189 lines
21 KiB
XML
189 lines
21 KiB
XML
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<mods:title id="5EE39007360249B3A77C9969294089F8">Review of the steatiticus - species group of the cuskeel genus Neobythites (Ophidiidae) from the Indo-Pacific, with description of two new species</mods:title>
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<mods:namePart id="702A1CC1B52CA6C2A751B8423C835599">Nielsen, Jørgen G.</mods:namePart>
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<treatment id="6570D216FFF88F3AFF69E5A236995368" ID-DOI="http://doi.org/10.5281/zenodo.5978332" ID-GBIF-Taxon="141251162" ID-Zenodo-Dep="5978332" LSID="urn:lsid:plazi:treatment:6570D216FFF88F3AFF69E5A236995368" httpUri="http://treatment.plazi.org/id/6570D216FFF88F3AFF69E5A236995368" lastPageNumber="165" pageId="8" pageNumber="165">
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<paragraph id="ED666300FFF88F3AFF69E5A2373E548A" blockId="8.[151,563,655,715]" box="[151,563,655,682]" pageId="8" pageNumber="165">
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<heading id="B62ED46CFFF88F3AFF69E5A2373E548A" bold="true" box="[151,563,655,682]" fontSize="11" level="1" pageId="8" pageNumber="165" reason="1">
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<taxonomicName id="2AD91883FFF88F3AFF69E5A2373E548A" ID-CoL="46C3V" authority="Nielsen, 1995" authorityName="Nielsen" authorityYear="1995" box="[151,563,655,682]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="meteori">
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<emphasis id="DFADBF12FFF88F3AFF69E5A2373E548A" bold="true" box="[151,563,655,682]" pageId="8" pageNumber="165">
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<emphasis id="DFADBF12FFF88F3AFF69E5A2348A5489" bold="true" box="[151,391,655,681]" italics="true" pageId="8" pageNumber="165">Neobythites meteori</emphasis>
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<bibRefCitation id="89481EF1FFF88F3AFE70E5BD373E548A" box="[398,563,656,682]" pageId="8" pageNumber="165" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen, 1995</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="ED666300FFF88F3AFF69E59F34BB54EB" blockId="8.[151,563,655,715]" box="[151,438,690,715]" pageId="8" pageNumber="165">
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(
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<figureCitation id="75E27F85FFF88F3AFF60E59F342A54EB" box="[158,295,690,715]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1503,1525]" captionStartId-1="5.[151,250,876,898]" captionTargetBox-0="[163,1422,185,1492]" captionTargetBox-1="[172,1414,190,861]" captionTargetId-0="figure@3.[163,1423,185,1492]" captionTargetId-1="figure@5.[171,1415,189,862]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURE 1. Indo-Pacific Neobythites steatiticus species group. All photographs at left side and F show preserved fishes; the remaining five photographs at right side show fresh fishes, photographed on board immediately after capture. A) Neobythites gloriae n. sp. USNM 309008, HT, 158 mm SL, inner Gulf of Oman; B) N. lombokensis, ZMUC P 77744, HT, 93 mm SL, SE off Lombok, preserved (B1) and fresh (B2; side reversed); C) N. longipes, ZMUC P77740, 98 mm SL, SE off Lombok, preserved (C1) and fresh (C2); D) N. malayanus, ZMUC P77742, 138 mm SL, SE off Lombok, preserved (D1) and fresh (D2); E) N. malhaensis, ZMUC P77840, PT, 135 mm SL, Saya de Malha bank, preserved; F) N. meteori, ZMH 5621, HT, 102 mm SL, off Sokotra, preserved; G) N. steatiticus, ZMUC P771752, 162 mm SL, off Myanmar, preserved (G1) and fresh (G2); H1) N. stefanovi, ZMUC P77841, PT, 198 mm SL, Gulf of Aden, preserved; H2) N. stefanovi, SMF 26785, 111 mm SL, Red Sea, fresh. Photographers: Oddgeir Alvheim (G2), Thomas Gloerfelt-Tarp (B2, C2, D2), Marcus Krag (A, B1, C1, D1, E, G1, H1), Ralph Thiel (F), Michael Türkay (H2). The blue bars indicate 1 cm length for the respective specimen." captionText-1="FIGURE 2. Distribution of studied specimens of Neobythites, with numbers indicating multiple sampling events." httpUri-0="https://zenodo.org/record/1186738/files/figure.png" httpUri-1="https://zenodo.org/record/1186740/files/figure.png" pageId="8" pageNumber="165">Figures 1–2</figureCitation>
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,
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<tableCitation id="A05B56BBFFF88F3AFECDE59F349C54EB" box="[307,401,690,715]" captionStart="TABLE 1" captionStartId="9.[147,218,545,567]" captionText="TABLE 1. Meristic, morphometric, and quantitative ocellus and otolith characters in eight steatiticus-group species.Clear differences from Neobythites gloriae n. sp. are indicated by bold italics." pageId="8" pageNumber="165">Tables 1</tableCitation>
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,
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<tableCitation id="A05B56BBFFF88F3AFE61E59E34A054EB" box="[415,429,691,715]" captionStart="TABLE 3" captionStartId="13.[151,242,1453,1476]" captionText="TABLE 3. Distribution of selected meristic characters in the steatiticus-species group, with data for the two populations of Neobythites stefanovi." pageId="8" pageNumber="165">3</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A5C3308BFFF88F3AFF69E5D737C5550F" pageId="8" pageNumber="165" type="reference_group">
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<paragraph id="ED666300FFF88F3AFF69E5D737505530" blockId="8.[151,712,762,815]" box="[151,605,762,784]" pageId="8" pageNumber="165">
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<treatmentCitationGroup id="CDC9442EFFF88F3AFF69E5D737505530" box="[151,605,762,784]" pageId="8" pageNumber="165">
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<taxonomicName id="2AD91883FFF88F3AFF69E5D737065530" ID-CoL="46C3V" authority="Nielsen, 1995: 7" authorityName="Nielsen" authorityPageNumber="7" authorityYear="1995" box="[151,523,762,784]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="meteori">
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<emphasis id="DFADBF12FFF88F3AFF69E5D7346C5530" box="[151,353,762,784]" italics="true" pageId="8" pageNumber="165">Neobythites meteori</emphasis>
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<treatmentCitation id="6C784511FFF88F3AFE96E5D737065530" author="" box="[360,523,762,784]" page="7" pageId="8" pageNumber="165">
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<bibRefCitation id="89481EF1FFF88F3AFE96E5D734FF5530" box="[360,498,762,784]" pageId="8" pageNumber="165" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen,1995</bibRefCitation>
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: 7
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</treatmentCitation>
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</taxonomicName>
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||
, fig. 6a.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="ED666300FFF88F3AFF69E43437C5550F" blockId="8.[151,712,762,815]" box="[151,712,793,815]" pageId="8" pageNumber="165">
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<treatmentCitationGroup id="CDC9442EFFF88F3AFF69E43437C5550F" box="[151,712,793,815]" pageId="8" pageNumber="165">
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<taxonomicName id="2AD91883FFF88F3AFF69E434376A550F" ID-CoL="46C4T" authority=": Kotthaus 1979: 13" authorityName="Kotthaus" authorityPageNumber="13" authorityYear="1979" box="[151,615,793,815]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="unimaculatus">
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<emphasis id="DFADBF12FFF88F3AFF69E4343490550F" box="[151,413,793,815]" italics="true" pageId="8" pageNumber="165">Neobythites unimaculatus</emphasis>
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:
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<treatmentCitation id="6C784511FFF88F3AFE54E434376A550F" author="" box="[426,615,793,815]" page="13" pageId="8" pageNumber="165">
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<bibRefCitation id="89481EF1FFF88F3AFE54E434374F550F" box="[426,578,793,815]" pageId="8" pageNumber="165" refId="ref12239" refString="Kotthaus, A. (1979) Fische des Indischen Ozeans, XXI. " Meteor " Forschungsergebnisse, Series D (28), 6 - 54." type="journal article">Kotthaus 1979</bibRefCitation>
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: 13
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</treatmentCitation>
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</taxonomicName>
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, fig. 462.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A5C3308BFFF88F3AFF69E471312E55B8" pageId="8" pageNumber="165" type="materials_examined">
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<paragraph id="ED666300FFF88F3AFF69E471312E55B8" blockId="8.[151,1437,860,1352]" pageId="8" pageNumber="165">
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<materialsCitation id="5DB1695DFFF88F3AFF69E471311255B8" ID-GBIF-Occurrence="1826816980" collectingDate="1964-12-20" collectionCode="ZMH" collectorName="NW Indian Ocean & RV Meteor & Agassiz" country="Yemen" elevation="256" latitude="11.565" location="southwestern Socotra Island" longLatPrecision="911" longitude="52.9" pageId="8" pageNumber="165" specimenCode="ZMH 5621" specimenCount="1" typeStatus="holotype">
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<typeStatus id="3262DDA2FFF88F3AFF69E47134095555" box="[151,260,860,885]" pageId="8" pageNumber="165" type="holotype">
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<emphasis id="DFADBF12FFF88F3AFF69E47134095555" bold="true" box="[151,260,860,885]" pageId="8" pageNumber="165">Holotype</emphasis>
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</typeStatus>
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||
.
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<specimenCode id="BD7FCB7BFFF88F3AFEECE47034825555" box="[274,399,860,885]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="8" pageNumber="165">ZMH 5621</specimenCode>
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(female,
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<quantity id="2A21CEE5FFF88F3AFE05E47137555554" box="[507,600,860,884]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="165" unit="mm" value="102.0">102 mm</quantity>
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SL), off
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<location id="E80635DBFFF88F3AFD40E47136F05555" LSID="urn:lsid:plazi:treatment:6570D216FFF88F3AFF69E5A236995368:E80635DBFFF88F3AFD40E47136F05555" box="[702,1021,860,885]" country="Yemen" latitude="11.565" longLatPrecision="911" longitude="52.9" name="southwestern Socotra Island" pageId="8" pageNumber="165">southwestern Socotra Island</location>
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,
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<collectingCountry id="95CE2390FFF88F3AFBF7E47031575554" box="[1033,1114,861,884]" name="Yemen" pageId="8" pageNumber="165">Yemen</collectingCountry>
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,
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<collectorName id="402C06D6FFF88F3AFB9BE47030395554" box="[1125,1332,860,885]" pageId="8" pageNumber="165">NW Indian Ocean</collectorName>
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,
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<geoCoordinate id="88ED05C7FFF88F3AFABEE47135A355B7" direction="north" orientation="latitude" pageId="8" pageNumber="165" precision="92" value="11.565">11°33.9' N</geoCoordinate>
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,
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<geoCoordinate id="88ED05C7FFF88F3AFF47E452341155B7" box="[185,284,895,920]" direction="east" orientation="longitude" pageId="8" pageNumber="165" precision="925" value="52.9">52°54' E</geoCoordinate>
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;
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<collectorName id="402C06D6FFF88F3AFED6E4AD34A955B8" box="[296,420,896,920]" pageId="8" pageNumber="165">
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RV
|
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<emphasis id="DFADBF12FFF88F3AFEAAE4AC34A955B8" box="[340,420,897,920]" italics="true" pageId="8" pageNumber="165">Meteor</emphasis>
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</collectorName>
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, st. 102,
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<collectorName id="402C06D6FFF88F3AFDF4E452376E55B8" box="[522,611,895,920]" pageId="8" pageNumber="165">Agassiz</collectorName>
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trawl;
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||
<elevation id="66F48433FFF88F3AFD4CE452363C55B7" box="[690,817,895,920]" metricMagnitude="2" metricUnit="m" metricValue="2.56" metricValueMax="3.37" metricValueMin="1.75" pageId="8" pageNumber="165" unit="m" value="256.0" valueMax="337.0" valueMin="175.0">
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<quantity id="2A21CEE5FFF88F3AFD4CE452363C55B7" box="[690,817,895,920]" metricMagnitude="2" metricUnit="m" metricValue="2.56" metricValueMax="3.37" metricValueMin="1.75" pageId="8" pageNumber="165" unit="m" value="256.0" valueMax="337.0" valueMin="175.0">175–337 m</quantity>
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</elevation>
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depth;
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<collectingDate id="8923BC28FFF88F3AFC7BE4AD311255B8" box="[901,1055,895,920]" pageId="8" pageNumber="165" value="1964-12-20">
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<date id="996745C0FFF88F3AFC7BE4AD311255B8" box="[901,1055,895,920]" pageId="8" pageNumber="165" value="1964-12-20">20 Dec. 1964</date>
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</collectingDate>
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</materialsCitation>
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.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5C3308BFFF88F3AFF39E48931055250" pageId="8" pageNumber="165" type="diagnosis">
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<paragraph id="ED666300FFF88F3AFF39E48931055250" blockId="8.[151,1437,860,1352]" pageId="8" pageNumber="165">
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||
<emphasis id="DFADBF12FFF88F3AFF39E4893437559D" bold="true" box="[199,314,932,957]" pageId="8" pageNumber="165">Diagnosis</emphasis>
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. Flat process on hind margin of preopercle; dorsal fin-rays 91; anal-fin rays 75; pectoral fin-rays 27; precaudal vertebrae 13; total vertebrae 53; pseudobranchial filaments 3; long gill rakers on anterior arch 6; head length 22.0 % SL; pelvic-fin length 11.0 % SL, pelvic fins not reaching anus; orbit length 4.1–4.9 % SL, 22.0 % head length, and and 2.0 times in upper-jaw length; longest gill filament 2.8 % SL and 13.0 % head length; ocellus spot placed above anus, the ocellus-spot distance being 39.0 % SL, and the spot covering 9 dorsal-fin rays; dorsal and anal fins not pigmented; no vertical bars on body; otolith length 4.6 % SL.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="A5C3308BFFF88F3AFF39E351310B52B5" box="[199,1030,1148,1173]" pageId="8" pageNumber="165" type="distribution">
|
||
<paragraph id="ED666300FFF88F3AFF39E351310B52B5" blockId="8.[151,1437,860,1352]" box="[199,1030,1148,1173]" pageId="8" pageNumber="165">
|
||
<emphasis id="DFADBF12FFF88F3AFF39E351345452B5" bold="true" box="[199,345,1148,1173]" pageId="8" pageNumber="165">Distribution</emphasis>
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||
. Off Sokotra Island, NW Indian Ocean, at
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||
<quantity id="2A21CEE5FFF88F3AFCC4E35136B452B4" box="[826,953,1148,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.56" metricValueMax="3.37" metricValueMin="1.75" pageId="8" pageNumber="165" unit="m" value="256.0" valueMax="337.0" valueMin="175.0">175–337 m</quantity>
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||
depth.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="A5C3308BFFF88F3AFF39E3B236995368" pageId="8" pageNumber="165" type="discussion">
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<paragraph id="ED666300FFF88F3AFF39E3B236995368" blockId="8.[151,1437,860,1352]" pageId="8" pageNumber="165">
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<emphasis id="DFADBF12FFF88F3AFF39E3B234365298" bold="true" box="[199,315,1183,1208]" pageId="8" pageNumber="165">Remarks.</emphasis>
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In his review,
|
||
<bibRefCitation id="89481EF1FFF88F3AFE13E3B237BA5298" box="[493,695,1183,1208]" pageId="8" pageNumber="165" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen (1995: 8)</bibRefCitation>
|
||
included a second specimen (from off Lombok) as
|
||
<taxonomicName id="2AD91883FFF88F3AFAE6E38D35E152FC" authorityName="Nielsen" authorityYear="1995" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="meteori">
|
||
<emphasis id="DFADBF12FFF88F3AFAE6E38D35E152FC" italics="true" pageId="8" pageNumber="165">Neobythites meteori</emphasis>
|
||
</taxonomicName>
|
||
without designated it as
|
||
<typeStatus id="3262DDA2FFF88F3AFDFFE3E8376852FD" box="[513,613,1221,1245]" pageId="8" pageNumber="165" type="paratype">paratype</typeStatus>
|
||
. This specimen is described as
|
||
<taxonomicName id="2AD91883FFF88F3AFC3FE3E8316152FC" authority="Uiblein & Nielsen, 2018" authorityName="Uiblein & Nielsen" authorityYear="2018" box="[961,1132,1220,1244]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="lombokensis" status="sp. nov.">
|
||
<emphasis id="DFADBF12FFF88F3AFC3FE3E8316152FC" box="[961,1132,1220,1244]" italics="true" pageId="8" pageNumber="165">N. lombokensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C49E0269FFF88F3AFB8CE3E931BD52FC" box="[1138,1200,1220,1245]" pageId="8" pageNumber="165" rank="species">
|
||
<emphasis id="DFADBF12FFF88F3AFB8CE3E931A752FD" bold="true" box="[1138,1194,1220,1245]" pageId="8" pageNumber="165">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in the current paper. A photo of the freshly caught specimen, later to become the
|
||
<typeStatus id="3262DDA2FFF88F3AFCA4E3CA36B15320" box="[858,956,1255,1280]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="2AD91883FFF88F3AFC18E3C4315252DF" authorityName="Nielsen" authorityYear="1995" box="[998,1119,1256,1280]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="meteori">
|
||
<emphasis id="DFADBF12FFF88F3AFC18E3C4315252DF" box="[998,1119,1256,1280]" italics="true" pageId="8" pageNumber="165">N. meteori</emphasis>
|
||
</taxonomicName>
|
||
, shows no dark margins or bands on dorsal and anal fins (
|
||
<emphasis id="DFADBF12FFF88F3AFE12E221370D5304" box="[492,512,1292,1316]" italics="true" pageId="8" pageNumber="165">cf</emphasis>
|
||
.
|
||
<bibRefCitation id="89481EF1FFF88F3AFDF0E22137BB5304" box="[526,694,1292,1317]" pageId="8" pageNumber="165" refId="ref12239" refString="Kotthaus, A. (1979) Fische des Indischen Ozeans, XXI. " Meteor " Forschungsergebnisse, Series D (28), 6 - 54." type="journal article">Kotthaus 1979</bibRefCitation>
|
||
). Also, the otolith of the
|
||
<typeStatus id="3262DDA2FFF88F3AFC32E22131235305" box="[972,1070,1292,1317]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="2AD91883FFF88F3AFBADE22031C45304" authorityName="Nielsen" authorityYear="1995" box="[1107,1225,1293,1316]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="8" pageNumber="165" phylum="Chordata" rank="species" species="meteori">
|
||
<emphasis id="DFADBF12FFF88F3AFBADE22031C45304" box="[1107,1225,1293,1316]" italics="true" pageId="8" pageNumber="165">N. meteori</emphasis>
|
||
</taxonomicName>
|
||
turned out to be in bad condition allowing us only to measure otolith length and height.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |