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<document id="94D74A7044F5CD7587D3EAD10965B4DD" ID-CLB-Dataset="7678" ID-DOI="10.1206/359.1" ID-GBIF-Dataset="f399e6d5-8c36-4871-ae88-64c70850516c" ID-ISSN="0003-0090" ID-Zenodo-Dep="4611111" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615989739501" checkinUser="felipe" docAuthor="Ochoa, José A., Ojanguren Affilastro, Andres A., Mattoni, Camilo I. &amp; Prendini, Lorenzo" docDate="2011" docId="083185411F05FFDB1A7AFD0FFB1D466D" docLanguage="en" docName="BulAmeMusNatHis.2011.359.1-90.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (359)" docSource="http://www.bioone.org/doi/abs/10.1206/359.1" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Orobothriurus MAURY 1976" docType="key" docVersion="5" lastPageNumber="15" masterDocId="F408FD391F0BFFD41A03FFC9FFD64231" masterDocTitle="Systematic Revision Of The Andean Scorpion Genus Orobothriurus Maury, 1976 (Bothriuridae), With Discussion Of The Altitude Record For Scorpions" masterLastPageNumber="90" masterPageNumber="1" pageNumber="14" updateTime="1698933505625" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AF7EE88F66C13F39FC62BC7FE130275B">Systematic Revision Of The Andean Scorpion Genus Orobothriurus Maury, 1976 (Bothriuridae), With Discussion Of The Altitude Record For Scorpions</mods:title>
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<mods:namePart id="DAC03DA53966A97521F8C9E7AA530E91">Ochoa, José A.</mods:namePart>
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<mods:namePart id="188312AA841143DE4A8EAE50A6F75EC1">Mattoni, Camilo I.</mods:namePart>
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<mods:namePart id="CE9D7545CB9F0652A728A6873D38B3BD">Prendini, Lorenzo</mods:namePart>
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<treatment id="083185411F05FFDB1A7AFD0FFB1D466D" ID-DOI="http://doi.org/10.5281/zenodo.6841987" ID-GBIF-Taxon="197619913" ID-Zenodo-Dep="6841987" LSID="urn:lsid:plazi:treatment:083185411F05FFDB1A7AFD0FFB1D466D" httpUri="http://treatment.plazi.org/id/083185411F05FFDB1A7AFD0FFB1D466D" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14" type="materials_methods">
<subSubSection id="C88267DC1F05FFDA1A7AFD0FFDE240C8" pageId="14" pageNumber="14" type="nomenclature">
<paragraph id="802734571F05FFDA1A7AFD0FFDE240C8" blockId="14.[121,591,710,761]" pageId="14" pageNumber="14">
KEY TO IDENTIFICATION OF THE SPECIES OF
<taxonomicName id="47984FD41F05FFDA1B39FD2DFDE240C8" ID-CoL="69YV" authorityName="MAURY" authorityYear="1976" box="[314,564,740,761]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B2ECE8451F05FFDA1B39FD2DFDE240C8" box="[314,564,740,761]" italics="true" pageId="14" pageNumber="14">OROBOTHRIURUS</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C88267DC1F05FFDB1A5EFCDEFB1D466D" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14" type="key">
<keyStep id="3B6C2CF21F05FFDA1A5EFCDEFDBD46EA" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1A5EFCDEFDBA41FD" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EFCDEFDBA41FD" pageId="14" pageNumber="14">
1. Tergites IVI entirely and densely pigmented (fig. 12D); metasomal segments IV, VM stripe contiguous with paired VL stripes posteriorly (fig. 11A); metasomal segment IV, VSM carinae present (fig. 16B); angle formed between pedipalp chela trichobothria
<emphasis id="B2ECE8451F05FFDA1819FC54FD834180" box="[538,597,925,945]" italics="true" pageId="14" pageNumber="14">ebEt</emphasis>
<subScript id="1C1C36121F05FFDA1855FC6FFD894185" attach="right" box="[598,607,934,948]" fontSize="6" pageId="14" pageNumber="14">5</subScript>
<emphasis id="B2ECE8451F05FFDA185CFC56FF7C41FD" italics="true" pageId="14" pageNumber="14"> Et</emphasis>
<subScript id="1C1C36121F05FFDA1AA9FC08FF6541FE" attach="left" box="[170,179,961,975]" fontSize="6" pageId="14" pageNumber="14">4</subScript>
less than 90
<emphasis id="B2ECE8451F05FFDA1B36FC70FEE841FD" bold="true" box="[309,318,953,972]" pageId="14" pageNumber="14">°</emphasis>
(fig. 25A)......
<taxonomicName id="47984FD41F05FFDA1804FC70FDBA41FD" ID-CoL="6T883" authorityName="Ochoa and Acosta" authorityYear="2003" box="[519,620,953,972]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="ampay">
<emphasis id="B2ECE8451F05FFDA1804FC70FDBA41FD" box="[519,620,953,972]" italics="true" pageId="14" pageNumber="14">O. ampay</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="802734571F05FFDA1A5EFC1CFDBD46EA" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EFC1CFDBD46EA" pageId="14" pageNumber="14">
Tergites IVI each with paired lateral spots of pigmentation delimiting unpigmented median longitudinal stripe (fig. 13C, EG); metasomal segments IV, VM stripe not contiguous with paired VL stripes posteriorly (at least on segments IV and V; figs. 11BD, 13A, D); metasomal segment IV, VSM carinae absent (fig. 17A); angle formed between pedipalp chela trichobothria
<emphasis id="B2ECE8451F05FFDA1B6AFB65FE72468E" box="[361,420,1196,1216]" italics="true" pageId="14" pageNumber="14">ebEt</emphasis>
<subScript id="1C1C36121F05FFDA1BA6FB7CFE7846F2" attach="right" box="[421,430,1205,1219]" fontSize="6" pageId="14" pageNumber="14">5</subScript>
<emphasis id="B2ECE8451F05FFDA1BB9FB64FE04468E" box="[442,466,1197,1215]" italics="true" pageId="14" pageNumber="14">Et</emphasis>
<subScript id="1C1C36121F05FFDA1BD1FB7CFE0D46F2" attach="left" box="[466,475,1205,1219]" fontSize="6" pageId="14" pageNumber="14">4</subScript>
greater than 90
<emphasis id="B2ECE8451F05FFDA1AA9FB0EFF6546EB" bold="true" box="[170,179,1223,1242]" pageId="14" pageNumber="14">°</emphasis>
(fig. 25B, D).................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1A5EFB2AFDBD4785" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1A5EFB2AFDBD4776" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EFB2AFDBD4776" pageId="14" pageNumber="14">2. Carapace, anterior margin with weak median projection (epistome) (fig. 14C, 15A, C, D); pedipalp chela, movable finger (♂) distinctly curved (fig. 26B, C)................ 3</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1A5EFA99FDBD4785" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EFA99FDBD4785" pageId="14" pageNumber="14"> Carapace, anterior margin sublinear or emarginate, with shallow median notch (figs. 14A, B, D, 15B, 16); pedipalp chela, movable finger (♂) straight (figs. 25B, D, 31C)... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1A5EFA74FDBD44EF" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1A5EFA74FDBD4467" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EFA74FDBD4467" pageId="14" pageNumber="14">
3. Metasomal segment V (
<specimenCount id="969EFFDE1F05FFDA1B8FFA70FE4847E1" box="[396,414,1465,1488]" pageId="14" pageNumber="14" type="female"></specimenCount>
) slightly elongated, length greater than or equal to twice its width (fig. 22C); metasomal segment I with two pairs of VSM macrosetae (fig. 19A, C, F); pedipalp chela, movable finger (♂) strongly curved (figs. 26C, 43C, 51A)......... 4
</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1A5EF996FDBD44EF" blockId="14.[93,620,791,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1A5EF996FDBD44EF" pageId="14" pageNumber="14">
Metasomal segment V (
<specimenCount id="969EFFDE1F05FFDA1B95F995FE7E4442" box="[406,424,1628,1651]" pageId="14" pageNumber="14" type="female"></specimenCount>
), length less than twice its width (fig. 22B); metasomal segment I with three pairs of VSM macrosetae (fig. 19B); pedipalp chela, movable finger (♂) slightly curved (fig. 26B)...
<taxonomicName id="47984FD41F05FFDA1BF0F902FDBD44EF" ID-CoL="75883" baseAuthorityName="Kraepelin" baseAuthorityYear="1911" box="[499,619,1738,1758]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="paessleri">
<emphasis id="B2ECE8451F05FFDA1BF0F902FDBD44EF" box="[499,619,1738,1758]" italics="true" pageId="14" pageNumber="14">O. paessleri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1898FF1DFB7F40A3" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1898FF1DFB7F4390" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FF1DFB7F4390" pageId="14" pageNumber="14">
4. Carapace, tergites, metasomal segments, ventral surface unpigmented (fig. 49); sternite VII finely granular (fig. 19F); metasomal segments IIV, VSM carinae present, at least on segments I and II (fig. 19F); hemispermatophore, frontal crest shorter than apex length, ventral border of distal part of apex straight (fig. 47C, D).......
<taxonomicName id="47984FD41F05FFDA1E23FE47FB7F4390" ID-CoL="6SW7Q" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1056,1193,397,417]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="tamarugal">
<emphasis id="B2ECE8451F05FFDA1E23FE47FB7F4390" bold="true" box="[1056,1193,397,417]" italics="true" pageId="14" pageNumber="14">O. tamarugal</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1898FE63FB7F40A3" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FE63FB7F40A3" pageId="14" pageNumber="14"> Carapace, tergites, metasomal segments, ventral surface pigmented (figs. 11B, 13E); sternite VII smooth or nearly so (fig. 19A, C); metasomal segments IIV, VSM carinae absent (fig. 19A, C); hemispermatophore, frontal crest slightly elongated, length greater than or equal to apex length, ventral border of distal part of apex slightly curved (figs. 28B, D, 38D, E)........................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1898FD53FB7F4612" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1898FD53FB7C417C" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FD53FB7C417C" pageId="14" pageNumber="14">
5. Carapace, tergites, metasomal segments, and pedipalps markedly pigmented (figs. 11B, 12C, 13E); telson vesicle, ventral surface (♂) granular in anterior third; pedipalp chela, trichobothrium
<emphasis id="B2ECE8451F05FFDA1926FCCDFC9D4129" box="[805,843,772,792]" italics="true" pageId="14" pageNumber="14">Esb</emphasis>
situated between trichobothria
<emphasis id="B2ECE8451F05FFDA18D3FCD6FD3A4102" box="[720,748,799,819]" italics="true" pageId="14" pageNumber="14">Eb</emphasis>
<subScript id="1C1C36121F05FFDA18EFFCE1FD234107" attach="left" box="[748,757,808,822]" fontSize="6" pageId="14" pageNumber="14">2</subScript>
and
<emphasis id="B2ECE8451F05FFDA1929FCD6FC904102" box="[810,838,799,819]" italics="true" pageId="14" pageNumber="14">Eb</emphasis>
<subScript id="1C1C36121F05FFDA1944FCE1FC864107" attach="right" box="[839,848,808,822]" fontSize="6" pageId="14" pageNumber="14">3</subScript>
; hemispermatophore, distal crest without folds (fig. 28B).....
<taxonomicName id="47984FD41F05FFDA1E17FCF3FB7C417C" ID-CoL="74W83" baseAuthorityName="Kraepelin" baseAuthorityYear="1911" box="[1044,1194,825,845]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="curvidigitus">
<emphasis id="B2ECE8451F05FFDA1E17FCF3FB7C417C" bold="true" box="[1044,1194,825,845]" italics="true" pageId="14" pageNumber="14">O. curvidigitus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1898FC9FFB7F4612" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FC9FFB7F4612" pageId="14" pageNumber="14">
Carapace and tergites faintly pigmented, metasomal segments and pedipalps unpigmented (fig. 41); telson vesicle, ventral surface (♂) entirely granular (fig. 23F); pedipalp chela, trichobothrium
<emphasis id="B2ECE8451F05FFDA19CDFC09FC2241E5" box="[974,1012,960,980]" italics="true" pageId="14" pageNumber="14">Esb</emphasis>
situated dorsal to trichobothrium
<emphasis id="B2ECE8451F05FFDA1998FC13FC6141DF" box="[923,951,986,1006]" italics="true" pageId="14" pageNumber="14">Eb</emphasis>
<subScript id="1C1C36121F05FFDA19BBFC2AFC1741C0" attach="right" box="[952,961,995,1009]" fontSize="6" pageId="14" pageNumber="14">2</subScript>
; hemispermatophore, distal crest with one or two small folds (fig. 40E).............
<taxonomicName id="47984FD41F05FFDA1E01FBD9FB7F4612" ID-CoL="6SVW2" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1026,1193,1039,1059]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="quewerukana">
<emphasis id="B2ECE8451F05FFDA1E01FBD9FB7F4612" bold="true" box="[1026,1193,1039,1059]" italics="true" pageId="14" pageNumber="14">O. quewerukana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1898FBE5FB7F46F4" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1898FBE5FB7F4644" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FBE5FB7F4644" pageId="14" pageNumber="14">6. Hemispermatophore, frontal crest reduced, length less than apex length (figs. 27, 36FH, KO).......................... 7</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1898FBB4FB7F46F4" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FBB4FB7F46F4" pageId="14" pageNumber="14"> Hemispermatophore, frontal crest elongated, length greater than apex length (fig. 36D, E, I, J)........................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F05FFDA1898FB07FB7F47DD" pageId="14" pageNumber="14">
<paragraph id="802734571F05FFDA1898FB07FB7F4757" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FB07FB7F4757" pageId="14" pageNumber="14">7. Metasomal segments IIII, dorsal surfaces each with two median subtriangular spots (occasionally absent on I), usually separated medially by narrow unpigmented line (fig. 12A, C); segment IV with three pairs of VSM macrosetae.................. 8</keyLead>
</paragraph>
<paragraph id="802734571F05FFDA1898FAA6FB7F47DD" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FAA6FB7F47DD" pageId="14" pageNumber="14"> Metasomal segments IIII, dorsal surfaces each without median spots, but with reticulate pigmentation along DL carinae (fig. 12B, 13C); segment IV with four or five pairs of VSM macrosetae (fig. 17A).......... 10</keyLead>
</paragraph>
</keyStep>
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<paragraph id="802734571F05FFDA1898FA3CFB7F4499" blockId="14.[667,1194,212,1759]" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDA1898FA3CFB7F4499" pageId="14" pageNumber="14">
8. Carapace, anterior margin with shallow median notch (fig. 14B); metasoma (♂), VL and VSM carinae obsolete, granular on segment I, absent on II (fig. 18F); hemispermatophore apex 52
<emphasis id="B2ECE8451F05FFDA199FF996FC644443" box="[924,946,1631,1650]" italics="true" pageId="14" pageNumber="14">%</emphasis>
54
<emphasis id="B2ECE8451F05FFDA19D5F996FC3A4443" box="[982,1004,1631,1650]" italics="true" pageId="14" pageNumber="14">%</emphasis>
of lamina length, terminal process of basal lobe well developed (figs. 32C, D, 36AC)....
<taxonomicName id="47984FD41F05FFDA19FDF95CFB7F4499" ID-CoL="74W7R" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1022,1193,1685,1704]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="compagnuccii">
<emphasis id="B2ECE8451F05FFDA19FDF95CFB7F4499" bold="true" box="[1022,1193,1685,1704]" italics="true" pageId="14" pageNumber="14">O. compagnuccii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F05FFDB1898F978FD5B435D" blockId="14.[667,1194,212,1759]" lastBlockId="15.[128,654,212,1758]" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14">
<keyLead id="3B6997621F05FFDB1898F978FD5B435D" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14">
Carapace, anterior margin with deep median notch (fig. 14A); metasoma (♂), VL and VSM carinae well developed, granular, occupying entire length of segment I and weakly developed but complete on II; hemispermatophore apex 66
<emphasis id="B2ECE8451F04FFDB1B6EFEEDFE554306" box="[365,387,292,311]" italics="true" pageId="15" pageNumber="15">%</emphasis>
72
<emphasis id="B2ECE8451F04FFDB1BA4FEEDFE6B4306" box="[423,445,292,311]" italics="true" pageId="15" pageNumber="15">%</emphasis>
of lamina length, terminal process of basal lobe small (fig. 32A, B, 36FH, KO).................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB1A83FEBCFD5B403F" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB1A83FEBCFD5B438C" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FEBCFD5B438C" pageId="15" pageNumber="15">
9. Metasomal segment IV with two or three pairs of ML macrosetae; segment III usually with one pair of DL macrosetae...
<taxonomicName id="47984FD41F04FFDB180DFE63FD5B438C" ID-CoL="74VW3" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[526,653,425,445]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="calchaqui">
<emphasis id="B2ECE8451F04FFDB180DFE63FD5B438C" bold="true" box="[526,653,425,445]" italics="true" pageId="15" pageNumber="15">O. calchaqui</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB1A83FE0FFD5B403F" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FE0FFD5B403F" pageId="15" pageNumber="15">
Metasomal segment IV with one pair of ML macrosetae; segment III usually without DL macrosetae...............
<taxonomicName id="47984FD41F04FFDB1811FE32FD5B403F" ID-CoL="757VR" authorityName="Acosta" authorityYear="2001" box="[530,653,506,526]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="famatina">
<emphasis id="B2ECE8451F04FFDB1811FE32FD5B403F" bold="true" box="[530,653,506,526]" italics="true" pageId="15" pageNumber="15">O. famatina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB1A83FDDEFD584614" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB1A83FDDEFD58412B" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FDDEFD58412B" pageId="15" pageNumber="15">
10. Pedipalp chela manus (♂), internal surface with apophysis (fig. 8B); pedipalp chela fixed finger (♂) slightly curved, creating small gap with movable finger when fingers closed (fig. 22A); metasomal segment V, VL and VM carinae complete (fig. 22E); metasomal segments with distinct VM pigmentation stripe (at least on segments IIV); hemispermatophore, ventral border of apex slightly curved (fig. 27B)............
<taxonomicName id="47984FD41F04FFDB1829FCCEFD58412B" ID-CoL="6T7W2" authorityName="Maury" authorityYear="1976" box="[554,654,775,794]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B2ECE8451F04FFDB1829FCCEFD58412B" bold="true" box="[554,654,775,794]" italics="true" pageId="15" pageNumber="15">O. parvus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB1A83FCEAFD584614" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FCEAFD584614" pageId="15" pageNumber="15">
Pedipalp chela manus (♂), internal surface without apophysis (fig. 25C); pedipalp chela fixed finger (♂) straight, no gap evident when fingers closed (fig. 25B); metasomal segment V, VL and VM carinae absent (♂; fig. 21C) or restricted to distal third (
<specimenCount id="969EFFDE1F04FFDB1807FC6CFDC0418D" box="[516,534,933,956]" pageId="15" pageNumber="15" type="female"></specimenCount>
; fig. 22F); metasomal segments irregularly pigmented, VM pigmentation stripe indistinct (fig. 13D); hemispermatophore, ventral border of apex straight (fig. 27C)...........
<taxonomicName id="47984FD41F04FFDB1826FBDBFD584614" ID-CoL="6SW82" authorityName="Acosta and Ochoa" authorityYear="2000" box="[549,654,1042,1061]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="wawita">
<emphasis id="B2ECE8451F04FFDB1826FBDBFD584614" bold="true" box="[549,654,1042,1061]" italics="true" pageId="15" pageNumber="15">O. wawita</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB1A83FBE7FD584413" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB1A83FBE7FD5B4700" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FBE7FD5B4700" pageId="15" pageNumber="15">
11. Metasoma segments IIII (
<specimenCount id="969EFFDE1F04FFDB1BD4FBE2FE3F4673" box="[471,489,1067,1090]" pageId="15" pageNumber="15" type="female"></specimenCount>
), VSM carinae present (fig. 19D); segment V, VL and VM carinae restricted to posterior two-thirds (fig. 21F); pedipalp chela manus, trichobothrium
<emphasis id="B2ECE8451F04FFDB1B01FB51FEFE469D" box="[258,296,1176,1196]" italics="true" pageId="15" pageNumber="15">Esb</emphasis>
situated between
<emphasis id="B2ECE8451F04FFDB1BE1FB51FE28469D" box="[482,510,1176,1196]" italics="true" pageId="15" pageNumber="15">Eb</emphasis>
<subScript id="1C1C36121F04FFDB1BFDFB68FDD1469E" attach="left" box="[510,519,1185,1199]" fontSize="6" pageId="15" pageNumber="15">2</subScript>
and
<emphasis id="B2ECE8451F04FFDB183FFB51FD8E469D" box="[572,600,1176,1196]" italics="true" pageId="15" pageNumber="15">Eb</emphasis>
<subScript id="1C1C36121F04FFDB185BFB68FDB7469E" attach="left" box="[600,609,1185,1199]" fontSize="6" pageId="15" pageNumber="15">3</subScript>
,
<emphasis id="B2ECE8451F04FFDB186DFB51FD5B469D" box="[622,653,1176,1196]" italics="true" pageId="15" pageNumber="15">Db</emphasis>
slightly closer to
<emphasis id="B2ECE8451F04FFDB1B71FB7DFE5B46F7" box="[370,397,1204,1222]" italics="true" pageId="15" pageNumber="15">Dt</emphasis>
than to
<emphasis id="B2ECE8451F04FFDB1BF0FB7AFDD946F6" box="[499,527,1203,1223]" italics="true" pageId="15" pageNumber="15">Eb</emphasis>
<subScript id="1C1C36121F04FFDB180CFB75FDCE46FB" attach="left" box="[527,536,1212,1226]" fontSize="6" pageId="15" pageNumber="15">3</subScript>
(fig. 39C); hemispermatophore, distal border of apex slightly rounded, frontal crest with internal lateral projection vestigial (fig. 40AC)..........................
<taxonomicName id="47984FD41F04FFDB1805FAD7FD5B4700" ID-CoL="7587R" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[518,653,1309,1329]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="huascaran">
<emphasis id="B2ECE8451F04FFDB1805FAD7FD5B4700" bold="true" box="[518,653,1309,1329]" italics="true" pageId="15" pageNumber="15">O. huascaran</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB1A83FAF3FD584413" blockId="15.[128,654,212,1758]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83FAF3FD584413" pageId="15" pageNumber="15">
Metasoma segments IIII (
<specimenCount id="969EFFDE1F04FFDB1BD4FAFEFE3F477F" box="[471,489,1335,1358]" pageId="15" pageNumber="15" type="female"></specimenCount>
), VSM carinae absent; metasomal segment V, VL, and VM carinae complete (fig. 20A, C, F); pedipalp chela manus, trichobothrium
<emphasis id="B2ECE8451F04FFDB1805FA40FDFA47AC" box="[518,556,1417,1437]" italics="true" pageId="15" pageNumber="15">Esb</emphasis>
situated dorsal to
<emphasis id="B2ECE8451F04FFDB1B24FA6DFE954789" box="[295,323,1444,1464]" italics="true" pageId="15" pageNumber="15">Eb</emphasis>
<subScript id="1C1C36121F04FFDB1B40FA65FE9A478B" attach="left" box="[323,332,1452,1466]" fontSize="6" pageId="15" pageNumber="15">2</subScript>
,
<emphasis id="B2ECE8451F04FFDB1B62FA6DFE564789" box="[353,384,1444,1464]" italics="true" pageId="15" pageNumber="15">Db</emphasis>
equidistant between
<emphasis id="B2ECE8451F04FFDB1870FA6CFD584786" box="[627,654,1445,1463]" italics="true" pageId="15" pageNumber="15">Dt</emphasis>
and
<emphasis id="B2ECE8451F04FFDB1AE5FA77FED447E3" box="[230,258,1470,1490]" italics="true" pageId="15" pageNumber="15">Eb</emphasis>
<subScript id="1C1C36121F04FFDB1B00FA0EFEDA47E4" attach="left" box="[259,268,1479,1493]" fontSize="6" pageId="15" pageNumber="15">3</subScript>
(figs. 25D, 46B, C); hemispermatophore, distal border of apex subtriangular, frontal crest with internal lateral projection complete (figs. 28A, 47A, B)......... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB1A83F9E2FB1D4072" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB1A83F9E2FB1D4306" blockId="15.[128,654,212,1758]" lastBlockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB1A83F9E2FB1D4306" pageId="15" pageNumber="15">
12. Sternite VII acarinate (fig. 18A); metasomal segment II (♂), VSM carinae absent (fig. 18A); segment V, VSM and VM carinae not discernible from dense surface granulation (fig. 21A); legs III and IV, telotarsi each with 3/4 spiniform macrosetae in pro- and retroventral rows; metasomal segments IIII, paired VL stripes contiguous with VM stripe posteriorly (but not on segments IV and V) (fig. 11D); segment IV with four pairs of VSM macrosetae...........
<taxonomicName id="47984FD41F04FFDB1E5BFEEDFB1D4306" ID-CoL="74VVR" authorityName="Ochoa and Acosta" authorityYear="2002" box="[1112,1227,292,311]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="atiquipa">
<emphasis id="B2ECE8451F04FFDB1E5BFEEDFB1D4306" bold="true" box="[1112,1227,292,311]" italics="true" pageId="15" pageNumber="15">O. atiquipa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB18BEFE89FB1D4072" blockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB18BEFE89FB1D4072" pageId="15" pageNumber="15"> Sternite VII with one or two pairs of carinae (fig. 18C, D); metasomal segment II (♂), VSM carinae present; segment V, VM, and VL carinae well defined (figs. 20A, C, F, 21E); legs III and IV, telotarsi each with 3/3 spiniform macrosetae in pro- and retroventral rows; metasomal segments IV, paired VL stripes not contiguous with VM stripe posteriorly (fig. 11C); segment IV with three pairs of VSM macrosetae....................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB18BEFD82FB1A415B" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB18BEFD82FB1A40F8" blockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB18BEFD82FB1A40F8" pageId="15" pageNumber="15">
13. Hemispermatophore apex elongated, angle of inclination 135.18
<emphasis id="B2ECE8451F04FFDB19A5FDAFFC794048" bold="true" box="[934,943,614,633]" pageId="15" pageNumber="15">°</emphasis>
139.1
<emphasis id="B2ECE8451F04FFDB19F2FDAFFC2C4048" bold="true" box="[1009,1018,614,633]" pageId="15" pageNumber="15">°</emphasis>
(fig. 36D, E); tergite VII entirely pigmented (fig. 13G); telson vesicle (♂), dorsal surface concave (fig. 23E)...............................
<taxonomicName id="47984FD41F04FFDB1E44FD7FFB1A40F8" ID-CoL="757W3" box="[1095,1228,693,713]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="grismadoi">
<emphasis id="B2ECE8451F04FFDB1E44FD7FFB1A40F8" bold="true" box="[1095,1228,693,713]" italics="true" pageId="15" pageNumber="15">O. grismadoi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB18BEFD1BFB1A415B" blockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB18BEFD1BFB1A415B" pageId="15" pageNumber="15">
Hemispermatophore apex short, angle of inclination 138.99
<emphasis id="B2ECE8451F04FFDB19AFFD25FC6340CE" bold="true" box="[940,949,748,767]" pageId="15" pageNumber="15">°</emphasis>
144.99
<emphasis id="B2ECE8451F04FFDB1E06FD25FBD840CE" bold="true" box="[1029,1038,748,767]" pageId="15" pageNumber="15">°</emphasis>
(figs. 36I, J, 47A, B); tergite VII with paired lateral spots of pigmentation delimiting unpigmented median longitudinal stripe (fig. 13F); telson vesicle (♂), dorsal surface flat (fig. 23A)....... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B6C2CF21F04FFDB18BEFCBAFB1D466D" pageId="15" pageNumber="15">
<paragraph id="802734571F04FFDB18BEFCBAFB1D41C0" blockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB18BEFCBAFB1D41C0" pageId="15" pageNumber="15">
14. Hemispermatophore, distal crest extremely developed, extending almost entire length of lamina (fig. 47A, B); telson length/width ratio (♂), 2.652.94 (mean 5 2.78), (
<specimenCount id="969EFFDE1F04FFDB1E43FC09FB8441E6" box="[1088,1106,960,983]" pageId="15" pageNumber="15" type="female"></specimenCount>
), 2.132.37 (mean 5 2.27).............
<taxonomicName id="47984FD41F04FFDB1E55FC17FB1D41C0" ID-CoL="6SVVQ" authorityName="Ochoa &amp; Ojanguren Affilastro &amp; Mattoni &amp; Prendini" authorityYear="2011" box="[1110,1227,990,1009]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="ramirezi">
<emphasis id="B2ECE8451F04FFDB1E55FC17FB1D41C0" bold="true" box="[1110,1227,990,1009]" italics="true" pageId="15" pageNumber="15">O. ramirezi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="802734571F04FFDB18BEFC33FB1D466D" blockId="15.[701,1228,212,1117]" pageId="15" pageNumber="15">
<keyLead id="3B6997621F04FFDB18BEFC33FB1D466D" pageId="15" pageNumber="15">
Hemispermatophore, distal crest restricted to distal half of lamina (fig. 36I); telson length/ width ratio (♂), 2.873.21 (mean 5 3.07), (
<specimenCount id="969EFFDE1F04FFDB18F8FB8FFCDB466C" box="[763,781,1094,1117]" pageId="15" pageNumber="15" type="female"></specimenCount>
), 2.442.55 (mean 5 2.49)...
<taxonomicName id="47984FD41F04FFDB1E5CFB80FB1D466D" ID-CoL="6T87R" baseAuthorityName="Pocock" baseAuthorityYear="1899" box="[1119,1227,1096,1116]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="B2ECE8451F04FFDB1E5CFB80FB1D466D" bold="true" box="[1119,1227,1096,1116]" italics="true" pageId="15" pageNumber="15">O. alticola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>