626 lines
94 KiB
XML
626 lines
94 KiB
XML
<document id="A6FA99846BC9E95B605722E757A97C51" ID-DOI="10.11646/zootaxa.5523.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13934059" ID-ZooBank="E032D32F-0F67-45D6-901B-704E79D1EB10" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1728986065471" checkinUser="plazi" docAuthor="Patel, Harshil, Thackeray, Tejas, Sheth, Chintan, Khandekar, Akshay & Agarwal, Ishan" docDate="2024" docId="039B87F1DD38F35D7FCDDD11FB18F977" docLanguage="en" docName="zootaxa.5523.2.1.pdf" docOrigin="Zootaxa 5523 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calotes sinyik Patel, Thackeray, Sheth, Khandekar & Agarwal, 2024, sp. nov." docType="treatment" docUuid="4EAFC175-AE18-4A92-B9E5-DA1CAB1D607F" docUuidSource="ZooBank" docVersion="3" lastPageNumber="166" masterDocId="FFA2FF89DD3DF3527F5ADA17FFFEFFEE" masterDocTitle="A new small-sized Calotes Cuvier (Squamata: Agamidae: Draconinae) from the Subansiri river basin, Arunachal Pradesh, India" masterLastPageNumber="170" masterPageNumber="151" pageNumber="156" updateTime="1729603822902" updateUser="juliana" zenodo-license-document="CLOSED">
|
||
<mods:mods id="3BF74900D3DB785FD3AA97076E9E9F7C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B4CA1677CDBE48593F7253E5A1FD9E32">
|
||
<mods:title id="A545D96E733933B106BCFAF00E126FB7">A new small-sized Calotes Cuvier (Squamata: Agamidae: Draconinae) from the Subansiri river basin, Arunachal Pradesh, India</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E15A5E638C9A6FFEAC59E671043D0C36" type="personal">
|
||
<mods:role id="35DC53F9505B141B8742EDD73A584EB8">
|
||
<mods:roleTerm id="65FBF24A2A6C389FED1943D85FCBC4CA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DDCA420B77442FFCA741B5BD55AC339D">Patel, Harshil</mods:namePart>
|
||
<mods:nameIdentifier id="747B268EB34DF2D4D555647FDF6CEDE3" type="ORCID">0000-0002-4679-5211</mods:nameIdentifier>
|
||
<mods:affiliation id="8B2FF2CC7B33245035FEC98401F390BE">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
|
||
<mods:nameIdentifier id="CF7F5EF5DE2B017D2D647A156F39617E" type="email">harshilpatel121@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="EB21294DEF5225FCC6CDA5F04CA85109" type="personal">
|
||
<mods:role id="74448A3737984F8C5D13BE8C5ADC9D1C">
|
||
<mods:roleTerm id="8489E8C5A5EB31448C0A0DE7059B22D3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="76C3EA2BDF9BF6718B0C44F52B07B921">Thackeray, Tejas</mods:namePart>
|
||
<mods:nameIdentifier id="6DD20BE55F184C7C172D3316796F5518" type="ORCID">0000-0002-9981-8763</mods:nameIdentifier>
|
||
<mods:affiliation id="713F757FA38DAA48A1DFE9231B3B4D3A">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
|
||
<mods:nameIdentifier id="1E899439D00112E98F0583E508DDCC35" type="email">tejasthackeray@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="282C60097CF1C0B0EBB1BB2CC0C566AC" type="personal">
|
||
<mods:role id="1DDCDEA72B97FCD82D1D1BB955B8D311">
|
||
<mods:roleTerm id="438496AEBE52E762358F83B8BC0FF597">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8EFA29FB95D719E8133C1291FEBDCDC1">Sheth, Chintan</mods:namePart>
|
||
<mods:nameIdentifier id="3E7B3BD4772BDF70AB603E278D255631" type="ORCID">0000-0003-2411-9850</mods:nameIdentifier>
|
||
<mods:affiliation id="DC5D1C852A8E3288CA0E28F623191295">Wildlife Conservation Society-India, Bangalore, 560097, India & School of Humanities, Faculty of Arts and Social Sciences, The University of Sydney, NSW 2006, Australia</mods:affiliation>
|
||
<mods:nameIdentifier id="EE967172D7CCD23DDD83A588E48ADAF3" type="email">cshe0361@uni.sydney.edu.au</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="02F335D680A685AAF9F08D3C26C56EA3" type="personal">
|
||
<mods:role id="432CFDAF9E45152FA10C3165094C7CBB">
|
||
<mods:roleTerm id="3E2F5985D2EE97A72EE2B71C70971AC2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1E459715D5587109BBA71F692D5A15E1">Khandekar, Akshay</mods:namePart>
|
||
<mods:nameIdentifier id="15FAE00D3771C75FB89796991B1B25F9" type="ORCID">0000-0002-7956-089X</mods:nameIdentifier>
|
||
<mods:affiliation id="67DEE55A6E6842038C1B66D0C926CD40">Thackeray Wildlife Foundation, Mumbai, 400051, India & Department of Zoology, Shivaji University, Kolhapur, 416004, India</mods:affiliation>
|
||
<mods:nameIdentifier id="0F8CDAC08E479E80F7B29344D1399A61" type="email">akshaykhandekar555@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="BC9383BA7F7705912076B2AF577D9633" type="personal">
|
||
<mods:role id="822577CE10F5120FA91E18CE4F8FE180">
|
||
<mods:roleTerm id="58AD91E82E1C2ADE6ADDC0160AB6EA86">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="50AFB35AAB4228180090ADEC439D2D10">Agarwal, Ishan</mods:namePart>
|
||
<mods:nameIdentifier id="2093A09290863B0185FCBCFD5412235E" type="ORCID">0000-0001-9734-5379</mods:nameIdentifier>
|
||
<mods:affiliation id="893EC2AF869EB79DA32AD29581AEE941">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
|
||
<mods:nameIdentifier id="E79C06790D7CADDB3A198508566B160F" type="email">ishan.agarwal@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="2B78E67A710937B4D345228ECA857D6A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="564CEB1D046900B533D5D9239D29D9EC" type="host">
|
||
<mods:titleInfo id="96E291280877D2D65E5C795099D4E035">
|
||
<mods:title id="10EF93F9C665E3889D84EEEDB06FDC04">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="620B0E48D3C233691E19143DB66EC434">
|
||
<mods:date id="FF9CC15E294492000C34F4F95F9AA5DE">2024</mods:date>
|
||
<mods:detail id="9BA5180DB21ABE771D90F02A7D4261F5" type="pubDate">
|
||
<mods:number id="D89527D33DABAD2FF14D68E2DAF9FC1B">2024-10-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B67507E249FC31A1B390DF3E201D2483" type="volume">
|
||
<mods:number id="4E185318F1282AF601E15C36F5D13A1B">5523</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A1358F0960A8D93B232414295129BD5F" type="issue">
|
||
<mods:number id="80847FAFA54B30214496053129A10F87">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="CB4C245C4322D3A8932F1FA3143564EA" unit="page">
|
||
<mods:start id="95F1380704C7EA37B73C384D33A604DA">151</mods:start>
|
||
<mods:end id="17799416A9B416E26E349D4FAEC04C44">170</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="981EC22D9F98BDFEBCFFA7AB65B06BDB">
|
||
<mods:url id="FD525EB5BEEE2D367B3C0CE4EFC785A1">http://dx.doi.org/10.11646/zootaxa.5523.2.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="F9D2AB1B17275FB6C6420545032BA109">journal article</mods:classification>
|
||
<mods:identifier id="5AB5D3F53339DE3B40E194B14857E80F" type="DOI">10.11646/zootaxa.5523.2.1</mods:identifier>
|
||
<mods:identifier id="6491FC9AE376AA02E31F65034376C2FB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="B61516D485DA38F48380AA766DE5B3B1" type="Zenodo-Dep">13934059</mods:identifier>
|
||
<mods:identifier id="AFC5EAB2FA51488931897B93D27D44B9" type="ZooBank">E032D32F-0F67-45D6-901B-704E79D1EB10</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="039B87F1DD38F35D7FCDDD11FB18F977" LSID="urn:lsid:zoobank.org:act:4EAFC175-AE18-4A92-B9E5-DA1CAB1D607F" httpUri="http://treatment.plazi.org/id/039B87F1DD38F35D7FCDDD11FB18F977" lastPageId="15" lastPageNumber="166" pageId="5" pageNumber="156">
|
||
<subSubSection id="C328656CDD38F3577FCDDD11FE5BF8CF" box="[151,421,1798,1825]" pageId="5" pageNumber="156" type="nomenclature">
|
||
<paragraph id="8B8D36E7DD38F3577FCDDD11FE5BF8CF" blockId="5.[151,992,1798,1897]" box="[151,421,1798,1825]" pageId="5" pageNumber="156">
|
||
<heading id="D0C5818BDD38F3577FCDDD11FE5BF8CF" bold="true" box="[151,421,1798,1825]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
|
||
<emphasis id="B946EAF5DD38F3577FCDDD11FE5BF8CF" bold="true" box="[151,421,1798,1825]" pageId="5" pageNumber="156">
|
||
<taxonomicName id="4C324D64DD38F3577FCDDD11FEBEF8CF" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[151,320,1798,1825]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="156" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD38F3577FCDDD11FEBEF8CF" bold="true" box="[151,320,1798,1825]" italics="true" pageId="5" pageNumber="156">Calotes sinyik</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A275578EDD38F3577E1DDD10FE5BF8CF" box="[327,421,1799,1825]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD38F3577FCDDD3DFDEEF887" pageId="5" pageNumber="156" type="description">
|
||
<paragraph id="8B8D36E7DD38F3577FCDDD3DFC1EF8AB" blockId="5.[151,992,1798,1897]" box="[151,992,1834,1861]" pageId="5" pageNumber="156">
|
||
<uri id="FFA33AE5DD38F3577FCDDD3DFC1EF8AB" box="[151,992,1834,1861]" pageId="5" pageNumber="156">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FF940C32DD38F3577EF4DD3DFC1EF8AB" box="[430,992,1834,1861]" pageId="5" pageNumber="156">4EAFC175-AE18-4A92-B9E5-DA1CAB1D607F</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD38F3577FCDDD59FDEEF887" blockId="5.[151,992,1798,1897]" box="[151,528,1870,1897]" pageId="5" pageNumber="156">
|
||
(
|
||
<figureCitation id="13092A62DD38F3577FC5DD59FEF5F887" box="[159,267,1870,1897]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@6.[208,1379,181,1909]" captionTargetPageId="6" captionText="FIGURE 3. Calotes sinyik sp. nov.: (A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar." figureDoi="http://doi.org/10.5281/zenodo.13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="5" pageNumber="156">Figures 3</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD38F3577E42DD58FEA3F887" box="[280,349,1871,1897]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="5" pageNumber="156">4A–C</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD38F3577E32DD59FE73F887" box="[360,397,1870,1897]" captionStart="FIGURE 5" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@10.[208,1379,181,1909]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934073" httpUri="https://zenodo.org/record/13934073/files/figure.png" pageId="5" pageNumber="156">5A</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD38F3577EC3DD59FE40F886" box="[409,446,1870,1896]" captionStart="FIGURE 6" captionStartId="11.[151,250,1735,1759]" captionTargetBox="[158,1430,181,1711]" captionTargetId="figure-20@11.[158,1430,181,1711]" captionTargetPageId="11" captionText="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="5" pageNumber="156">6A</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD38F3577E93DD58FE10F887" box="[457,494,1871,1897]" captionStart="FIGURE 7" captionStartId="12.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@12.[208,1379,181,1909]" captionTargetPageId="12" captionText="FIGURE 7. Ventral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B), C. zolaiking (paratype, ESV 105); and (C) Calotes paulus (NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934079" httpUri="https://zenodo.org/record/13934079/files/figure.png" pageId="5" pageNumber="156">7A</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD38F3577EA3DD58FDF9F887" box="[505,519,1871,1897]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-20@13.[203,1384,181,1919]" captionTargetPageId="13" captionText="FIGURE 8. Calotes sinyik sp. nov. in life: (A) adult female (holotype, NRC-AA-8424), (B) juvenile (paratype, NRC-AA- 8425), (C) ventral view of adult female (holotype, NRC-AA-8424). Photos by Chintan Sheth." figureDoi="http://doi.org/10.5281/zenodo.13934083" httpUri="https://zenodo.org/record/13934083/files/figure.png" pageId="5" pageNumber="156">8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD38F35A7FCDDD87FEE1FF3B" lastPageId="8" lastPageNumber="159" pageId="5" pageNumber="156" type="materials_examined">
|
||
<paragraph id="8B8D36E7DD38F3577FCDDD87FD44F81C" blockId="5.[151,1437,1936,2034]" pageId="5" pageNumber="156">
|
||
<materialsCitation id="3B5A3CBADD38F3577FCDDD87FD44F81C" collectingDate="2023-10-07" collectionCode="AP" collectorName="Harshil Patel & Ishan Agarwal & Chintan Sheth" country="India" elevation="1270" latitude="28.2621" location="Lingbuk Riamukh" longLatPrecision="1" longitude="94.159" municipality="Payeng Circle" pageId="5" pageNumber="156" specimenCode="NRC-AA-8424" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
|
||
<emphasis id="B946EAF5DD38F3577FCDDD87FEF5F844" bold="true" box="[151,267,1936,1962]" pageId="5" pageNumber="156">
|
||
<typeStatus id="54898845DD38F3577FCDDD87FEF8F844" box="[151,262,1936,1962]" pageId="5" pageNumber="156" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DB949E9CDD38F3577E4CDD87FE3CF844" box="[278,450,1936,1962]" pageId="5" pageNumber="156">NRC-AA-8424</specimenCode>
|
||
(AP 078), adult
|
||
<specimenCount id="9D34FD6EDD38F3577DD1DD87FD24F844" box="[651,730,1936,1962]" pageId="5" pageNumber="156" type="female">female</specimenCount>
|
||
, from near
|
||
<location id="8EED603CDD38F3577C3FDD87FBCBF844" LSID="urn:lsid:plazi:treatment:039B87F1DD38F35D7FCDDD11FB18F977:8EED603CDD38F3577C3FDD87FBCBF844" box="[869,1077,1936,1962]" country="India" latitude="28.2621" longLatPrecision="1" longitude="94.159" municipality="Payeng Circle" name="Lingbuk Riamukh" pageId="5" pageNumber="156" stateProvince="Arunachal Pradesh">Lingbuk Riamukh</location>
|
||
(
|
||
<geoCoordinate id="EE065020DD38F3577B1DDD87FB2AF844" box="[1095,1236,1936,1962]" degrees="28.26210" direction="north" orientation="latitude" pageId="5" pageNumber="156" precision="1" value="28.2621">28.26210°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE065020DD38F3577BB9DD87FA92F845" box="[1251,1388,1936,1963]" degrees="94.15900" direction="east" orientation="longitude" pageId="5" pageNumber="156" precision="1" value="94.159">94.15900°E</geoCoordinate>
|
||
;
|
||
<emphasis id="B946EAF5DD38F3577A21DD86FA62F844" box="[1403,1436,1937,1962]" italics="true" pageId="5" pageNumber="156">ca.</emphasis>
|
||
<elevation id="001FD1D4DD38F3577FCDDDA3FEE7F820" box="[151,281,1972,1998]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="156" unit="m" value="1270.0">
|
||
<quantity id="4CCA9B02DD38F3577FCDDDA3FF13F820" box="[151,237,1972,1998]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="156" unit="m" value="1270.0">1270 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.),
|
||
<collectingMunicipality id="6BE9AC9DDD38F3577E69DDA3FE2AF820" box="[307,468,1972,1998]" pageId="5" pageNumber="156">Payeng Circle</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="6BE9AC9DDD38F3577EBADDA3FD09F820" box="[480,759,1972,1998]" pageId="5" pageNumber="156">Upper Subansiri District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49F6F805DD38F3577C5BDDA3FC27F820" box="[769,985,1972,1998]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="156">Arunachal Pradesh</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3257677DD38F3577CBFDDA3FBDCF820" box="[997,1058,1972,1998]" name="India" pageId="5" pageNumber="156">India</collectingCountry>
|
||
; collected by
|
||
<collectorName id="26C75331DD38F3577BE5DDA3FAAAF820" box="[1215,1364,1972,1998]" pageId="5" pageNumber="156">Harshil Patel</collectorName>
|
||
,
|
||
<collectorName id="26C75331DD38F3577A3ADDA3FF09F81C" pageId="5" pageNumber="156">Ishan Agarwal</collectorName>
|
||
and
|
||
<collectorName id="26C75331DD38F3577E74DDCFFE30F81C" box="[302,462,2008,2034]" pageId="5" pageNumber="156">Chintan Sheth</collectorName>
|
||
on
|
||
<date id="FF8C1027DD38F3577EADDDCFFD48F81C" box="[503,694,2007,2034]" pageId="5" pageNumber="156" value="2023-10-07">
|
||
<collectingDate id="EFC8E9CFDD38F3577EADDDCFFD48F81C" box="[503,694,2007,2034]" pageId="5" pageNumber="156" value="2023-10-07">
|
||
7
|
||
<superScript id="7C479BAFDD38F3577D5FDDC0FDECF80B" attach="left" box="[517,530,2007,2021]" fontSize="6" pageId="5" pageNumber="156">th</superScript>
|
||
October 2023
|
||
</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF4D666FDD3BF3547FCDDD9AFBB3F827" ID-DOI="http://doi.org/10.5281/zenodo.13934067" ID-Zenodo-Dep="13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="6" pageNumber="157" startId="6.[151,250,1933,1957]" targetBox="[208,1379,181,1909]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B8D36E7DD3BF3547FCDDD9AFBB3F827" blockId="6.[151,1436,1933,1994]" pageId="6" pageNumber="157">
|
||
<emphasis id="B946EAF5DD3BF3547FCDDD9AFEECF84B" bold="true" box="[151,274,1933,1957]" pageId="6" pageNumber="157">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C324D64DD3BF3547E42DD9AFE58F84B" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[280,422,1933,1957]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="157" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD3BF3547E42DD9AFE58F84B" box="[280,422,1933,1957]" italics="true" pageId="6" pageNumber="157">Calotes sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD3BF3547EF6DD9AFDFBF84B" bold="true" box="[428,517,1933,1957]" pageId="6" pageNumber="157">
|
||
<taxonomicNameLabel id="A275578EDD3BF3547EF6DD9AFDFEF84B" box="[428,512,1933,1957]" pageId="6" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
:
|
||
</emphasis>
|
||
(A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D666FDD3AF3557FCDDD98FB1BF81E" ID-DOI="http://doi.org/10.5281/zenodo.13934071" ID-Zenodo-Dep="13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,250,1935,1959]" targetBox="[207,1379,181,1911]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B8D36E7DD3AF3557FCDDD98FB1BF81E" blockId="7.[151,1436,1935,2032]" pageId="7" pageNumber="158">
|
||
<emphasis id="B946EAF5DD3AF3557FCDDD98FEEBF849" bold="true" box="[151,277,1935,1959]" pageId="7" pageNumber="158">FIGURE 4.</emphasis>
|
||
Head scalation of three small-bodied species
|
||
<taxonomicName id="4C324D64DD3AF3557DA8DD87FCC1F846" box="[754,831,1936,1960]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="158" phylum="Chordata" rank="genus">
|
||
<emphasis id="B946EAF5DD3AF3557DA8DD87FCC1F846" box="[754,831,1936,1960]" italics="true" pageId="7" pageNumber="158">Calotes</emphasis>
|
||
</taxonomicName>
|
||
from Northeast India: (A–C)
|
||
<taxonomicName id="4C324D64DD3AF3557B2DDD87FB2CF846" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[1143,1234,1936,1960]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD3AF3557B2DDD87FB2CF846" box="[1143,1234,1936,1960]" italics="true" pageId="7" pageNumber="158">C. sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD3AF3557B80DD87FAD2F846" bold="true" box="[1242,1324,1936,1960]" pageId="7" pageNumber="158">
|
||
<taxonomicNameLabel id="A275578EDD3AF3557B80DD87FAD2F846" box="[1242,1324,1936,1960]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype, NRC-AA-8424); (D–F)
|
||
<taxonomicName id="4C324D64DD3AF3557ED6DDA3FDF8F822" box="[396,518,1972,1996]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD3AF3557ED6DDA3FDF8F822" box="[396,518,1972,1996]" italics="true" pageId="7" pageNumber="158">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
(paratype, ESV 105); (G–I)
|
||
<taxonomicName id="4C324D64DD3AF3557C7FDDA3FC78F822" box="[805,902,1972,1996]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="158" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD3AF3557C7FDDA3FC78F822" box="[805,902,1972,1996]" italics="true" pageId="7" pageNumber="158">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
(NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36E7DD35F35A7F9DDA80FEE1FF3B" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="159">
|
||
<materialsCitation id="3B5A3CBADD35F35A7F9DDA80FEE2FF3B" collectingDate="2023-10-07" collectionCode="AP" collectorName="Harshil Patel" country="India" elevation="1270" latitude="28.26868" location="Upper Subansiri District" longLatPrecision="4" longitude="94.159" municipality="Payeng Circle" pageId="8" pageNumber="159" specimenCode="NRC-AA-8425" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
|
||
<emphasis id="B946EAF5DD35F35A7F9DDA80FEC5FF5F" bold="true" box="[199,315,151,177]" pageId="8" pageNumber="159">
|
||
<typeStatus id="54898845DD35F35A7F9DDA80FEC8FF5F" box="[199,310,151,177]" pageId="8" pageNumber="159" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DB949E9CDD35F35A7E16DA81FE06FF5F" box="[332,504,150,177]" pageId="8" pageNumber="159">NRC-AA-8425</specimenCode>
|
||
(AP 077), juvenile, same collection data as holotype except (
|
||
<geoCoordinate id="EE065020DD35F35A7A51DA80FA66FF5F" box="[1291,1432,151,177]" degrees="28.26868" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="1" value="28.26868">28.26868°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE065020DD35F35A7FCDDAADFEEAFF3B" box="[151,276,186,213]" degrees="94.1590" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="5" value="94.159">94.1590°E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD35F35A7F9DDAC8FEB3FE8A" pageId="8" pageNumber="159" type="etymology">
|
||
<paragraph id="8B8D36E7DD35F35A7F9DDAC8FEB3FE8A" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="159">
|
||
<emphasis id="B946EAF5DD35F35A7F9DDAC8FEB0FF17" bold="true" box="[199,334,223,249]" pageId="8" pageNumber="159">Etymology.</emphasis>
|
||
The specific epithet is the local name of the Subansiri River in the Tagin language, used as a noun in apposition. The Tagin are one of the major tribes of
|
||
<collectingRegion id="49F6F805DD35F35A7C47DB14FC0BFEF3" box="[797,1013,259,285]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="159">Arunachal Pradesh</collectingRegion>
|
||
and mainly inhabit Upper Subansiri District. Suggested common name is Sinyik or Subansiri dragon lizard. Agamids are commonly referred to as /sopin/ in Tagin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD35F35A7F9DDB79FD1FFC03" pageId="8" pageNumber="159" type="diagnosis">
|
||
<paragraph id="8B8D36E7DD35F35A7F9DDB79FEF2FD47" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="159">
|
||
<emphasis id="B946EAF5DD35F35A7F9DDB79FEBCFE66" bold="true" box="[199,322,366,392]" pageId="8" pageNumber="159">Diagnosis.</emphasis>
|
||
A small-sized
|
||
<taxonomicName id="4C324D64DD35F35A7EB2DB78FDC0FE67" box="[488,574,367,393]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="B946EAF5DD35F35A7EB2DB78FDC0FE67" box="[488,574,367,393]" italics="true" pageId="8" pageNumber="159">Calotes</emphasis>
|
||
</taxonomicName>
|
||
, snout to vent length up to
|
||
<quantity id="4CCA9B02DD35F35A7C28DB79FC3FFE67" box="[882,961,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="159" unit="mm" value="65.0">65 mm</quantity>
|
||
(
|
||
<emphasis id="B946EAF5DD35F35A7C88DB78FC1EFE66" box="[978,992,367,392]" italics="true" pageId="8" pageNumber="159">n</emphasis>
|
||
= 2). Body feebly compressed laterally with a weak dorsal crest; scales on top of head highly heterogeneous, keeled, those above orbit and surrounding parietal are among the largest in size; three spines on each side of head, a short one above orbit, one above tympanum and one on temporal region; dorsolateral scales of body heterogeneous in size, composed of medium-sized, weakly pointed, strongly keeled scales, intermixed with similar but slightly larger scales which are distinct on flanks, upper rows directed backwards and upwards and a few lower rows directed backwards; 54 or 55 midbody scale rows; 42 or 43 vertebral scales; a weakly developed antehumeral fold; tympanum very small in adult; tail rounded in crosssection; nine or 10 supralabials and 10–12 infralabials; lamellae bicarinate, 24 or 25 on fourth finger and 26–28 on fourth toe.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD35F35A7F9DD8A5FD1FFC03" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="159">
|
||
<emphasis id="B946EAF5DD35F35A7F9DD8A5FD6CFD23" bold="true" box="[199,658,690,717]" pageId="8" pageNumber="159">Comparisons with regional congeners.</emphasis>
|
||
The new species can be easily separated from all regional congeners except
|
||
<taxonomicName id="4C324D64DD35F35A7FB2D8C0FE71FD1F" authorityName="Biswas" authorityYear="1975" box="[232,399,727,753]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="bhutanensis">
|
||
<emphasis id="B946EAF5DD35F35A7FB2D8C0FE71FD1F" box="[232,399,727,753]" italics="true" pageId="8" pageNumber="159">C. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C324D64DD35F35A7EC4D8C0FDF4FD1F" box="[414,522,727,753]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7EC4D8C0FDF4FD1F" box="[414,522,727,753]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D64DD35F35A7D1BD8C0FD37FD1F" box="[577,713,727,753]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7D1BD8C0FD37FD1F" box="[577,713,727,753]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
by its small body size (maximum SVL up to
|
||
<quantity id="4CCA9B02DD35F35A7B96D8C1FAE2FD1F" box="[1228,1308,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="159" unit="mm" value="65.0">65 mm</quantity>
|
||
<emphasis id="B946EAF5DD35F35A7A79D8C0FA95FD1E" box="[1315,1387,727,752]" italics="true" pageId="8" pageNumber="159">versus</emphasis>
|
||
<quantity id="4CCA9B02DD35F35A7A28D8C0FF0AFCFB" metricMagnitude="-1" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.58" metricValueMin="0.9099999999999999" pageId="8" pageNumber="159" unit="mm" value="124.5" valueMax="158.0" valueMin="91.0">91– 158 mm</quantity>
|
||
) and from all regional congeners except
|
||
<taxonomicName id="4C324D64DD35F35A7DEDD8ECFCDCFCFB" box="[695,802,763,789]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7DEDD8ECFCDCFCFB" box="[695,802,763,789]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D64DD35F35A7C0CD8ECFC23FCFB" box="[854,989,763,789]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7C0CD8ECFC23FCFB" box="[854,989,763,789]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
by the presence of heterogeneous scales on the dorsolateral aspect of body, a weakly developed antehumeral fold, and a weakly developed dorsal crest. The new species is most similar to
|
||
<taxonomicName id="4C324D64DD35F35A7EB2D954FDACFCB3" box="[488,594,835,861]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7EB2D954FDACFCB3" box="[488,594,835,861]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D64DD35F35A7DDCD954FCF3FCB3" box="[646,781,835,861]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7DDCD954FCF3FCB3" box="[646,781,835,861]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
but differs from both species with respect to the following: greater number of midbody scale rows (54 or 55
|
||
<emphasis id="B946EAF5DD35F35A7D88D970FCE4FC6E" box="[722,794,871,896]" italics="true" pageId="8" pageNumber="159">versus</emphasis>
|
||
<quantity id="4CCA9B02DD35F35A7C79D970FC77FC6F" box="[803,905,871,897]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" metricValueMax="1.1684" metricValueMin="1.0668" pageId="8" pageNumber="159" unit="in" value="44.0" valueMax="46.0" valueMin="42.0">42–46 in</quantity>
|
||
<taxonomicName id="4C324D64DD35F35A7CC8D970FBFFFC6F" box="[914,1025,871,897]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7CC8D970FBFFFC6F" box="[914,1025,871,897]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="4CCA9B02DD35F35A7B66D971FB5FFC6F" box="[1084,1185,870,897]" metricMagnitude="0" metricUnit="m" metricValue="1.2827000000000002" metricValueMax="1.3208000000000002" metricValueMin="1.2446000000000002" pageId="8" pageNumber="159" unit="in" value="50.5" valueMax="52.0" valueMin="49.0">49–52 in</quantity>
|
||
<taxonomicName id="4C324D64DD35F35A7BF1D970FACBFC6F" box="[1195,1333,871,897]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7BF1D970FACBFC6F" box="[1195,1333,871,897]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
), greater number of vertebral scales (42 or 43
|
||
<emphasis id="B946EAF5DD35F35A7D77D99CFD8BFC4A" box="[557,629,907,932]" italics="true" pageId="8" pageNumber="159">versus</emphasis>
|
||
<quantity id="4CCA9B02DD35F35A7D20D99CFD22FC4B" box="[634,732,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" metricValueMax="8.128" metricValueMin="6.096" pageId="8" pageNumber="159" unit="in" value="28.0" valueMax="32.0" valueMin="24.0">24–32 in</quantity>
|
||
<taxonomicName id="4C324D64DD35F35A7DB8D99CFCB3FC4B" box="[738,845,907,933]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7DB8D99CFCB3FC4B" box="[738,845,907,933]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="4CCA9B02DD35F35A7CDBD99DFC1DFC4B" box="[897,995,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="8.89" metricValueMin="8.128" pageId="8" pageNumber="159" unit="in" value="33.5" valueMax="35.0" valueMin="32.0">32–35 in</quantity>
|
||
<taxonomicName id="4C324D64DD35F35A7CB2D99CFB91FC4B" box="[1000,1135,907,933]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7CB2D99CFB91FC4B" box="[1000,1135,907,933]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
), and in dorsolateral scales of the body being heterogeneous only in size (
|
||
<emphasis id="B946EAF5DD35F35A7DF4D9B8FD08FC26" box="[686,758,943,968]" italics="true" pageId="8" pageNumber="159">versus</emphasis>
|
||
dorsolateral scales of body heterogeneous in both size and shape in
|
||
<taxonomicName id="4C324D64DD35F35A7FA1D9C4FE98FC03" box="[251,358,979,1005]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD35F35A7FA1D9C4FE98FC03" box="[251,358,979,1005]" italics="true" pageId="8" pageNumber="159">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D64DD35F35A7EC6D9C4FDDAFC03" box="[412,548,979,1005]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD35F35A7EC6D9C4FDDAFC03" box="[412,548,979,1005]" italics="true" pageId="8" pageNumber="159">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
) (see
|
||
<figureCitation id="13092A62DD35F35A7D32D9C4FD2BFC03" box="[616,725,978,1005]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="10.[151,250,1933,1957]" captionStartId-1="11.[151,250,1735,1759]" captionStartId-2="12.[151,250,1933,1957]" captionTargetBox-0="[208,1379,181,1909]" captionTargetBox-1="[158,1430,181,1711]" captionTargetBox-2="[208,1379,181,1909]" captionTargetId-0="figure-30@10.[208,1379,181,1909]" captionTargetId-1="figure-20@11.[158,1430,181,1711]" captionTargetId-2="figure-30@12.[208,1379,181,1909]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 5. Dorsal scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." captionText-1="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." captionText-2="FIGURE 7. Ventral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B), C. zolaiking (paratype, ESV 105); and (C) Calotes paulus (NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi-0="http://doi.org/10.5281/zenodo.13934073" figureDoi-1="http://doi.org/10.5281/zenodo.13934077" figureDoi-2="http://doi.org/10.5281/zenodo.13934079" httpUri-0="https://zenodo.org/record/13934073/files/figure.png" httpUri-1="https://zenodo.org/record/13934077/files/figure.png" httpUri-2="https://zenodo.org/record/13934079/files/figure.png" pageId="8" pageNumber="159">Figs. 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD35F35D7F9DD9E1FCDFFCB3" lastPageId="15" lastPageNumber="166" pageId="8" pageNumber="159" type="description">
|
||
<paragraph id="8B8D36E7DD35F35B7F9DD9E1FDD0FABB" blockId="8.[151,1437,150,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="160" pageId="8" pageNumber="159">
|
||
<emphasis id="B946EAF5DD35F35A7F9DD9E1FDE4FBFF" bold="true" box="[199,538,1014,1041]" pageId="8" pageNumber="159">
|
||
Description of the
|
||
<typeStatus id="54898845DD35F35A7EF6D9E0FDEBFBFF" box="[428,533,1015,1041]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult female in generally good condition except tail bent toward left in a sigmoid manner; left and right manus slightly adpressed with third and fourth fingers curved; second, third and fourth toe of the right pes curved; eyes sunken, orbital spine on left side and tips of temporal spines on both sides curved; a
|
||
<quantity id="4CCA9B02DD35F35A7E52DE74FEA1FB93" box="[264,351,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="159" unit="mm" value="7.8">7.8 mm</quantity>
|
||
long incision in sternal region for tissue collection (
|
||
<figureCitation id="13092A62DD35F35A7CE9DE74FBC6FB93" box="[947,1080,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@6.[208,1379,181,1909]" captionTargetPageId="6" captionText="FIGURE 3. Calotes sinyik sp. nov.: (A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar." figureDoi="http://doi.org/10.5281/zenodo.13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="8" pageNumber="159">Fig. 3A, 3B</figureCitation>
|
||
)—all artefacts of preservation. Mensural and meristic data is summarised at the end of the description. SVL
|
||
<quantity id="4CCA9B02DD35F35A7B52DE90FB8EFB4F" box="[1032,1136,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="6.459999999999999" pageId="8" pageNumber="159" unit="mm" value="64.6">64.6 mm</quantity>
|
||
, head relatively long (HL/ SVL 0.33), broad (HW/HL 0.71), not depressed (HD/HL 0.55), noticeably broader than neck (
|
||
<figureCitation id="13092A62DD35F35A7BE7DEBCFAB8FB2B" box="[1213,1350,1195,1221]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4A, 4B</figureCitation>
|
||
). Snout short (SE/HL 0.39), longer than orbit diameter (OD/SE 0.67). Orbit large (OD/HL 0.26); pupil round, eyelids covered with small rounded scales, a single row of scales bordering eyelids slightly elongate, eight supraciliaries on each side, slightly elongate, subimbricate, slightly protruding laterally on supraorbital ridge, similar in size. Snout obtusely pointed; rostral wider (
|
||
<quantity id="4CCA9B02DD35F35A7D5ADF2CFDA7FABB" box="[512,601,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="159" unit="mm" value="2.3">2.3 mm</quantity>
|
||
) than deep (
|
||
<quantity id="4CCA9B02DD35F35A7DBCDF2CFCBFFABB" box="[742,833,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="159" unit="mm" value="1.1">1.1 mm</quantity>
|
||
), contacted laterally by first supralabial, an elongated large prenasal, and two large scales dorsally. Canthus rostralis and supraciliary edge sharp. Nostril circular, oriented slightly downwards, laterally positioned and placed at centre of a large, undivided, protrudent nasal plate (
|
||
<figureCitation id="13092A62DD35F35A7A35DF94FF45FA2F" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4C</figureCitation>
|
||
), which is bordered by eight scales on both sides, including one prenasal, two supranasal, three postnasals, and two subnasals on both sides; separated on both sides from rostral and first supralabial by prenasal and first subnasal. Supralabials ten, roughly rectangular, more or less equal sized, posterior most being longest. Supralabials separated from the border of orbit by three rows of scales at midorbital position; middle row distinctly enlarged consisting of equal sized, roughly rectangular scales; scales on lower and upper rows heterogeneous in shape and size. Loreal region concave, scales of loreal region heterogeneous in shape and size, weakly tuberculate. Scales on postorbital and temporal region heterogeneous in shape and size, subimbricate, tuberculate; a row of four much larger longitudinally keeled scales, second longest and first widest, extending from posterior midorbital region to just before scales bordering the supra-tympanic spine (
|
||
<figureCitation id="13092A62DD35F35A7C50DCD0FC9BF90F" box="[778,869,1735,1761]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4C</figureCitation>
|
||
). Tympanum small in size (TD
|
||
<quantity id="4CCA9B02DD35F35A7B89DCD0FAD1F90F" box="[1235,1327,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="159" unit="mm" value="1.1">1.1 mm</quantity>
|
||
), covered with a single scale (
|
||
<figureCitation id="13092A62DD35F35A7E28DCFCFE35F8EB" box="[370,459,1771,1797]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4C</figureCitation>
|
||
). Canthal scales enlarged, overlapping, keeled or weakly tuberculate, becoming slightly larger along supraciliaries. Scales on dorsal surface of snout and forehead heterogeneous in shape and size (
|
||
<figureCitation id="13092A62DD35F35A7A6DDD18FA6EF8C7" box="[1335,1424,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4A</figureCitation>
|
||
), smaller scales weakly tuberculate, larger with distinct keels except those on internarial region which are roughly rounded, uniformly large and weakly tuberculate. Supraorbitals increase in size from supraciliaries to inner edge of orbit, of which enlarged scales follow curvature of orbits posterolaterally, larger scales are keeled and smaller scales tuberculate with one or two tubercles on each scale, forming two or three rows adjacent to supraciliaries (
|
||
<figureCitation id="13092A62DD35F35A7A6FDD88FA6EF857" box="[1333,1424,1951,1977]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="8" pageNumber="159">Fig. 4A</figureCitation>
|
||
). Two small scale rows divide enlarged scales of inner orbits at narrowest point of frontal. Scales on occipital region heterogeneous in shape and size; smaller scales weakly tuberculate, larger ones with keels. Parietal scale roughly rectangular, with a pineal eye (
|
||
<figureCitation id="13092A62DD34F35B7EB0DAACFDBDFF3B" box="[490,579,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="9" pageNumber="160">Fig. 4A</figureCitation>
|
||
). Parietal scale bordered laterally by two or three enlarged, roughly elongated keeled scales, with much smaller scales anteriorly and posteriorly. Single temporal spine on each side, as long as the nuchal spines; very short orbital spine and a supra-tympanic spine, as long as the temporal spine, also present on each side (
|
||
<figureCitation id="13092A62DD34F35B7E6FDB30FE33FEAF" box="[309,461,295,321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="9" pageNumber="160">Figs. 4A, 4C</figureCitation>
|
||
). Mental subtriangular, wider (
|
||
<quantity id="4CCA9B02DD34F35B7C76DB30FC7BFEAF" box="[812,901,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="160" unit="mm" value="1.8">1.8 mm</quantity>
|
||
) than long (
|
||
<quantity id="4CCA9B02DD34F35B7B4BDB30FB90FEAF" box="[1041,1134,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="160" unit="mm" value="1.3">1.3 mm</quantity>
|
||
), bordered posterolaterally by a row of five elongated postmental chin shields on either side, gradually changing shape and size posteriorly to blend with surrounding rows of gular scales (
|
||
<figureCitation id="13092A62DD34F35B7DE0DB78FCEAFE67" box="[698,788,367,393]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="9" pageNumber="160">Fig. 4B</figureCitation>
|
||
). Infralabials roughly rectangular, more or less similar in size, posterior most being largest, ten on the left side and 11 on the right side. First pair of postmental chin shields in contact laterally with first infralabials, remaining shields separated from infralabials by one or two rows of small gular scales. Posterior gular region with considerably enlarged, flat, strongly keeled, pointed, imbricate scales, those on jowls slightly larger in size; anterior gular scales smaller, rounded, weakly keeled and subimbricate; all gular scales directed posteromedially except a few median rows which are directed posteriorly. No distinct gular pouch present, transverse gular fold absent (
|
||
<figureCitation id="13092A62DD34F35B7D1BD850FD65FD8F" box="[577,667,583,609]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="9" pageNumber="160">Fig. 4B</figureCitation>
|
||
). Ventral scales similar to those on posterior gular region, enlarged, flat, strongly keeled, pointed, imbricate, homogenous in shape but heterogeneous in size, generally increasing in size laterally; arranged in regular longitudinal rows that are directed posteriorly, those on lateral aspect are directed posteromedially (
|
||
<figureCitation id="13092A62DD34F35B7E06D8A4FE3FFD23" box="[348,449,691,717]" captionStart="FIGURE 6" captionStartId="11.[151,250,1735,1759]" captionTargetBox="[158,1430,181,1711]" captionTargetId="figure-20@11.[158,1430,181,1711]" captionTargetPageId="11" captionText="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="9" pageNumber="160">Figs. 6A</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD34F35B7E97D8A4FE0DFD23" box="[461,499,691,717]" captionStart="FIGURE 7" captionStartId="12.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@12.[208,1379,181,1909]" captionTargetPageId="12" captionText="FIGURE 7. Ventral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B), C. zolaiking (paratype, ESV 105); and (C) Calotes paulus (NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934079" httpUri="https://zenodo.org/record/13934079/files/figure.png" pageId="9" pageNumber="160">7A</figureCitation>
|
||
). Approximately 55 scales around midbody. Nuchal crest most prominent on neck, composed of nine or ten long, erect, laterally compressed, spines; remaining scales in vertebral row slightly enlarged relative to adjacent dorsal scale rows, much more flattened than spines in the nuchal crest (
|
||
<figureCitation id="13092A62DD34F35B7BE1D8ECFAEAFCFB" box="[1211,1300,763,789]" captionStart="FIGURE 4" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[207,1379,181,1911]" captionTargetId="figure-20@7.[207,1379,181,1911]" captionTargetPageId="7" captionText="FIGURE 4. Head scalation of three small-bodied species Calotes from Northeast India: (A–C) C. sinyik sp. nov. (holotype, NRC-AA-8424); (D–F) C. zolaiking (paratype, ESV 105); (G–I) C. paulus (NCBS AQ197); A,D,G: dorsal view; B,E,H: ventral view; C,F,I: lateral view. Scale bars 10 mm; photos by Akshay Khandekar (A–F) and Tarun Karmakar (G–I)." figureDoi="http://doi.org/10.5281/zenodo.13934071" httpUri="https://zenodo.org/record/13934071/files/figure.png" pageId="9" pageNumber="160">Fig. 4C</figureCitation>
|
||
) but with a pronounced median keel providing a serrated appearance in profile (
|
||
<figureCitation id="13092A62DD34F35B7CCFD908FC11FCD7" box="[917,1007,799,825]" captionStart="FIGURE 6" captionStartId="11.[151,250,1735,1759]" captionTargetBox="[158,1430,181,1711]" captionTargetId="figure-20@11.[158,1430,181,1711]" captionTargetPageId="11" captionText="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="9" pageNumber="160">Fig. 6A</figureCitation>
|
||
); 43 middorsal scales from first raised nuchal scale to above level of cloaca. Dorsal scales rectangular, heterogeneous in size on dorsolateral aspect; all scales with a moderate median keel, slightly pointed, roughly arranged in transverse rows (
|
||
<figureCitation id="13092A62DD34F35B7BCFD970FB07FC6F" box="[1173,1273,870,897]" captionStart="FIGURE 5" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@10.[208,1379,181,1909]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934073" httpUri="https://zenodo.org/record/13934073/files/figure.png" pageId="9" pageNumber="160">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD34F35B7A5FD970FAD4FC6F" box="[1285,1322,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1735,1759]" captionTargetBox="[158,1430,181,1711]" captionTargetId="figure-20@11.[158,1430,181,1711]" captionTargetPageId="11" captionText="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="9" pageNumber="160">6A</figureCitation>
|
||
); keels on those of upper flanks mostly oriented obliquely upward, horizontal or obliquely upward on midflanks and horizontal on lower flanks (
|
||
<figureCitation id="13092A62DD34F35B7E0CD9B8FE4FFC27" box="[342,433,943,969]" captionStart="FIGURE 6" captionStartId="11.[151,250,1735,1759]" captionTargetBox="[158,1430,181,1711]" captionTargetId="figure-20@11.[158,1430,181,1711]" captionTargetPageId="11" captionText="FIGURE 6. Lateral scalation on trunk of three small-bodied species Calotes from Northeast India: (A) C. sinyik sp. nov. (holotype, NRC-AA-8424); (B) C. zolaiking (paratype, ESV 105); and (C) C. paulus (NCBSAQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C)." figureDoi="http://doi.org/10.5281/zenodo.13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="9" pageNumber="160">Fig. 6A</figureCitation>
|
||
); scales bordering middorsal vertebral scales pointed with weak keels. A shallow oblique skin fold in front, and curving around anterior forelimb insertion. Scales of forelimbs and ventral hindlimbs form approximate rows, those on dorsal hindlimbs do not form regular rows and are heterogeneous in size. Dorsal scales larger than ventral scales on forelimbs and hindlimbs, all moderately keeled dorsally, very weakly keeled ventrally, scales on the posterior side of the thigh much smaller, weekly keeled and pointed. Forelimbs long ((UAL +LAL+HaL)/SVL 0.55); hindlimbs considerably long ((ULL+LLL+FoL)/SVL 0.82). Digits slender, elongate and ending in a strong, slightly curved claw. Lamellae entire, bicarinate; 25 on fourth finger, 28 on fourth toe. Relative finger lengths in mm: IV (10.1)> III (9.5)> II (7.4)> V (5.9)> I (4.6) and relative toe lengths: IV (14.3)> III (11.2)> V (8.7)> II (7.4)> I (4.9). Tail entire, rounded, more than twice head-body length (TL/SVL 2.35), not swollen at base (
|
||
<figureCitation id="13092A62DD34F35B7FAFDEE4FE77FAE3" box="[245,393,1267,1293]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@6.[208,1379,181,1909]" captionTargetPageId="6" captionText="FIGURE 3. Calotes sinyik sp. nov.: (A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar." figureDoi="http://doi.org/10.5281/zenodo.13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="9" pageNumber="160">Figs. 3A, 3B</figureCitation>
|
||
); uniformly covered with similar sized, strongly keeled, weakly pointed, regularly arranged, posteriorly directed imbricate scales, except for the tip, which is covered with six rows of small, strongly keeled, laterally compressed, erected scales.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD34F35B7F9DDF48FDDBFA0B" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="160">
|
||
Mensural (in mm) and meristic data for the
|
||
<typeStatus id="54898845DD34F35B7DEBDF48FCE9FA97" box="[689,791,1375,1401]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DB949E9CDD34F35B7C78DF48FC6FFA97" box="[802,913,1375,1401]" collectionCode="SVL" pageId="9" pageNumber="160">SVL 64.6</specimenCode>
|
||
; TrL 29.5; TL 152; TaD 6.1; TW 7.6; HL 21.2; HW 15.1; HD 11.6; SE 8.3; NE 4.4; EE 5.7;
|
||
<specimenCode id="DB949E9CDD34F35B7DDCDF94FD31FA73" box="[646,719,1410,1437]" collectionCode="IO" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="9" pageNumber="160">IO 9.5</specimenCode>
|
||
;
|
||
<specimenCode id="DB949E9CDD34F35B7D80DF94FCC3FA73" box="[730,829,1411,1437]" collectionCode="SFI" country="Slovenia" httpUri="http://grbio.org/cool/bbss-z5bi" name="Slovenian Forestry Institute" pageId="9" pageNumber="160">SFI 23.4</specimenCode>
|
||
;
|
||
<collectionCode id="ED23AE22DD34F35B7C12DF94FC7FFA73" box="[840,897,1411,1437]" pageId="9" pageNumber="160">UAL</collectionCode>
|
||
11.0; LAL 11.1; HaL 13.3; 4FL 10.1;
|
||
<specimenCode id="DB949E9CDD34F35B7A72DF94FA69FA73" box="[1320,1431,1411,1437]" collectionCode="ULL" pageId="9" pageNumber="160">ULL 14.1</specimenCode>
|
||
; LLL 16.1; FOL 23.3; 4TL 14.3; TD 1.1; OD 5.6; SnS 10; HeadSTr 13;
|
||
<specimenCode id="DB949E9CDD34F35B7CE5DFB0FC07FA2F" box="[959,1017,1447,1473]" collectionCode="CS" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="9" pageNumber="160">CS 7</specimenCode>
|
||
; SL (L/
|
||
<collectionCode id="ED23AE22DD34F35B7B0BDFB0FB9BFA2F" box="[1105,1125,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="160">R</collectionCode>
|
||
) 10/10; IL (L/
|
||
<collectionCode id="ED23AE22DD34F35B7A5DDFB0FAE5FA2F" box="[1287,1307,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="160">R</collectionCode>
|
||
) 10/11; VS 43; MSR 55; 4Flam 25; 4TLam 28.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD34F35B7F9DDFF9FE19F833" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="160">
|
||
<emphasis id="B946EAF5DD34F35B7F9DDFF9FDC5F9E6" bold="true" box="[199,571,1518,1544]" pageId="9" pageNumber="160">
|
||
Coloration in life (
|
||
<figureCitation id="13092A62DD34F35B7EF0DFF9FDD2F9E6" box="[426,556,1518,1544]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-20@13.[203,1384,181,1919]" captionTargetPageId="13" captionText="FIGURE 8. Calotes sinyik sp. nov. in life: (A) adult female (holotype, NRC-AA-8424), (B) juvenile (paratype, NRC-AA- 8425), (C) ventral view of adult female (holotype, NRC-AA-8424). Photos by Chintan Sheth." figureDoi="http://doi.org/10.5281/zenodo.13934083" httpUri="https://zenodo.org/record/13934083/files/figure.png" pageId="9" pageNumber="160">Fig. 8A, C</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Coloration in life variable, primarily in mottled shades of light and darker brown with black variegation dorsally, tip of some scales on flank white, lower flanks lighter in colour. Forelimbs and hindlimbs darker than dorsum, with buff markings on the back side of upper arms and thighs, forelimbs with a whitish patch on the elbow. Region below orbit and lateral surface of snout and head with indistinct narrow light and dark brown radial stripes extending from eye, upper and lower lips greenish with white and brown mottling, eyelids light brown, orbital spine lemon green in colouration. Tail brown with faint light and darker brown bands. Venter with numerous dark brown or grey streaks and spots, a mid-ventral line runs from behind throat to above vent Ventral scales yellow on middle of throat and pectoral region; light brown from centre of belly to vent; sides of throat with dark variegations and orange scales; sides of belly light brown to grey with a light purple tinge. Several scales on the belly have a white tip. While photographing this individual it initially appeared dark greenish brown dorsally with green labials and the characteristic black or dark brown variegation. However, within a six seconds the lizard changed to a light brown shade with dark variegations and several seconds later to a dark brown colouration. This shows that the individual was capable of changing colour to suit its dark surroundings. The juvenile however, did not exhibit this behaviour.
|
||
</paragraph>
|
||
<caption id="DF4D666FDD37F3587FCDDD9AFD26F803" ID-DOI="http://doi.org/10.5281/zenodo.13934073" ID-Zenodo-Dep="13934073" httpUri="https://zenodo.org/record/13934073/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1933,1957]" targetBox="[208,1379,181,1909]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8B8D36E7DD37F3587FCDDD9AFD26F803" blockId="10.[151,1437,1933,2029]" pageId="10" pageNumber="161">
|
||
<emphasis id="B946EAF5DD37F3587FCDDD9AFEE8F84B" bold="true" box="[151,278,1933,1957]" pageId="10" pageNumber="161">FIGURE 5.</emphasis>
|
||
Dorsal scalation on trunk of three small-bodied species
|
||
<taxonomicName id="4C324D64DD37F3587C2BDD9AFC40F84B" box="[881,958,1933,1957]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="genus">
|
||
<emphasis id="B946EAF5DD37F3587C2BDD9AFC40F84B" box="[881,958,1933,1957]" italics="true" pageId="10" pageNumber="161">Calotes</emphasis>
|
||
</taxonomicName>
|
||
from Northeast India: (A)
|
||
<taxonomicName id="4C324D64DD37F3587BB8DD9AFAC0F84B" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[1250,1342,1933,1957]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD37F3587BB8DD9AFAC0F84B" box="[1250,1342,1933,1957]" italics="true" pageId="10" pageNumber="161">C. sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD37F3587A12DD9AFA62F84B" bold="true" box="[1352,1436,1933,1957]" pageId="10" pageNumber="161">
|
||
<taxonomicNameLabel id="A275578EDD37F3587A12DD9AFA62F84B" box="[1352,1436,1933,1957]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype, NRC-AA-8424); (B)
|
||
<taxonomicName id="4C324D64DD37F3587E86DDA6FDA8F827" box="[476,598,1969,1993]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD37F3587E86DDA6FDA8F827" box="[476,598,1969,1993]" italics="true" pageId="10" pageNumber="161">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
(paratype, ESV 105); and (C)
|
||
<taxonomicName id="4C324D64DD37F3587CDDDDA6FC16F827" box="[903,1000,1969,1993]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD37F3587CDDDDA6FC16F827" box="[903,1000,1969,1993]" italics="true" pageId="10" pageNumber="161">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
(NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D666FDD36F3597FCDDCD0FD26F8C9" ID-DOI="http://doi.org/10.5281/zenodo.13934077" ID-Zenodo-Dep="13934077" httpUri="https://zenodo.org/record/13934077/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1735,1759]" targetBox="[158,1430,181,1711]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8B8D36E7DD36F3597FCDDCD0FD26F8C9" blockId="11.[151,1437,1735,1831]" pageId="11" pageNumber="162">
|
||
<emphasis id="B946EAF5DD36F3597FCDDCD0FEE8F931" bold="true" box="[151,278,1735,1759]" pageId="11" pageNumber="162">FIGURE 6.</emphasis>
|
||
Lateral scalation on trunk of three small-bodied species
|
||
<taxonomicName id="4C324D64DD36F3597C29DCD0FC3EF931" box="[883,960,1735,1759]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="genus">
|
||
<emphasis id="B946EAF5DD36F3597C29DCD0FC3EF931" box="[883,960,1735,1759]" italics="true" pageId="11" pageNumber="162">Calotes</emphasis>
|
||
</taxonomicName>
|
||
from Northeast India: (A)
|
||
<taxonomicName id="4C324D64DD36F3597BB8DCDFFAC0F931" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[1250,1342,1735,1759]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD36F3597BB8DCDFFAC0F931" box="[1250,1342,1735,1759]" italics="true" pageId="11" pageNumber="162">C. sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD36F3597A12DCD0FA62F931" bold="true" box="[1352,1436,1735,1759]" pageId="11" pageNumber="162">
|
||
<taxonomicNameLabel id="A275578EDD36F3597A12DCD0FA62F931" box="[1352,1436,1735,1759]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype, NRC-AA-8424); (B)
|
||
<taxonomicName id="4C324D64DD36F3597E86DCFBFDA8F8ED" box="[476,598,1771,1795]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD36F3597E86DCFBFDA8F8ED" box="[476,598,1771,1795]" italics="true" pageId="11" pageNumber="162">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
(paratype, ESV 105); and (C)
|
||
<taxonomicName id="4C324D64DD36F3597CDDDCFBFC16F8ED" box="[903,1000,1771,1795]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="162" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD36F3597CDDDCFBFC16F8ED" box="[903,1000,1771,1795]" italics="true" pageId="11" pageNumber="162">C. paulus</emphasis>
|
||
</taxonomicName>
|
||
(NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D666FDD31F35E7FCDDD9AFCDCF803" ID-DOI="http://doi.org/10.5281/zenodo.13934079" ID-Zenodo-Dep="13934079" httpUri="https://zenodo.org/record/13934079/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1933,1957]" targetBox="[208,1379,181,1909]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8B8D36E7DD31F35E7FCDDD9AFCDCF803" blockId="12.[151,1436,1933,2029]" pageId="12" pageNumber="163">
|
||
<emphasis id="B946EAF5DD31F35E7FCDDD9AFEE8F84B" bold="true" box="[151,278,1933,1957]" pageId="12" pageNumber="163">FIGURE 7.</emphasis>
|
||
Ventral scalation on trunk of three small-bodied species
|
||
<taxonomicName id="4C324D64DD31F35E7C29DD9AFC3EF84B" box="[883,960,1933,1957]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="163" phylum="Chordata" rank="genus">
|
||
<emphasis id="B946EAF5DD31F35E7C29DD9AFC3EF84B" box="[883,960,1933,1957]" italics="true" pageId="12" pageNumber="163">Calotes</emphasis>
|
||
</taxonomicName>
|
||
from Northeast India: (A)
|
||
<taxonomicName id="4C324D64DD31F35E7BB9DD9AFAC0F84B" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[1251,1342,1933,1957]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="163" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD31F35E7BB9DD9AFAC0F84B" box="[1251,1342,1933,1957]" italics="true" pageId="12" pageNumber="163">C. sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD31F35E7A12DD9AFA62F84B" bold="true" box="[1352,1436,1933,1957]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="A275578EDD31F35E7A12DD9AFA62F84B" box="[1352,1436,1933,1957]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype, NRC-AA-8424); (B),
|
||
<taxonomicName id="4C324D64DD31F35E7EBFDDA6FDA1F827" box="[485,607,1969,1993]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="163" phylum="Chordata" rank="species" species="zolaiking">
|
||
<emphasis id="B946EAF5DD31F35E7EBFDDA6FDA1F827" box="[485,607,1969,1993]" italics="true" pageId="12" pageNumber="163">C. zolaiking</emphasis>
|
||
</taxonomicName>
|
||
(paratype, ESV 105); and (C)
|
||
<taxonomicName id="4C324D64DD31F35E7CCFDDA6FBD2F827" box="[917,1068,1969,1993]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="163" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B946EAF5DD31F35E7CCFDDA6FBD2F827" box="[917,1068,1969,1993]" italics="true" pageId="12" pageNumber="163">Calotes paulus</emphasis>
|
||
</taxonomicName>
|
||
(NCBS AQ197). Scale bars 10 mm; photos by Akshay Khandekar (A & B) and Tarun Karmakar (C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D666FDD30F35F7FCDDD80FBB3F83D" ID-DOI="http://doi.org/10.5281/zenodo.13934083" ID-Zenodo-Dep="13934083" httpUri="https://zenodo.org/record/13934083/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1943,1967]" targetBox="[203,1384,181,1919]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8B8D36E7DD30F35F7FCDDD80FBB3F83D" blockId="13.[151,1437,1943,2003]" pageId="13" pageNumber="164">
|
||
<emphasis id="B946EAF5DD30F35F7FCDDD80FEEBF841" bold="true" box="[151,277,1943,1967]" pageId="13" pageNumber="164">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C324D64DD30F35F7E45DD80FE4EF841" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[287,432,1943,1967]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="164" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD30F35F7E45DD80FE4EF841" box="[287,432,1943,1967]" italics="true" pageId="13" pageNumber="164">Calotes sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD30F35F7EE3DD80FDF2F841" bold="true" box="[441,524,1943,1967]" pageId="13" pageNumber="164">
|
||
<taxonomicNameLabel id="A275578EDD30F35F7EE3DD80FDF2F841" box="[441,524,1943,1967]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life: (A) adult female (holotype, NRC-AA-8424), (B) juvenile (paratype, NRC-AA-8425), (C) ventral view of adult female (holotype, NRC-AA-8424). Photos by Chintan Sheth.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D666FDD33F35C7FCDDD80FEBCF819" ID-DOI="http://doi.org/10.5281/zenodo.13934087" ID-Zenodo-Dep="13934087" httpUri="https://zenodo.org/record/13934087/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1943,1967]" targetBox="[203,1384,181,1919]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8B8D36E7DD33F35C7FCDDD80FEBCF819" blockId="14.[151,1437,1943,2039]" pageId="14" pageNumber="165">
|
||
<emphasis id="B946EAF5DD33F35C7FCDDD80FEEDF841" bold="true" box="[151,275,1943,1967]" pageId="14" pageNumber="165">FIGURE 9.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C324D64DD33F35C7EDDDD80FDE8F841" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[391,534,1943,1967]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD33F35C7EDDDD80FDE8F841" box="[391,534,1943,1967]" italics="true" pageId="14" pageNumber="165">Calotes sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD33F35C7D47DD80FD90F841" bold="true" box="[541,622,1943,1967]" pageId="14" pageNumber="165">
|
||
<taxonomicNameLabel id="A275578EDD33F35C7D47DD80FD90F841" box="[541,622,1943,1967]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at the type locality: (A) general view of the hilly forest near Lingbuk Riamukh, Upper Subansiri, Arunachal Pradesh, and (B) microhabitat from where individuals of the new species were collected. Photos by Harshil Patel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36E7DD32F35D7F9DDA81FD8DFF17" blockId="15.[151,1437,150,1689]" pageId="15" pageNumber="166">
|
||
<emphasis id="B946EAF5DD32F35D7F9DDA81FD5AFF5E" bold="true" box="[199,676,150,177]" pageId="15" pageNumber="166">
|
||
Coloration in preservative (
|
||
<figureCitation id="13092A62DD32F35D7D54DA81FD6AFF5F" box="[526,660,150,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@6.[208,1379,181,1909]" captionTargetPageId="6" captionText="FIGURE 3. Calotes sinyik sp. nov.: (A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar." figureDoi="http://doi.org/10.5281/zenodo.13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="15" pageNumber="166">Fig. 3A, 3B</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Coloration in preservation more or less similar to colouration in life but faded and predominantly brownish with upper arm and thighs slightly bluish in shade. Transverse stripes on body, limbs and tail are darker than in life.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD32F35D7F9DDB15FCDFFCB3" blockId="15.[151,1437,150,1689]" pageId="15" pageNumber="166">
|
||
<emphasis id="B946EAF5DD32F35D7F9DDB15FE3BFEF2" bold="true" box="[199,453,258,284]" pageId="15" pageNumber="166">
|
||
Variation (
|
||
<figureCitation id="13092A62DD32F35D7E1FDB15FE62FEF2" box="[325,412,258,284]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1957]" captionTargetBox="[208,1379,181,1909]" captionTargetId="figure-30@6.[208,1379,181,1909]" captionTargetPageId="6" captionText="FIGURE 3. Calotes sinyik sp. nov.: (A) dorsal view, (B) ventral view of adult female holotype (NRC-AA-8424) and (C) dorsal view of juvenile paratype (NRC-AA-8425). Scale bars 10 mm; photos by Akshay Khandekar." figureDoi="http://doi.org/10.5281/zenodo.13934067" httpUri="https://zenodo.org/record/13934067/files/figure.png" pageId="15" pageNumber="166">Fig. 3C</figureCitation>
|
||
,
|
||
<figureCitation id="13092A62DD32F35D7EFDDB15FE48FEF2" box="[423,438,258,284]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-20@13.[203,1384,181,1919]" captionTargetPageId="13" captionText="FIGURE 8. Calotes sinyik sp. nov. in life: (A) adult female (holotype, NRC-AA-8424), (B) juvenile (paratype, NRC-AA- 8425), (C) ventral view of adult female (holotype, NRC-AA-8424). Photos by Chintan Sheth." figureDoi="http://doi.org/10.5281/zenodo.13934083" httpUri="https://zenodo.org/record/13934083/files/figure.png" pageId="15" pageNumber="166">8</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54898845DD32F35D7EA0DB14FDA4FEF3" box="[506,602,259,285]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
|
||
(juvenile, NRC-AA-8425), agrees with the
|
||
<typeStatus id="54898845DD32F35D7B61DB14FB63FEF3" box="[1083,1181,259,285]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
|
||
in general morphology and scalation with following exceptions: nine SL on both sides, IL 11 on the left side and 12 on the right side; 42 vertebral scales; approximately 54 scales around midbody; 24 lamellae under fourth finger and 26 under fourth toe; 11 scales between anterior supraciliaries. The live colouration is unlike the
|
||
<typeStatus id="54898845DD32F35D7B74DB78FB6EFE67" box="[1070,1168,367,393]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
|
||
with green, brown and blue coloured variegated markings on head and dorsum (
|
||
<figureCitation id="13092A62DD32F35D7C48DB84FC97FE43" box="[786,873,403,429]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-20@13.[203,1384,181,1919]" captionTargetPageId="13" captionText="FIGURE 8. Calotes sinyik sp. nov. in life: (A) adult female (holotype, NRC-AA-8424), (B) juvenile (paratype, NRC-AA- 8425), (C) ventral view of adult female (holotype, NRC-AA-8424). Photos by Chintan Sheth." figureDoi="http://doi.org/10.5281/zenodo.13934083" httpUri="https://zenodo.org/record/13934083/files/figure.png" pageId="15" pageNumber="166">Fig. 8B</figureCitation>
|
||
). Dorsal scales light green or brown, brown scales have a fine brown peppering over a cream colour. Around nine brown stripes (1–3 scales broad) emerge radially from the orbit spaced apart by 1–4 scales. Scales from the upper labials to the sub-orbital edge are lime green with light edges. Dorso-anterior portion of the head mostly green with brown mottling, two dark greenish-brown stripes present on the dorsal scales of the head in line with the orbits. The distal portion of the head largely brown. Dorsal surface of the body has green, blue and brown scale colouration. Brown scales arranged in large blotches or diamonds spaced apart by one or two green or blue scales. This continues along the tail which is brown distally. A few dorsal scales are tipped white. An apical notch or pore present on all scales. Belly scales mostly cream coloured peppered with brown. Legs mostly with green and brown scales, except at the joint – forelimb joints with white scales and hindlimb joints with buff coloured scales. Further variation can be found in the mensural (in mm) and meristic characters in
|
||
<typeStatus id="54898845DD32F35D7ECAD8ECFE0EFCFB" box="[400,496,763,789]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
|
||
documented here: SVL 26.5; TrL 9.9; TL 53.0; TaD 2.1; TW 2.1; HL 9.0; HW 6.8; HD 4.8; SE 3.7; NE 2.0; EE 2.7; IO 4.9; SFI 10.4; UAL 5.2; LAL 4.9; HaL 6.0; 4FL 4.5; ULL 5.7; LLL 5.9; FOL 9.6; 4TL 5.7; TD 0.7; OD 3.1; SnS 11; HeadSTr 13; CS 7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328656CDD32F35D7F9DD971FB18F977" pageId="15" pageNumber="166" type="distribution">
|
||
<paragraph id="8B8D36E7DD32F35D7F9DD971FD4DFA2F" blockId="15.[151,1437,150,1689]" pageId="15" pageNumber="166">
|
||
<emphasis id="B946EAF5DD32F35D7F9DD971FDA7FC6E" bold="true" box="[199,601,870,897]" pageId="15" pageNumber="166">Distribution and Natural History.</emphasis>
|
||
<taxonomicName id="4C324D64DD32F35D7D3AD970FD00FC6F" authorityName="Patel & Thackeray & Sheth & Khandekar & Agarwal" authorityYear="2024" box="[608,766,871,897]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="166" phylum="Chordata" rank="species" species="sinyik" status="sp. nov.">
|
||
<emphasis id="B946EAF5DD32F35D7D3AD970FD00FC6F" box="[608,766,871,897]" italics="true" pageId="15" pageNumber="166">Calotes sinyik</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAF5DD32F35D7C5FD970FCA1FC6F" bold="true" box="[773,863,871,897]" pageId="15" pageNumber="166">
|
||
<taxonomicNameLabel id="A275578EDD32F35D7C5FD970FCA1FC6F" box="[773,863,871,897]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from the
|
||
<typeStatus id="54898845DD32F35D7B29D970FB5DFC6F" box="[1139,1187,871,897]" pageId="15" pageNumber="166">type</typeStatus>
|
||
locality near Lingbuk Riamukh in Upper Subansiri District at ~
|
||
<quantity id="4CCA9B02DD32F35D7D28D99CFD34FC4B" box="[626,714,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="166" unit="m" value="1270.0">1270 m</quantity>
|
||
asl. (
|
||
<figureCitation id="13092A62DD32F35D7C51D99CFCB1FC4B" box="[779,847,907,933]" captionStart="FIGURE 1" captionStartId="1.[151,250,1617,1641]" captionTargetBox="[151,1436,309,1599]" captionTargetId="figure-87@1.[153,1435,311,1593]" captionTargetPageId="1" captionText="FIGURE 1. Relief map of northeast India and adjacent areas showing Indian state boundaries; the distribution of Calotes sinyik sp. nov. (star), C. paulus (solid black circles), C. zolaiking (solid white circles) and C. sp. Medog (solid black square). India boundary layer as per Survey of India’s outline map." figureDoi="http://doi.org/10.5281/zenodo.13934061" httpUri="https://zenodo.org/record/13934061/files/figure.png" pageId="15" pageNumber="166">Fig. 1</figureCitation>
|
||
). The region has a temperate climate characterized by hot humid summers and cool to mild winters that is influenced by the southwest monsoon (
|
||
<bibRefCitation id="EFA34B16DD32F35D7BE4D9B8FA6FFC27" author="Kottek, M. & Grieser, J. & Beck, C. & Rudolf, B. & Rubel, F." box="[1214,1425,943,969]" pageId="15" pageNumber="166" pagination="259 - 263" refId="ref9695" refString="Kottek, M., Grieser, J., Beck, C., Rudolf, B. & Rubel, F. (2006) World Map of the Koppen-Geiger climate classification updated. Meteorologische Zeitschrift, 15, 259 - 263. https: // doi. org / 10.1127 / 0941 - 2948 / 2006 / 0130" type="journal article" year="2006">
|
||
Kottek
|
||
<emphasis id="B946EAF5DD32F35D7A4ED9B8FAB8FC27" box="[1300,1350,943,969]" italics="true" pageId="15" pageNumber="166">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
). Mean annual precipitation is ~
|
||
<quantity id="4CCA9B02DD32F35D7EB5D9C4FDA2FC03" box="[495,604,979,1005]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="166" unit="mm" value="1900.0">1900 mm</quantity>
|
||
which is unequally distributed—precipitation of the driest quarter is only
|
||
<quantity id="4CCA9B02DD32F35D7FCDD9E0FF19FBFF" box="[151,231,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="166" unit="mm" value="42.0">42 mm</quantity>
|
||
and of the wettest quarter about
|
||
<quantity id="4CCA9B02DD32F35D7D3AD9E0FD32FBFF" box="[608,716,1015,1041]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="166" unit="mm" value="1000.0">1000 mm</quantity>
|
||
(
|
||
<bibRefCitation id="EFA34B16DD32F35D7D81D9E0FC27FBFF" author="Fick, S. E. & Hijmans, R. J." box="[731,985,1015,1041]" pageId="15" pageNumber="166" pagination="4302 - 4315" refId="ref9361" refString="Fick, S. E. & Hijmans, R. J. (2017) WorldClim 2: new 1 km spatial resolution climate surfaces for global land areas. International Journal of Climatology, 37 (12), 4302 - 4315. https: // doi. org / 10.1002 / joc. 5086" type="journal article" year="2017">Fick & Hijmans 2017</bibRefCitation>
|
||
). The mean temperature of the warmest quarter is 23°C and maximum temperature of the warmest month reaches about 27.6°C (
|
||
<bibRefCitation id="EFA34B16DD32F35D7B33DE0CFA9EFBDB" author="Fick, S. E. & Hijmans, R. J." box="[1129,1376,1051,1077]" pageId="15" pageNumber="166" pagination="4302 - 4315" refId="ref9361" refString="Fick, S. E. & Hijmans, R. J. (2017) WorldClim 2: new 1 km spatial resolution climate surfaces for global land areas. International Journal of Climatology, 37 (12), 4302 - 4315. https: // doi. org / 10.1002 / joc. 5086" type="journal article" year="2017">Fick & Hijmans 2017</bibRefCitation>
|
||
). The vegetation is largely recovering
|
||
<emphasis id="B946EAF5DD32F35D7D59DE28FDC5FBB7" box="[515,571,1087,1113]" italics="true" pageId="15" pageNumber="166">jhum</emphasis>
|
||
(swidden or shifting cultivation areas that are periodically cleared of woody and shrubby vegetation for cultivation). The vegetation around these upland
|
||
<emphasis id="B946EAF5DD32F35D7CA3DE74FBCFFB93" box="[1017,1073,1123,1149]" italics="true" pageId="15" pageNumber="166">jhum</emphasis>
|
||
habitats are not the typical sub-tropical evergreen or temperate broad-leaved forest vegetation that are observed in Western Arunachal (
|
||
<bibRefCitation id="EFA34B16DD32F35D7A49DE90FA6AFB4F" author="Sheth, C." box="[1299,1428,1159,1185]" pageId="15" pageNumber="166" refId="ref10923" refString="Sheth, C. (2021) Land-cover survey of Singchung Bugun Village Community Reserve. Unpublished report submitted to Wildlife Conservation Society-India and Rainforest Trust. s. n., s. l., 51 pp." type="book" year="2021">Sheth 2021</bibRefCitation>
|
||
) but lies along a transition zone of these two comprising both sub-tropical evergreen and temperate broad-leaved vegetation
|
||
<typeStatus id="54898845DD32F35D7E48DED8FEAFFB07" box="[274,337,1231,1257]" pageId="15" pageNumber="166">types</typeStatus>
|
||
. Several species of bamboo (Bambusoideae), banana (
|
||
<taxonomicName id="4C324D64DD32F35D7CF4DED8FBDFFB07" box="[942,1057,1231,1257]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
|
||
) and zingerables (
|
||
<taxonomicName id="4C324D64DD32F35D7BB6DED8FA6FFB07" authorityName="Martinov" authorityYear="1820" box="[1260,1425,1231,1257]" class="Liliopsida" family="Zingiberaceae" kingdom="Plantae" order="Zingiberales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="family">Zingiberaceae</taxonomicName>
|
||
), along with Eastern Himalayan Sago Palms (
|
||
<taxonomicName id="4C324D64DD32F35D7DC8DEE4FCF7FAE3" box="[658,777,1267,1293]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C324D64DD32F35D7C7EDEE4FB94FAE3" authority="C. F. Wei" authorityName="C. F. Wei" box="[804,1130,1267,1293]" class="Liliopsida" family="Arecaceae" genus="Arenga" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="micrantha">
|
||
<emphasis id="B946EAF5DD32F35D7C7EDEE4FC0CFAE3" box="[804,1010,1267,1293]" italics="true" pageId="15" pageNumber="166">Arenga micrantha</emphasis>
|
||
C. F. Wei
|
||
</taxonomicName>
|
||
) were observed within the understorey indicating a long history of cultivation and food production by the indigenous Tagin tribe of the area (Blench 2012). The overstorey comprised large old-growth woody trees as well as recovering vegetation typical of regenerative
|
||
<emphasis id="B946EAF5DD32F35D7E72DF48FE9EFA97" box="[296,352,1375,1401]" italics="true" pageId="15" pageNumber="166">jhum</emphasis>
|
||
agricultural areas (
|
||
<figureCitation id="13092A62DD32F35D7D6DDF48FD85FA97" box="[567,635,1375,1401]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-30@14.[203,1384,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. Habitat of Calotes sinyik sp. nov. at the type locality: (A) general view of the hilly forest near Lingbuk Riamukh, Upper Subansiri, Arunachal Pradesh, and (B) microhabitat from where individuals of the new species were collected. Photos by Harshil Patel." figureDoi="http://doi.org/10.5281/zenodo.13934087" httpUri="https://zenodo.org/record/13934087/files/figure.png" pageId="15" pageNumber="166">Fig. 9</figureCitation>
|
||
). Lingbuk Riamukh is situated on the eastern side of the river Subansiri in a sub-basin of the Subansiri known as Sichi. The Sichi basin ranges from
|
||
<quantity id="4CCA9B02DD32F35D7B57DF94FB6DFA73" box="[1037,1171,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.16" metricValueMax="4.0" metricValueMin="0.32" pageId="15" pageNumber="166" unit="m" value="2160.0" valueMax="4000.0" valueMin="320.0">320–4000m</quantity>
|
||
above sea level with a limited area lying within the
|
||
<quantity id="4CCA9B02DD32F35D7E86DFB0FD91FA2F" box="[476,623,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="15" pageNumber="166" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200–1400m</quantity>
|
||
zone.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E7DD32F35D7F9DDFDCFB18F977" blockId="15.[151,1437,150,1689]" pageId="15" pageNumber="166">
|
||
The
|
||
<typeStatus id="54898845DD32F35D7FA1DFDCFEA3FA0B" box="[251,349,1483,1509]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54898845DD32F35D7ECFDFDCFE0BFA0B" box="[405,501,1483,1509]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
|
||
were collected <
|
||
<quantity id="4CCA9B02DD32F35D7D9ADFDCFD04FA0B" box="[704,762,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="166" unit="km" value="1.0">1 km</quantity>
|
||
apart on a single transect from ~1400–2030 hrs by three of us (
|
||
<collectionCode id="ED23AE22DD32F35D7FE4DFF8FF1DF9E7" box="[190,227,1519,1545]" pageId="15" pageNumber="166">HP</collectionCode>
|
||
,
|
||
<collectionCode id="ED23AE22DD32F35D7FB4DFF8FEEDF9E7" box="[238,275,1519,1545]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="15" pageNumber="166">CS</collectionCode>
|
||
,
|
||
<collectionCode id="ED23AE22DD32F35D7E44DFF8FEBEF9E7" box="[286,320,1519,1545]" country="USA" lsid="urn:lsid:biocol.org:col:14893" name="University of Iowa" pageId="15" pageNumber="166" type="Herbarium">IA</collectionCode>
|
||
).
|
||
<location id="8EED603CDD32F35D7E08DFF8FE76F9E7" LSID="urn:lsid:plazi:treatment:039B87F1DD38F35D7FCDDD11FB18F977:8EED603CDD32F35D7E08DFF8FE76F9E7" box="[338,392,1519,1545]" county="Lingbuk Riamukh. The" municipality="Subansiri River" name="Both" pageId="15" pageNumber="166">Both</location>
|
||
lizards were spotted asleep at night, the
|
||
<typeStatus id="54898845DD32F35D7C16DFF8FC50F9E7" box="[844,942,1519,1545]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
|
||
on a leafless twig and the
|
||
<typeStatus id="54898845DD32F35D7B8EDFF8FACAF9E7" box="[1236,1332,1519,1545]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
|
||
on a thin branch of a fallen dry tree, about
|
||
<quantity id="4CCA9B02DD32F35D7D5ADC05FD74F9C3" box="[512,650,1554,1581]" metricMagnitude="0" metricUnit="m" metricValue="1.575" metricValueMax="1.8" metricValueMin="1.35" pageId="15" pageNumber="166" unit="cm" value="157.5" valueMax="180.0" valueMin="135.0">135–180 cm</quantity>
|
||
above the ground with the body roughly horizontally oriented and head directed slightly upwards.
|
||
<location id="8EED603CDD32F35D7EE5DC20FE1AF9BF" LSID="urn:lsid:plazi:treatment:039B87F1DD38F35D7FCDDD11FB18F977:8EED603CDD32F35D7EE5DC20FE1AF9BF" box="[447,484,1591,1617]" county="Lingbuk Riamukh. The" municipality="Subansiri River" name="We" pageId="15" pageNumber="166">We</location>
|
||
did not record the new species at similar elevations immediately across the
|
||
<collectingMunicipality id="6BE9AC9DDD32F35D7A68DC20FF29F99B" pageId="15" pageNumber="166">Subansiri River</collectingMunicipality>
|
||
, nor were they observed further upstream or downstream from
|
||
<collectingCounty id="62EC4E6BDD32F35D7CFADC4CFB59F99B" box="[928,1191,1627,1653]" pageId="15" pageNumber="166">Lingbuk Riamukh. The</collectingCounty>
|
||
only sympatric lizard we recorded at the type locality was
|
||
<taxonomicName id="4C324D64DD32F35D7D6BDC68FC9CF977" authority="Li" authorityName="Li" box="[561,866,1663,1689]" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" pageId="15" pageNumber="166" phylum="Chordata" rank="species" species="cayuensis">
|
||
<emphasis id="B946EAF5DD32F35D7D6BDC68FCBCF976" box="[561,834,1663,1689]" italics="true" pageId="15" pageNumber="166">Cyrtodactylus cayuensis</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
on vegetation along road cuttings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |