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<document id="FFF3B5479588DA02615E9D537270C748" ID-DOI="10.5281/zenodo.3673491" ID-GBIF-Dataset="d8f38b1a-d989-4559-8f58-95878de25602" ID-ISSN="1942-1354" ID-Zenodo-Dep="3673491" ID-ZooBank="B2EA9141-6419-4A1A-85CC-BCE190C17802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027722378" checkinUser="felipe" docAuthor="Harris, Steven C. &amp; Armitage, Brian J." docDate="2019" docId="17184D65FFB3651252BEFBD9FDC77DE8" docLanguage="en" docName="InsectaMundi.0707.1-54.pdf.imf" docOrigin="Insecta Mundi 707 (707)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Rhyacopsyche holzenthali Harris and Armitage 2019, new species" docType="treatment" docVersion="11" lastPageNumber="17" masterDocId="EB21351DFFA16500527EFFFCFF967A60" masterDocTitle="The Trichoptera of Panama X. The Quebrada Rambala drainage, with description of 19 new species of microcaddisflies (Trichoptera: Hydroptilidae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="17" updateTime="1698718896375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="DD72F5F9DC6DBFC6B4C41282F1CDA9F9">The Trichoptera of Panama X. The Quebrada Rambala drainage, with description of 19 new species of microcaddisflies (Trichoptera: Hydroptilidae)</mods:title>
</mods:titleInfo>
<mods:name id="409E4A0390D8CB323A5D892676F3F17E" type="personal">
<mods:role id="9530906C9ED222D286F893AB1FB9FBED">
<mods:roleTerm id="F603DB98457DF682ECB75BA49DDBC1EE">Author</mods:roleTerm>
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<mods:namePart id="8AEFA561682A1FD68E5484E21658F007">Harris, Steven C.</mods:namePart>
<mods:affiliation id="A9D8B15967160142287B110A29DD176E">Department of Biology and Geosciences Clarion University Clarion, PA 16214 USA</mods:affiliation>
<mods:nameIdentifier id="9BBAF19B63DC673BE9928407518793DE" type="email">sharris@clarion.edu</mods:nameIdentifier>
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<mods:role id="D6C32B84EB9D5EAFA03CDD2FD38BD83D">
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<mods:namePart id="C6D73CD6527D8BC28A5158153F4B1FE4">Armitage, Brian J.</mods:namePart>
<mods:affiliation id="0B633278B7B57F03E874E09EEEC26C7A">Museo de Peces de Agua Dulce e Invertebrados (MUPADI) Universidad Autónoma de Chiriquí (UNACHI) David, Republic of Panama</mods:affiliation>
<mods:nameIdentifier id="484881B2D3891C87978F7D263B674894" type="email">tobikera89@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="4ADC3A97D141F9CB831356FD09E4B120">Insecta Mundi</mods:title>
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<mods:part id="8CB7880C75CFA5B4D2859D1ABA899B60">
<mods:date id="73E279E365E1D2C7D9496CC7B73CC880">2019</mods:date>
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<mods:number id="D10FB71D6AE81BC93F114E67BEC9E9E0">2019-05-31</mods:number>
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<mods:number id="F0FAFF010A36DF4F9C4215530AAD6ECB">707</mods:number>
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<mods:classification id="347CF23516E6BED65ECF413E7C091D8C">journal article</mods:classification>
<mods:identifier id="C44A94BFBB45399FCED1199DBF91F59F" type="DOI">10.5281/zenodo.3673491</mods:identifier>
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<mods:identifier id="953A0231388F980A0258E127E783267C" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="6D41B3C3758D5B3EBEB46290A5A20A7D" type="Zenodo-Dep">3673491</mods:identifier>
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<treatment id="17184D65FFB3651252BEFBD9FDC77DE8" ID-DOI="http://doi.org/10.5281/zenodo.3680023" ID-GBIF-Taxon="162161865" ID-Zenodo-Dep="3680023" LSID="urn:lsid:plazi:treatment:17184D65FFB3651252BEFBD9FDC77DE8" httpUri="http://treatment.plazi.org/id/17184D65FFB3651252BEFBD9FDC77DE8" lastPageNumber="17" pageId="18" pageNumber="17">
<subSubSection id="D7ABAFF8FFB3651252BEFBD9FC697E5E" box="[192,1023,1061,1086]" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="9F0EFC73FFB3651252BEFBD9FC697E5E" blockId="18.[192,1023,1061,1119]" box="[192,1023,1061,1086]" pageId="18" pageNumber="17">
<heading id="C4464B1FFFB3651252BEFBD9FC697E5E" bold="true" box="[192,1023,1061,1086]" fontSize="10" level="2" pageId="18" pageNumber="17" reason="3">
<emphasis id="ADC52061FFB3651252BEFBD9FC697E5E" bold="true" box="[192,1023,1061,1086]" pageId="18" pageNumber="17">
<taxonomicName id="58B187F0FFB3651252BEFBD9FCDA7E5E" authority="Harris and Armitage" authorityName="Harris and Armitage" authorityYear="2019" box="[192,844,1061,1086]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="holzenthali" status="sp. nov.">
<emphasis id="ADC52061FFB3651252BEFBD9FDB37E5D" bold="true" box="[192,549,1061,1085]" italics="true" pageId="18" pageNumber="17">Rhyacopsyche holzenthali</emphasis>
Harris and Armitage
</taxonomicName>
,
<taxonomicNameLabel id="B6F69D1AFFB365125124FBDAFC697E5E" box="[858,1023,1062,1086]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFB3651252BEFBBAFE837E3F" box="[192,277,1094,1119]" pageId="18" pageNumber="17" type="description">
<paragraph id="9F0EFC73FFB3651252BEFBBAFE837E3F" blockId="18.[192,1023,1061,1119]" box="[192,277,1094,1119]" pageId="18" pageNumber="17">
<figureCitation id="078AE0F6FFB3651252BEFBBAFE837E3F" box="[192,277,1094,1119]" captionStart="Figure 22" captionStartId="46.[192,275,1877,1899]" captionTargetBox="[206,1440,216,1831]" captionTargetId="figure@46.[189,1442,213,1845]" captionTargetPageId="46" captionText="Figure 22. Rhyacopsyche holzenthali, sp. n., Male genitalia. A) Lateral view. B) Ventral view. C) Dorsal view. D) Phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3673535" httpUri="https://zenodo.org/record/3673535/files/figure.png" pageId="18" pageNumber="17">Fig. 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFB3651252BEFB88FE637E8D" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="9F0EFC73FFB3651252BEFB88FE637E8D" blockId="18.[192,1440,1140,1261]" pageId="18" pageNumber="17">
<emphasis id="ADC52061FFB3651252BEFB88FEC67EEC" bold="true" box="[192,336,1140,1164]" pageId="18" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="58B187F0FFB365125327FB88FD007EEC" authorityName="Harris and Armitage" authorityYear="2019" box="[345,662,1140,1164]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="holzenthali">
<emphasis id="ADC52061FFB365125327FB88FD007EEC" box="[345,662,1140,1164]" italics="true" pageId="18" pageNumber="17">Rhyacopsyche holzenthali</emphasis>
</taxonomicName>
appears most closely related to
<taxonomicName id="58B187F0FFB365125655FB89FB677EEC" authority="Flint" authorityName="Flint" authorityYear="1991" box="[1067,1265,1140,1164]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="ADC52061FFB365125655FB89FB3E7EEC" box="[1067,1192,1140,1164]" italics="true" pageId="18" pageNumber="17">R. andina</emphasis>
Flint
</taxonomicName>
,
<taxonomicName id="58B187F0FFB365125681FB89FD907ECC" authority="Wasmund and Holzenthal" authorityName="Wasmund and Holzenthal" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="intraspira">
<emphasis id="ADC52061FFB365125681FB89FA367EEB" box="[1279,1440,1140,1164]" italics="true" pageId="18" pageNumber="17">R. intraspira</emphasis>
Wasmund and Holzenthal
</taxonomicName>
,
<taxonomicName id="58B187F0FFB36512506DFB69FC927ECC" authority="(Flint)" baseAuthorityName="Flint" box="[531,772,1172,1196]" class="Insecta" family="Hydroptilidae" genus="Hydroptila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="ADC52061FFB36512506DFB69FD3C7ECC" box="[531,682,1173,1196]" italics="true" pageId="18" pageNumber="17">R. mexicana</emphasis>
(Flint)
</taxonomicName>
, and
<taxonomicName id="58B187F0FFB365125138FB69FB847ECC" authority="Flint" authorityName="Flint" authorityYear="1971" box="[838,1042,1172,1196]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="torulosa">
<emphasis id="ADC52061FFB365125138FB69FC5B7ECC" box="[838,973,1172,1196]" italics="true" pageId="18" pageNumber="17">R. torulosa</emphasis>
Flint
</taxonomicName>
in the shape of inferior appendages in lateral view. However, in these species the dorsum of segment IX is deeply divided, which is the new species is entire.
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFB3651252BEFAFFFE227CEB" pageId="18" pageNumber="17" type="description">
<paragraph id="9F0EFC73FFB3651252BEFAFFFD2E7C3E" blockId="18.[192,1440,1283,1630]" pageId="18" pageNumber="17">
<emphasis id="ADC52061FFB3651252BEFAFFFE9C7F7B" bold="true" box="[192,266,1283,1307]" pageId="18" pageNumber="17">Male.</emphasis>
Length
<quantity id="58495196FFB36512530FFAFFFD937F7A" box="[369,517,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="18" pageNumber="17" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
. 39 antennal segments, tufts of hair between and behind the scapes, brown in alcohol with wings mottled with white hairs. Abdominal segment VIII annular. Segment IX produced dorsally into shelf-like lobe, tapering distally and bearing row of dark, peg-like setae ventrally, truncate lobe mesally; in dorsal view truncate posteriorly with small notch mesally, cluster of dark peg-like setae on margin, anteriorly deeply incised, truncate mesally. Tergum X narrow and shelf-like, retracted within segment IX posteriorly, anteriorly articulating with subgenital plate, which is round distally and ventrally, and bears several stout setae from venter. Inferior appendages narrow basally, dorsal margin expanded at midlength, then abruptly narrowing to finger-like apex which bears an elongate dorsal spine; in dorsal and ventral views, nearly parallel-sided until narrowing apically and curving along the outer margin. Phallus in dorsal view tubular, thin membranous sheath subapically, both central and lateral processes linear.
</paragraph>
<paragraph id="9F0EFC73FFB3651252BEF98FFE227CEB" blockId="18.[192,436,1651,1675]" box="[192,436,1651,1675]" pageId="18" pageNumber="17">
<emphasis id="ADC52061FFB3651252BEF98FFEBB7CEB" bold="true" box="[192,301,1651,1675]" pageId="18" pageNumber="17">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFB3651252BEF95DFAEB7D7A" pageId="18" pageNumber="17" type="materials_examined">
<paragraph id="9F0EFC73FFB3651252BEF95DFAEB7D7A" blockId="18.[192,1440,1697,1818]" pageId="18" pageNumber="17">
<emphasis id="ADC52061FFB3651252BEF95DFE1D7CD9" bold="true" box="[192,395,1697,1721]" pageId="18" pageNumber="17">Type material.</emphasis>
<materialsCitation id="2FD9F62EFFB3651253EAF95DFE4C7C9A" ID-GBIF-Occurrence="2565952439" collectingDate="2017-02-06" collectingDateMax="2017-02-12" collectingDateMin="2017-02-06" collectionCode="SCH" collectorName="E. Carlson" country="Panama" county="Quebrada Rambala" elevation="120" latitude="8.91627" location="Rambala Jungle Lodge" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" pageId="18" pageNumber="17" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<emphasis id="ADC52061FFB3651253EAF95DFBC07CD9" bold="true" box="[404,1110,1697,1721]" pageId="18" pageNumber="17">
<typeStatus id="400A42D1FFB3651253EAF95DFD857CD9" box="[404,531,1697,1721]" pageId="18" pageNumber="17" type="holotype">Holotype</typeStatus>
, male—
<collectingCountry id="E7A6BCE3FFB3651250FEF95DFD647CD9" box="[640,754,1697,1721]" name="Panama" pageId="18" pageNumber="17">Panama</collectingCountry>
,
<collectingRegion id="5D753291FFB36512517FF95DFBC07CD9" box="[769,1110,1697,1721]" country="Panama" name="Bocas del Toro" pageId="18" pageNumber="17">Bocas del Toro Province</collectingRegion>
</emphasis>
,
<collectingCounty id="766F84FFFFB365125618F95DFACD7CD9" box="[1126,1371,1697,1721]" pageId="18" pageNumber="17">
Quebrada
<collectingMunicipality id="7F6A6609FFB365125697F95DFACD7CD9" box="[1257,1371,1697,1721]" pageId="18" pageNumber="17">Rambala</collectingMunicipality>
</collectingCounty>
, near
<location id="9A6EAAA8FFB3651252BEF93DFE457CB9" LSID="urn:lsid:plazi:treatment:17184D65FFB3651252BEFBD9FDC77DE8:9A6EAAA8FFB3651252BEF93DFE457CB9" box="[192,467,1729,1753]" country="Panama" county="Quebrada Rambala" latitude="8.91627" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" name="Rambala Jungle Lodge" pageId="18" pageNumber="17" stateProvince="Bocas del Toro">
<location id="9A6EAAA8FFB3651252BEF93DFEBA7CB9" LSID="urn:lsid:plazi:treatment:17184D65FFB3651252BEFBD9FDC77DE8:9A6EAAA8FFB3651252BEF93DFEBA7CB9" box="[192,300,1729,1753]" country="Panama" county="Quebrada Rambala" latitude="8.91627" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" name="Rambala" pageId="18" pageNumber="17" stateProvince="Bocas del Toro">Rambala</location>
Jungle Lodge
</location>
,
<locationDeviation id="8EBCE551FFB3651253A3F93EFD4D7CB9" box="[477,731,1729,1754]" location="Rambala" pageId="18" pageNumber="17">
<quantity id="58495196FFB3651253A3F93EFDB87CB9" box="[477,558,1729,1754]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="18" pageNumber="17" unit="km" value="3.7">3.7 km</quantity>
SSE Rambala
</locationDeviation>
,
<geoCoordinate id="FA859AB4FFB36512509AF93DFCF47CBA" box="[740,866,1729,1754]" degrees="8.91627" direction="north" orientation="latitude" pageId="18" pageNumber="17" precision="1" value="8.91627">8.91627°N</geoCoordinate>
and
<geoCoordinate id="FA859AB4FFB3651251E4F93DFBB87CB9" box="[922,1070,1729,1753]" degrees="82.15469" direction="west" orientation="longitude" pageId="18" pageNumber="17" precision="1" value="-82.15469">82.15469°W</geoCoordinate>
,
<quantity id="58495196FFB365125649F93EFB147CB9" box="[1079,1154,1730,1754]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="17" unit="m" value="120.0">
<elevation id="149C1B40FFB365125649F93EFB147CB9" box="[1079,1154,1730,1754]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="17" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<collectingMethod id="46F08464FFB3651256F2F93DFAB37CB9" box="[1164,1317,1729,1753]" pageId="18" pageNumber="17">Malaise trap</collectingMethod>
,
<date id="EB0FDAB3FFB365125751F93DFEDE7C9A" pageId="18" pageNumber="17" value="2017-02-06" valueMax="2017-02-12" valueMin="2017-02-06">
<collectingDate id="FB4B235BFFB365125751F93DFEDE7C9A" pageId="18" pageNumber="17" value="2017-02-06" valueMax="2017-02-12" valueMin="2017-02-06">February 612, 2017</collectingDate>
</date>
,
<collectorName id="324499A5FFB36512532AF91EFE4C7C9A" box="[340,474,1762,1786]" pageId="18" pageNumber="17">E. Carlson</collectorName>
</materialsCitation>
.
<materialsCitation id="2FD9F62EFFB36512539AF91EFC8C7C9A" ID-GBIF-Occurrence="2565952514" box="[484,794,1762,1786]" collectingDate="2017-02-06" collectingDateMax="2017-02-12" collectingDateMin="2017-02-06" collectionCode="SCH" collectorName="E. Carlson" country="Panama" county="Quebrada Rambala" elevation="120" latitude="8.91627" location="Rambala Jungle Lodge" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" pageId="18" pageNumber="17" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<emphasis id="ADC52061FFB36512539AF91EFDEF7C9A" bold="true" box="[484,633,1762,1786]" pageId="18" pageNumber="17">
<typeStatus id="400A42D1FFB36512539AF91EFDE47C9A" box="[484,626,1762,1786]" pageId="18" pageNumber="17" type="paratype">Paratypes</typeStatus>
.
</emphasis>
ibid.,
<specimenCount id="89B737FAFFB3651250BBF91EFC8C7C9A" box="[709,794,1762,1786]" pageId="18" pageNumber="17" type="male">1 male</specimenCount>
</materialsCitation>
;
<materialsCitation id="2FD9F62EFFB36512515BF91EFB697C9A" ID-GBIF-Occurrence="2565952512" box="[805,1279,1762,1786]" collectingDate="2015-03-31" collectingDateMax="2015-04-11" collectingDateMin="2015-03-31" collectionCode="SCH" collectorName="E. Carlson" country="Panama" county="Quebrada Rambala" elevation="120" latitude="8.91627" location="Rambala Jungle Lodge" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" pageId="18" pageNumber="17" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
ibid.,
<date id="EB0FDAB3FFB365125114F91EFB087C9A" box="[874,1182,1762,1786]" pageId="18" pageNumber="17" value="2015-03-31" valueMax="2015-04-11" valueMin="2015-03-31">
<collectingDate id="FB4B235BFFB365125114F91EFB087C9A" box="[874,1182,1762,1786]" pageId="18" pageNumber="17" value="2015-03-31" valueMax="2015-04-11" valueMin="2015-03-31">March 31April 11, 2015</collectingDate>
</date>
,
<specimenCount id="89B737FAFFB3651256D4F91EFB697C9A" box="[1194,1279,1762,1786]" pageId="18" pageNumber="17" type="male">1 male</specimenCount>
</materialsCitation>
;
<materialsCitation id="2FD9F62EFFB365125775F91EFE1A7D7A" ID-GBIF-Occurrence="2565952506" collectingDate="2015-03-28" collectionCode="SCH" collectorName="E. Carlson" country="Panama" county="Quebrada Rambala" elevation="120" latitude="8.91627" location="Rambala Jungle Lodge" longLatPrecision="1" longitude="-82.15469" municipality="Rambala" pageId="18" pageNumber="17" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="paratype">
ibid.,
<date id="EB0FDAB3FFB36512572EF91EFEBC7D7A" pageId="18" pageNumber="17" value="2015-03-28">
<collectingDate id="FB4B235BFFB36512572EF91EFEBC7D7A" pageId="18" pageNumber="17" value="2015-03-28">March 28, 2015</collectingDate>
</date>
,
<specimenCount id="89B737FAFFB365125349F8FEFE1A7D7A" box="[311,396,1794,1818]" pageId="18" pageNumber="17" type="male">1 male</specimenCount>
</materialsCitation>
;
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ibid., tributary of Quebrada
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,
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<elevation id="149C1B40FFB36512510CF8FEFC567D7A" box="[882,960,1794,1818]" metricMagnitude="2" metricUnit="m" metricValue="1.34" pageId="18" pageNumber="17" unit="m" value="134.0">134 m</elevation>
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,
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<collectingDate id="FB4B235BFFB3651251B3F8FEFB557D7A" box="[973,1219,1794,1818]" pageId="18" pageNumber="17" value="2015-09-29">September 29, 2015</collectingDate>
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(
<collectionCode id="F9A064B6FFB36512574CF8FEFAE77D7A" box="[1330,1393,1794,1818]" country="Switzerland" lsid="urn:lsid:biocol.org:col:13675" name="Museum zu Allerheiligen" pageId="18" pageNumber="17" type="Herbarium">SCH</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="D7ABAFF8FFB3651252BEF8CCFDC77DE8" pageId="18" pageNumber="17" type="etymology">
<paragraph id="9F0EFC73FFB3651252BEF8CCFDC77DE8" blockId="18.[192,1440,1840,1929]" pageId="18" pageNumber="17">
<emphasis id="ADC52061FFB3651252BEF8CCFEC57D28" bold="true" box="[192,339,1840,1864]" pageId="18" pageNumber="17">Etymology</emphasis>
. We take great pleasure in naming this species for our colleague Ralph Holzenthal, who has contributed much to our knowledge of neotropical caddisflies and, along with Anne Wasmund, recently revised the genus
<taxonomicName id="58B187F0FFB3651253DEF88CFDDC7DE8" authorityName="F Mueller" authorityYear="1879" box="[416,586,1904,1928]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="ADC52061FFB3651253DEF88CFDDC7DE8" box="[416,586,1904,1928]" italics="true" pageId="18" pageNumber="17">Rhyacopsyche</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>