageId="63" pageNumber="64">E</emphasis>
= 1.5 cm;
<emphasis bold="true" pageId="63" pageNumber="64">G</emphasis>
= 5 mm;
<emphasis bold="true" pageId="63" pageNumber="64">H</emphasis>
= 0.4 mm. Drawn by Lucy T. Smith.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="distribution">
<paragraph pageId="63" pageNumber="64">Distribution</paragraph>
<paragraph pageId="63" pageNumber="64">
(Figure
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution of Solanum runsoriense C. H. Wright." figureDoi="10.3897/phytokeys.66.8457.figure22" httpUri="https://binary.pensoft.net/fig/95595" pageId="63" pageNumber="64">22</figureCitation>
). In the mountains of eastern Africa in Ethiopia, Kenya, Uganda, and the Democratic Republic of the Congo.
</paragraph>
<caption ID-Zenodo-Dep="980373" doi="10.3897/phytokeys.66.8457.figure22" httpUri="https://binary.pensoft.net/fig/95595" pageId="63" pageNumber="64" start="Figure 22" startId="F22">
<paragraph pageId="63" pageNumber="64">
<emphasis bold="true" pageId="63" pageNumber="64">Figure 22.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum runsoriense</emphasis>
</taxonomicName>
C.H.Wright.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="common names and uses">
<paragraph pageId="63" pageNumber="64">Common names and uses.</paragraph>
<paragraph pageId="63" pageNumber="64">None recorded.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="ecology">
<paragraph pageId="63" pageNumber="64">Ecology and habitat.</paragraph>
<paragraph pageId="63" pageNumber="64">Montane forests, bamboo forests; often in open areas and forest edges; 1500-3700 m elevation.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="preliminary conservation status">
<paragraph pageId="63" pageNumber="64">Preliminary conservation status</paragraph>
<paragraph pageId="63" pageNumber="64">
(
<bibRefCitation author="IUCN" journalOrPublisher="Annales de Biologie" pageId="109" pageNumber="110" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B56" refString="IUCN, 2014. Guidelines for Using the IUCN Red List Categories and Criteria. Version 11., http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 11" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2014">IUCN 2014</bibRefCitation>
). Least Concern (LC). EOO 1,346,193 km2 (LC), AOO 200 km2 (EN). Although
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum runsoriense</emphasis>
</taxonomicName>
as delimited here occupies a large geographical range, it is found at high elevations on isolated mountain systems. Genetic differentiation at the population level across its range could therefore be important for conservation and should be assessed.
</paragraph>
</subSubSection>
<subSubSection lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64" type="discussion">
<paragraph pageId="63" pageNumber="64">Discussion.</paragraph>
<paragraph lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum runsoriense</emphasis>
</taxonomicName>
is a montane forest edge species with large, openly branching inflorescences of purple (
<normalizedToken originalValue="“blue”">&quot;blue&quot;</normalizedToken>
) flowers. It is not likely to be confused with any other species on the African continent; it the only species of the ANS clade with short-styled and long-styled flowers (see below, Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Solanum runsoriense C. H. Wright. A Flowering branch from short-styled plant B Bud from short-styled flower C Open short-styled flower D Flowering branch of long-styled plant E Open long-styled flower with long-exserted style F Infructescence G Detail of dense dendritic pubescence of leaf undersides H Elongate dendritic trichome. (Based on: A-C Napier 5130; D, E, G, H Friis et al. 3610; F Mooney 7016). Scale bar: A, D, F = 4 cm; B, C = 7 mm; E = 1.5 cm; G = 5 mm; H = 0.4 mm. Drawn by Lucy T. Smith." figureDoi="10.3897/phytokeys.66.8457.figure21" httpUri="https://binary.pensoft.net/fig/95594" pageId="63" pageNumber="64">21</figureCitation>
).
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum runsoriense</emphasis>
</taxonomicName>
is partly sympatric with
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum terminale" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="terminale">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum terminale</emphasis>
</taxonomicName>
but occurs at higher elevations and apparently in wetter forests.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum terminale" order="Solanales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="terminale">
<emphasis italics="true" pageId="63" pageNumber="64">Solanum terminale</emphasis>
</taxonomicName>
has more deeply stellate corollas, flowers tightly clustered in groups rather than evenly and widely spaced on the inflorescence branches, and bright
<pageBreakToken pageId="64" pageNumber="65" start="start">red</pageBreakToken>
rather than blackish purple berries. Pubescence is extremely variable in
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
; populations from Ethiopia (described as
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="benderianum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum benderianum</emphasis>
</taxonomicName>
) are almost completely glabrous on stems and leaves, while populations from the Ruwenzoris in Democratic Republic of the Congo and from the Aberdare Mountains in Kenya are densely pubescent with golden dendritic trichomes. Individual populations are isolated in what has been recognised as the Afromontane (Wright 1978, 1981) or Tropic-montane Flora (
<bibRefCitation author="Linder, HP" journalOrPublisher="Frontiers in Ecology and Evolution" pageId="110" pageNumber="111" pagination="1 - 14" publicationUrl="10.3389/fevo.2014.00038" refId="B71" refString="Linder, HP, 2014. The evolution of African plant diversity. Frontiers in Ecology and Evolution 2 (38): 1 - 14, DOI: 10.3389/fevo.2014.00038" title="The evolution of African plant diversity." url="10.3389/fevo.2014.00038" volume="2" year="2014">Linder 2014</bibRefCitation>
) and limited gene exchange may allow these variants to become fixed. Flower, fruit and seed morphology are constant throughout the species range.
</paragraph>
<paragraph pageId="64" pageNumber="65">
<bibRefCitation author="Edmonds, JM" editor="Beentje, H" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="108" pageNumber="109" pagination="105 - 125" refId="B44" refString="Edmonds, JM, 2012. Solanum spp. In: Beentje, H, Ed., Flora of Tropical East Africa Solanaceae. Royal Botanic Gardens, Kew: 105 - 125" title="Solanum spp." volumeTitle="Flora of Tropical East Africa Solanaceae." year="2012">Edmonds (2012)</bibRefCitation>
placed
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum macrothyrsum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="macrothyrsum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum macrothyrsum</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
; she cited
<bibRefCitation author="Dammer, U" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="107" pageNumber="108" pagination="176 - 196" refId="B26" refString="Dammer, U, 1906 1907. Solanaceae africanae I. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 38: 176 - 196" title="Solanaceae africanae I." volume="38" year="1906">Dammer (1906)</bibRefCitation>
and
<bibRefCitation author="Jaeger, PM-L" journalOrPublisher="University of Birmingham, Birmingham, United Kingdom" pageId="109" pageNumber="110" refId="B59" refString="Jaeger, PM-L, 1985. Systematic studies in the genus Solanum in Africa. University of Birmingham, Birmingham, United Kingdom" title="Systematic studies in the genus Solanum in Africa" year="1985">Jaeger (1985)</bibRefCitation>
both of whom suggested there were affinities between the two taxa. The two taxa share openly branched inflorescences and
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum macrothyrsum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="macrothyrsum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum macrothyrsum</emphasis>
</taxonomicName>
is superfically similar vegetatively to more glabrous forms of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
, but
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum macrothyrsum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="macrothyrsum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum macrothyrsum</emphasis>
</taxonomicName>
has deeply stellate corollas and anthers that open with two terminal pores that never extend to slits as the flower ages, while
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
has shallowly stellate corollas and anther pores that extend to longitudinal slits with age.
</paragraph>
<paragraph pageId="64" pageNumber="65">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
has the only member of the ANS clade that is heterostylous; in all but one of the specimens we have examined flowers on a given stem are either short-styled or long-styled. The Ethiopian collection
<emphasis italics="true" pageId="64" pageNumber="65">Gilbert 138</emphasis>
(K000788688) is the only one we have seen with fruit and short-styled flowers on same plant. No other inflorescences with short-styled flowers had fruits, and when specimens were fruiting, all flowers in the inflorescence had set fruit. Collections are often of some sheets with flowers (short-styled) and duplicates with fruit (e.g.,
<emphasis italics="true" pageId="64" pageNumber="65">Schimper 1227</emphasis>
); it is not clear if these represent branches from a single plant or different individuals.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
could be either monoecious or dioecious, but most monoecious solanums have a mixture of short- and long-styled flowers in the same inflorescence (
<bibRefCitation author="Diggle, P" journalOrPublisher="American Journal of Botany" pageId="108" pageNumber="109" pagination="20 - 30" publicationUrl="10.3732/ajb.91.12.2030" refId="B37" refString="Diggle, P, Miller, J, 2004. Architectual effects mimic floral sexual dimorphism in Solanum (Solanaeae). American Journal of Botany 91: 20 - 30, DOI: 10.3732/ajb.91.12.2030" title="Architectual effects mimic floral sexual dimorphism in Solanum (Solanaeae)." url="10.3732/ajb.91.12.2030" volume="91" year="2004">Diggle and Miller 2004</bibRefCitation>
;
<bibRefCitation author="Miller, J" journalOrPublisher="American Journal of Botany" pageId="110" pageNumber="111" pagination="707 - 715" publicationUrl="10.3732/ajb.90.5.707" refId="B83" refString="Miller, J, Diggle, P, 2003. Diversification of andromonoecy in Solanum section Lasiocarpa (Solanaceae): the roles of phenotypic plascticity and architecture. American Journal of Botany 90: 707 - 715, DOI: 10.3732/ajb.90.5.707" title="Diversification of andromonoecy in Solanum section Lasiocarpa (Solanaceae): the roles of phenotypic plascticity and architecture." url="10.3732/ajb.90.5.707" volume="90" year="2003">Miller and Diggle 2003</bibRefCitation>
;
<bibRefCitation author="Whalen, MD" editor="D'Arcy, WG" journalOrPublisher="Columbia University Press, New York" pageId="112" pageNumber="113" pagination="284 - 302" refId="B112" refString="Whalen, MD, Costich, DE, 1986. Andromonoecy in Solanum. In: D'Arcy, WG, Ed., Solanaceae: biology and systematics. Columbia University Press, New York: 284 - 302" title="Andromonoecy in Solanum." volumeTitle="Solanaceae: biology and systematics." year="1986">Whalen and Costich 1986</bibRefCitation>
) so we suspect
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
is another instance of dioecy in the genus (see
<bibRefCitation author="Symon, DE" journalOrPublisher="Taxon" pageId="111" pageNumber="112" pagination="909 - 910" publicationUrl="10.2307/1218308" refId="B100" refString="Symon, DE, 1970. Dioecious Solanum. Taxon 19: 909 - 910, DOI: 10.2307/1218308" title="Dioecious Solanum." url="10.2307/1218308" volume="19" year="1970">Symon 1970</bibRefCitation>
;
<bibRefCitation author="Symon, DE" editor="Hawkes, JD" journalOrPublisher="Academic Press, London" pageId="111" pageNumber="112" pagination="385 - 397" refId="B101" refString="Symon, DE, 1979. Sex forms in Solanum (Solanaceae) and the role of pollen collecting insects. In: Hawkes, JD, Lester, RN, Skelding, AD, Eds., The biology and taxonomy of the Solanaceae. Academic Press, London: 385 - 397" title="Sex forms in Solanum (Solanaceae) and the role of pollen collecting insects." volumeTitle="The biology and taxonomy of the Solanaceae." year="1979">Symon 1979</bibRefCitation>
;
<bibRefCitation author="Anderson, GJ" journalOrPublisher="Evolution" pageId="106" pageNumber="107" pagination="204 - 219" publicationUrl="10.2307/2409175" refId="B2" refString="Anderson, GJ, Symon, DE, 1989. Functional dioecy and andromonoecy in Solanum. Evolution 43: 204 - 219, DOI: 10.2307/2409175" title="Functional dioecy and andromonoecy in Solanum." url="10.2307/2409175" volume="43" year="1989">Anderson and Symon 1989</bibRefCitation>
;
<bibRefCitation author="Knapp, S" journalOrPublisher="Plant Systematics and Evolution" pageId="109" pageNumber="110" pagination="113 - 139" publicationUrl="10.1007/BF00984731" refId="B65" refString="Knapp, S, Persson, V, Blackmore, S, 1998. Pollen morphology and functional dioecy in Solanum (Solanaceae). Plant Systematics and Evolution 210: 113 - 139, DOI: 10.1007/BF00984731" title="Pollen morphology and functional dioecy in Solanum (Solanaceae)." url="10.1007/BF00984731" volume="210" year="1998">Knapp et al. 1998</bibRefCitation>
;
<bibRefCitation author="Martine, CT" journalOrPublisher="Australian Systematic Botany" pageId="110" pageNumber="111" pagination="107 - 120" publicationUrl="10.1071/SB07039" refId="B79" refString="Martine, CT, Anderson, GJ, Les, DH, 2009. Gender-bending aubergines: Molecular phylogenetics of cryptically dioecious Solanum in Australia. Australian Systematic Botany 22: 107 - 120, DOI: 10.1071/SB07039" title="Gender-bending aubergines: Molecular phylogenetics of cryptically dioecious Solanum in Australia." url="10.1071/SB07039" volume="22" year="2009">Martine et al. 2009</bibRefCitation>
). Field observations on the breeding system of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum runsoriense</emphasis>
</taxonomicName>
are a priority, and microscopic examination of pollen morphology to see if long-styled flowers have inaperurate pollen like other dioecious solanums (e.g.,
<bibRefCitation author="Knapp, S" journalOrPublisher="Plant Systematics and Evolution" pageId="109" pageNumber="110" pagination="113 - 139" publicationUrl="10.1007/BF00984731" refId="B65" refString="Knapp, S, Persson, V, Blackmore, S, 1998. Pollen morphology and functional dioecy in Solanum (Solanaceae). Plant Systematics and Evolution 210: 113 - 139, DOI: 10.1007/BF00984731" title="Pollen morphology and functional dioecy in Solanum (Solanaceae)." url="10.1007/BF00984731" volume="210" year="1998">Knapp et al. 1998</bibRefCitation>
) might help to determine the breeding system of this species.
</paragraph>
<paragraph lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<bibRefCitation author="Engler, A" journalOrPublisher="Abhandlungen der Koniglichen Akademie der Wissenschaften in Berlin" pageId="108" pageNumber="109" pagination="1 - 461" refId="B45" refString="Engler, A, 1892. Ueber die Hochgebirgsflora des tropisches Africa. Abhandlungen der Koniglichen Akademie der Wissenschaften in Berlin 1891: 1 - 461" title="Ueber die Hochgebirgsflora des tropisches Africa." volume="1891" year="1892">Engler (1892)</bibRefCitation>
was the first to publish the name
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="benderianum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum benderianum</emphasis>
</taxonomicName>
, but without description; he attributed the name to Schimper and cited
<normalizedToken originalValue="Schimpers">Schimper's</normalizedToken>
1863 manuscript list of plants collected in Ethiopia.
<bibRefCitation author="Wright, CH" journalOrPublisher="Flora of Tropical Africa" pageId="112" pageNumber="113" pagination="207 - 261" refId="B120" refString="Wright, CH, 1906. Solanaceae. Flora of Tropical Africa 4 (2): 207 - 261" title="Solanaceae." volume="4" year="1906">Wright (1906)</bibRefCitation>
and
<bibRefCitation author="Dammer, U" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="107" pageNumber="108" pagination="176 - 196" refId="B26" refString="Dammer, U, 1906 1907. Solanaceae africanae I. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 38: 176 - 196" title="Solanaceae africanae I." volume="38" year="1906">Dammer (1906)</bibRefCitation>
both used
<normalizedToken originalValue="Englers">Engler's</normalizedToken>
epithet and provided descriptions, but
<normalizedToken originalValue="Wrights">Wright's</normalizedToken>
treatment was published in February 1906, a few months before
<normalizedToken originalValue="Dammers">Dammer's</normalizedToken>
that appeared in August 1906 making
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum benderianum" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="benderianum">
<emphasis italics="true" pageId="64" pageNumber="65">Solanum benderianum</emphasis>
</taxonomicName>
Dammer an isonym and illegitimate. Both botanists attributed the name to Schimper, cited
<bibRefCitation author="Engler, A" journalOrPublisher="Abhandlungen der Koniglichen Akademie der Wissenschaften in Berlin" pageId="108" pageNumber="109" pagination="1 - 461" refId="B45" refString="Engler, A, 1892. Ueber die Hochgebirgsflora des tropisches Africa. Abhandlungen der Koniglichen Akademie der Wissenschaften in Berlin 1891: 1 - 461" title="Ueber die Hochgebirgsflora des tropisches Africa." volume="1891" year="1892">
<normalizedToken originalValue="Englers">Engler's</normalizedToken>
(1892)
</bibRefCitation>
publication and based their descriptions on two collections (presumably different duplicates),
<emphasis italics="true" pageId="64" pageNumber="65">Schimper 1227</emphasis>
and
<emphasis italics="true" pageId="64" pageNumber="65">Scott-Elliot 7733</emphasis>
.
<bibRefCitation author="Wright, CH" journalOrPublisher="Flora of Tropical Africa" pageId="112" pageNumber="113" pagination="207 - 261" refId="B120" refString="Wright, CH, 1906. Solanaceae. Flora of Tropical Africa 4 (2): 207 - 261" title="Solanaceae." volume="4" year="1906">Wright (1906)</bibRefCitation>
also cited
<emphasis italics="true" pageId="64" pageNumber="65">Wellby s.n.</emphasis>
from Ethiopia.
<bibRefCitation author="Bitter, G" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="106" pageNumber="107" pagination="416 - 506" refId="B5" refString="Bitter, G, 1917. Solana Africana II. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 54: 416 - 506" title="Solana Africana II." volume="54" year="1917">Bitter (1917)</bibRefCitation>
later used
<emphasis italics="true" pageId="64" pageNumber="65">Scott-Elliot 7733</emphasis>
as the type of his var.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. ruwenzoriense" order="Solanales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="variety" species="benderianum" variety="ruwenzoriense">
<emphasis italics="true" pageId="64" pageNumber="65">ruwenzoriense</emphasis>
</taxonomicName>
. We have lectotypified these three names with the three different specimens used in various combinations in their protologues
<pageBreakToken pageId="65" pageNumber="66" start="start">in</pageBreakToken>
order to avoid making them homotypic.
<bibRefCitation author="Edmonds, JM" editor="Beentje, H" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="108" pageNumber="109" pagination="105 - 125" refId="B44" refString="Edmonds, JM, 2012. Solanum spp. In: Beentje, H, Ed., Flora of Tropical East Africa Solanaceae. Royal Botanic Gardens, Kew: 105 - 125" title="Solanum spp." volumeTitle="Flora of Tropical East Africa Solanaceae." year="2012">Edmonds (2012)</bibRefCitation>
incorrectly used
<bibRefCitation author="Wright, CH" journalOrPublisher="Flora of Tropical Africa" pageId="112" pageNumber="113" pagination="207 - 261" refId="B120" refString="Wright, CH, 1906. Solanaceae. Flora of Tropical Africa 4 (2): 207 - 261" title="Solanaceae." volume="4" year="1906">
<normalizedToken originalValue="Wrights">Wright's</normalizedToken>
(1906)
</bibRefCitation>
mis-spelling of the name &quot;
<emphasis italics="true" pageId="65" pageNumber="66">bendirianum</emphasis>
&quot;; we consider this a correctable spelling mistake because Wright cited
<normalizedToken originalValue="Englers">Engler's</normalizedToken>
publication in which the name was spelled in the way Schimper intended, to honour his son-in-law &quot;Herr Bender&quot;.
</paragraph>
<paragraph pageId="65" pageNumber="66">
<bibRefCitation author="De Wildeman, E" journalOrPublisher="A. Buyens, Gand &amp; J. Le Chevalier, Paris" pageId="108" pageNumber="109" pagination="299 - 452" refId="B34" refString="De Wildeman, E, 1922. Plantae Bequaertianae. A. Buyens, Gand &amp; J. Le Chevalier, Paris: 299 - 452" title="Plantae Bequaertianae" year="1922">De Wildeman (1922)</bibRefCitation>
specified no herbarium or sheet in his description of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum longipedicellatum" order="Solanales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="longipedicellatum">
<emphasis italics="true" pageId="65" pageNumber="66">Solanum longipedicellatum</emphasis>
</taxonomicName>
(a later homonym of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum longipedicellatum" order="Solanales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="longipedicellatum">
<emphasis italics="true" pageId="65" pageNumber="66">Solanum longipedicellatum</emphasis>
</taxonomicName>
Bitter, a synonym of the potato species
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum stoloniferum" order="Solanales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="stoloniferum">
<emphasis italics="true" pageId="65" pageNumber="66">Solanum stoloniferum</emphasis>
</taxonomicName>
Schltdl.,
<bibRefCitation author="Spooner, DE" journalOrPublisher="Systematic Botany Monographs" pageId="111" pageNumber="112" pagination="1 - 209" publicationUrl="10.2307/25027915" refId="B98" refString="Spooner, DE, van den Berg, RG, Rodriguez, A, Bamberg, R, Hijmans, RJ, Lara-Cabrera, SI, 2004. Wild potatoes (Solanum section Petota; Solanaceae) of North and Central America. Systematic Botany Monographs 68: 1 - 209, DOI: 10.2307/25027915" title="Wild potatoes (Solanum section Petota; Solanaceae) of North and Central America." url="10.2307/25027915" volume="68" year="2004">Spooner et al. 2004</bibRefCitation>
). Of the two sheets of
<emphasis italics="true" pageId="65" pageNumber="66">Bequaert 4676</emphasis>
at BR, we have chosen that (BR0000008993151) annotated incorrectly as
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
by the late R.N. Lester as the lectotype because it has a label with the number and collecting locality. The isolectotype at BR (BR0000008993120) has no original label, but has a scrawled sheet in De
<normalizedToken originalValue="Wildemans">Wildeman's</normalizedToken>
handwriting stating the differences between his species and
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum runsoriense" order="Solanales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="runsoriense">
<emphasis italics="true" pageId="65" pageNumber="66">Solanum runsoriense</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="materials_examined">
<paragraph pageId="65" pageNumber="66">Selected specimens examined.</paragraph>
<paragraph pageId="65" pageNumber="66">
<materialsCitation collectingDate="1948-05-12" collectingDateMax="1952-08-03" collectingDateMin="1948-05-12" collectorName="de Wilde" country="Democratic Republic of the Congo" location="Upper Ruamoli valley" specimenCount="1" stateProvince="Nord-Kivu">
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingCountry name="Democratic Republic of the Congo">Democratic Republic of the Congo</collectingCountry>
</emphasis>
.
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingRegion country="Democratic Republic of the Congo" name="Nord-Kivu">Nord-Kivu</collectingRegion>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:8B70CB0DFFD35A1A4703597DEDDC38AD" country="Democratic Republic of the Congo" name="Upper Ruamoli valley" stateProvince="Nord-Kivu">Upper Ruamoli valley</location>
,
<collectingDate value="1952-08-03">3 Aug 1952</collectingDate>
,
<emphasis italics="true" pageId="65" pageNumber="66">Ross 800</emphasis>
(BM); Katwah Kitumo, pistes des Beulele et a Mahonge (P.N.A. [Parc Nacional Albert = Virunga]),
<collectingDate value="1948-05-12">12 May 1948</collectingDate>
,
<emphasis italics="true" pageId="65" pageNumber="66">
<collectorName>de Wilde</collectorName>
55
</emphasis>
(BR)
</materialsCitation>
.
</paragraph>
<paragraph pageId="65" pageNumber="66">
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingCountry name="Ethiopia">Ethiopia</collectingCountry>
</emphasis>
.
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingRegion country="Ethiopia" name="Adis Abeba">Addis Ababa</collectingRegion>
</emphasis>
: Addis Ababa, 1956,
<emphasis italics="true" pageId="65" pageNumber="66">Mooney 6712</emphasis>
(K).
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingRegion country="Ethiopia" name="Amara">Amhara</collectingRegion>
</emphasis>
: Semien, Uulkefit, Lumalumo,
<date value="1909-07-10">10 Jul 1909</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Chiovenda 877</emphasis>
(FT); West Shewa region W of-Washa Forest,
<date value="1984-12-03">3 Dec 1984</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Edwards &amp; Tewolde 3572</emphasis>
(ETH); Choke Mountains, Gojjam, vicinity of the upper Ghiedeb valley,
<date value="1957-08-05">5 Aug 1957</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Evans 413</emphasis>
(BM, ETH, FT, K); Debre Workto Mota,
<date value="1981-10-30">30 Oct 1981</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Mesfin Tadese</emphasis>
&amp;
<emphasis italics="true" pageId="65" pageNumber="66">Kagnew 1684</emphasis>
(ETH, K); Mussolini Pass, between Debre Berhan and Dobre Sina, about
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.0" unit="km" value="200.0">200 km</quantity>
NNE of Addis Ababa,
<date value="1965-07-23">23 Jul 1965</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">de Wilde &amp; de Wilde-Duyfjes 7369</emphasis>
(BR).
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingRegion country="Ethiopia" name="Oromiya">Oromia</collectingRegion>
</emphasis>
: Menegasha State Forest, Mount Wuchacha, 48 kms due W of Addis Ababa on Ambo road, turn off just W of Menengasha village,
<date value="1971-10-21">21 Oct 1971</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Ash 1299</emphasis>
(EA, K); Ghidami,
<date value="1939-03">Mar 1939</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Benedetto 390</emphasis>
(FT); Bale Zone, Dolo Mena (Masslo) to Goba,
<date value="1984-10-29">29 Oct 1984</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Friis et al. 3610</emphasis>
(ETH, K); Mount Chilalo, a few kilometres S of mountain, track leading from road South of Asella to Ticho,
<date value="1968-11-25">25 Nov 1968</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Gilbert &amp; Gilbert 1098</emphasis>
(EA, K); Ticcio track; Arsi region,
<date value="1966-11-28">28 Nov 1966</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Gilbert 138</emphasis>
(ETH, K); Mt. Zuquala,
<date value="1994-11">Nov 1994</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Hylander, K. 139</emphasis>
(ETH); Mount Maigudo,
<date value="1954-10-24">24 Oct 1954</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Mooney 6156</emphasis>
(EA, FT, K, S); Debre Berhan, Wofasha,
<date value="1955-03-23">23 Mar 1955</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Mooney 6475</emphasis>
(FT, K); ca.
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
N of Koffale, Gobe Livestock Farm,
<date value="1971-10-13">13 Oct 1971</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Thulin 1502</emphasis>
(ETH, K); Mount Wuchacha, about
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.5" unit="km" value="15.0">15 km</quantity>
W of Addis Ababa,
<date value="1965-10-27">27 Oct 1965</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">de Wilde &amp; de Wilde-Duyfjes 8480</emphasis>
(BR, ETH, K); Arsi, Mount Borulucciu, Asella to Ticcio, E slope of Mount Borulucciu,
<date value="1965-12-06">6 Dec 1965</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">de Wilde et al. 9213</emphasis>
(BR, K).
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingRegion country="Ethiopia" name="Ye Debub Biheroch Bihereseboch na Hizboch">Southern Nations</collectingRegion>
,
<collectingRegion country="Ethiopia" name="Nationalities">Nationalities</collectingRegion>
and Peoples Region
</emphasis>
: Amaro Mountains, Mount Delo, E slope,
<date value="1953-01-29">29 Jan 1953</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Gillett 15034</emphasis>
(K); road from Wondo to Agere Selam [Hagere Selam],
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
from Wondo,
<date value="1968-01-24">24 Jan 1968</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">Westphal &amp; Westphal 3157</emphasis>
(BR); Agere Selam [Hagere Selam], about
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
S of Wondo,
<date value="1965-10-21">21 Oct 1965</date>
,
<emphasis italics="true" pageId="65" pageNumber="66">de Wilde &amp; de Wilde-Duyfjes 8360</emphasis>
(BR, ETH).
</paragraph>
<paragraph lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<materialsCitation collectingDate="1932-10" collectingDateMax="1975-01-12" collectingDateMin="1932-10" country="Kenya" county="Aberdare National Park" location="Aberdare National Park" municipality="Uplands District" specimenCount="1" stateProvince="Nyandarua">
<emphasis bold="true" pageId="65" pageNumber="66">
<collectingCountry name="Kenya">Kenya</collectingCountry>
</emphasis>
.
<emphasis bold="true" pageId="65" pageNumber="66">Central</emphasis>
:
<collectingCounty>Aberdare National Park</collectingCounty>
, Aberdare Mtns., ca.
<locationDeviation location="Jerusalem Gate">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
W of Jerusalem Gate
</locationDeviation>
(West), Dist.
<collectingRegion country="Kenya" name="Nyandarua">Nyandarua</collectingRegion>
,
<collectingDate value="1975-01-12">12 Jan 1975</collectingDate>
,
<emphasis italics="true" pageId="65" pageNumber="66">Croat 28375</emphasis>
(K, MO); Katamayu,
<collectingMunicipality>Uplands District</collectingMunicipality>
,
<collectingDate value="1933-08">Aug 1933</collectingDate>
,
<emphasis italics="true" pageId="65" pageNumber="66">Napier 5130</emphasis>
(K, MA);
<normalizedToken originalValue="Muranga">
<collectingRegion country="Kenya" name="Murang'a">Murang'a</collectingRegion>
</normalizedToken>
, Tuso Fishing Camp,
<collectingDate value="1932-10">
<pageBreakToken pageId="66" pageNumber="67" start="start">Oct</pageBreakToken>
1932
</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Jex-Blake 3304</emphasis>
(EA);
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:1284C36BF7F0ED72C9152D5D9FC88B4A" country="Kenya" county="Aberdare National Park" municipality="Uplands District" name="Aberdare National Park" stateProvince="Nyandarua">Aberdare National Park</location>
,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:42E17D875CD1626500C9C6C951C57CBC" country="Kenya" county="Aberdare National Park" municipality="Uplands District" name="North Kinangop-Nyeri" stateProvince="Nyandarua">North Kinangop-Nyeri</location>
road,
<collectingDate value="1960-07-30">30 Jul 1960</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Polhill 250</emphasis>
(BR, EA, K)
</materialsCitation>
.
<materialsCitation collectingDate="1934-10-28" collectingDateMax="1995-03-29" collectingDateMin="1934-10-28" country="Kenya" county="Mount Nyiru" location="Mbarta forest" municipality="Mount Nyiru" specimenCount="1" stateProvince="Samburu">
<emphasis bold="true" pageId="66" pageNumber="67">Rift Valley</emphasis>
:
<collectingRegion country="Kenya" name="Samburu">Samburu</collectingRegion>
,
<collectingCounty>Mount Nyiru</collectingCounty>
,
<collectingDate value="1955-12-30">30 Dec 1955</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Adamson 543</emphasis>
(EA, K);
<collectingRegion country="Kenya" name="Samburu">Samburu</collectingRegion>
,
<collectingMunicipality>Mount Nyiru</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:8698F0013BBB8393C3A5E1ACF7632094" country="Kenya" county="Mount Nyiru" municipality="Mount Nyiru" name="Mbarta forest" stateProvince="Samburu">Mbarta forest</location>
zone,
<collectingDate value="1995-03-29">29 Mar 1995</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Bytebier et al 28</emphasis>
(BR, EA, K);
<collectingRegion country="Kenya" name="Laikipia">Laikipia</collectingRegion>
, district around Nyasi, Lakipia Plateau and Aberdare range, 1908,
<emphasis italics="true" pageId="66" pageNumber="67">Routledge s.n.</emphasis>
(K); Naivasha, Naivasha-Nyeri track,
<collectingDate value="1934-10-28">28 Oct 1934</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Taylor 1408</emphasis>
(BM)
</materialsCitation>
.
</paragraph>
<paragraph pageId="66" pageNumber="67">
<materialsCitation collectingDate="1930-04" collectingDateMax="1993-03-29" collectingDateMin="1930-04" collectorName="Sheil, Musingizi, Thanes" country="Uganda" county="Mount Elgon" elevation="300" location="Nabulalo" municipality="Mount Elgon" specimenCount="1" stateProvince="Bulambuli">
<emphasis bold="true" pageId="66" pageNumber="67">
<collectingCountry name="Uganda">Uganda</collectingCountry>
. Eastern
</emphasis>
:
<collectingCounty>Mount Elgon</collectingCounty>
,
<collectingDate value="1930-04">Apr 1930</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Liebenberg 1637</emphasis>
(K);
<collectingMunicipality>Mount Elgon</collectingMunicipality>
, western section,
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</elevation>
</quantity>
E of
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:33FB925CAF34BDE2C32FA60A128EA32B" country="Uganda" county="Mount Elgon" municipality="Mount Elgon" name="Nabulalo" stateProvince="Bulambuli">Nabulalo</location>
,
<locationDeviation location="Maika">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
SE of Maika
</locationDeviation>
,
<collectingDate value="1993-03-29">29 Mar 1993</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">
<collectorName>Sheil</collectorName>
&amp;
<collectorName>Musingizi</collectorName>
1813
</emphasis>
(K);
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:1B565ADE6758193B4F400FE3CD86DE02" country="Uganda" county="Mount Elgon" municipality="Mount Elgon" name="Mount Elgon" stateProvince="Bulambuli">Mount Elgon</location>
,
<collectingRegion country="Uganda" name="Bulambuli">Bulambuli</collectingRegion>
,
<collectingDate value="1932-09-06">6 Sep 1932</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">
<collectorName>Thanes</collectorName>
650
</emphasis>
(K)
</materialsCitation>
.
<materialsCitation collectingDate="1906-02-21" collectingDateMax="1970-06-03" collectingDateMin="1906-02-21" collectorName="Kendall, Richardson, Lye, K., Nyamitaba, R, Livingstone, Ruwenzori E" country="Uganda" county="Ruwenzori Mountains" location="Kilembe" municipality="Kaleveru" specimenCount="1" stateProvince="Western">
<emphasis bold="true" pageId="66" pageNumber="67">
<collectingRegion>Western</collectingRegion>
</emphasis>
:
<collectingCounty>Ruwenzori Mountains</collectingCounty>
,
<collectingMunicipality>Kaleveru</collectingMunicipality>
slopes, 6 kms west of
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:A4C281B240FFF8B55FC77A5056650208" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Kilembe" stateProvince="Western">Kilembe</location>
,
<collectingDate value="1970-06-03">3 Jun 1970</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Katende 345 A</emphasis>
(EA); Mahoma,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:5E64BE4FF50B390F576293894BB72538" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Ruwenzori Mountains" stateProvince="Western">Ruwenzori Mountains</location>
<collectingDate value="1960-07-19">19 Jul 1960</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">
<collectorName>Kendall</collectorName>
&amp;
<collectorName>Richardson</collectorName>
25
</emphasis>
(EA, K);
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:3C258EA47849D3E5B8F53FD5DB7B52C2" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Mount Ruwenzori" stateProvince="Western">Mount Ruwenzori</location>
, below Kanyasabo rock shelter,
<collectingDate value="1969-01-03">3 Jan 1969</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">
<collectorName>Lye, K.</collectorName>
1340
</emphasis>
(K);
<collectorName>Nyamitaba, R</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:89B3A7095823DD13CC438FC62C93E6AC" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Mobuku valley" stateProvince="Western">Mobuku valley</location>
,
<collectingDate value="1952-07-11">11 Jul 1952</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Osmaston 1549</emphasis>
(BM);
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:4909329908BEE5E718A6DF196A1A43B3" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Lake Mahoma" stateProvince="Western">Lake Mahoma</location>
,
<collectingDate value="1960-07">Jul 1960</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">
<collectorName>Richardson</collectorName>
&amp;
<collectorName>Livingstone</collectorName>
s.n.
</emphasis>
(DUKE);
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:8386503F3A16E60B48101A8541657C84" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Mobuka valley" stateProvince="Western">Mobuka valley</location>
, above Kichundu,
<collectingDate value="1952-07-15">15 Jul 1952</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Ross 578</emphasis>
(BM); Ruwenzori,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:C4CB0B790BFAF1BC102415B8C61D9382" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Nyamgasani valley" stateProvince="Western">Nyamgasani valley</location>
,
<collectingDate value="1935-01">Jan 1935</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Synge 1444</emphasis>
(BM); Toro, Ruwenzori,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:0EDF44434E20B4A81BD4665BBF8B98E3" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Namwamba valley" stateProvince="Western">Namwamba valley</location>
,
<collectingDate value="1935-01-06">6 Jan 1935</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Taylor 2913</emphasis>
(BM). Bwamba, Ibonde Pass,
<collectingDate value="1932-10-01">1 Oct 1932</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Thomas 767</emphasis>
(EA); Nyabitaba Hut,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:2B8A98EB47D47A2C88318C3754749423" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Bujuku Valley" stateProvince="Western">Bujuku Valley</location>
below hut, slope down to bridge,
<collectingDate value="1967-01-16">16 Jan 1967</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Wood 816</emphasis>
(EA);
<collectorName>Ruwenzori E</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:41E31C342BCB555EB49B60D72E6A18CE:8ACA5BB048FA74C7575E59CCF525631C" country="Uganda" county="Ruwenzori Mountains" municipality="Kaleveru" name="Mt. Ruwenzori" stateProvince="Western">Mt. Ruwenzori</location>
,
<collectingDate value="1906-02-21">21 Feb 1906</collectingDate>
,
<emphasis italics="true" pageId="66" pageNumber="67">Wollaston s.n.</emphasis>
(BM)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>