304 lines
38 KiB
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304 lines
38 KiB
XML
<document id="98C2917BB7EE1A3A7A068C488F583DBD" ID-DOI="10.11646/zootaxa.4088.2.1" ID-GBIF-Dataset="6535244e-22b6-435b-9db9-16cc0a9fa1f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="258925" ID-ZooBank="43023B2C-A031-47D4-8FBE-CB2D782D825C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457512252588" checkinUser="donat" docAuthor="Oliver, Paul M. & Doughty, Paul" docDate="2016" docId="655187F3051EB051F88A9C37FEF61CFA" docLanguage="en" docName="15565-5090-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oedura bella Oliver & Doughty, 2016, sp. nov." docType="treatment" docVersion="12" lastPageNumber="168" masterDocId="9968FF8B0510B040F81D9E5BFFC6191F" masterDocTitle="Systematic revision of the marbled velvet geckos (Oedura marmorata species complex, Diplodactylidae) from the Australian arid and semi-arid zones" masterLastPageNumber="176" masterPageNumber="151" pageNumber="165" updateTime="1698177407615" updateUser="plazi">
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<mods:mods id="939F77248B00ABBCD5C52CF39DE9AE36" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="54EDD13ACAE70CA73A255562F43BD62D">
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<mods:title id="576E51C6B0A7DF4C94E8233E32A59D9A">Systematic revision of the marbled velvet geckos (Oedura marmorata species complex, Diplodactylidae) from the Australian arid and semi-arid zones</mods:title>
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<mods:name id="90A99E24843374FF6E33E331013D496E" type="personal">
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<mods:namePart id="46A49659CE2D2B0437C9B4DBE7C6A9B3">Oliver, Paul M.</mods:namePart>
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<mods:name id="EB821EC83FBEAC037BA9EACE9E8B881E" type="personal">
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<mods:roleTerm id="DB00DD59758CC0EAD73C7E8912432E47">Author</mods:roleTerm>
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<mods:namePart id="A4F74AEC1EB3E9E4971B7AA156ECA25B">Doughty, Paul</mods:namePart>
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<mods:typeOfResource id="2F91EE084D9C7E6CEA8163E650C392C5">text</mods:typeOfResource>
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<mods:title id="4B784E841E21269ED8A03975257297EA">Zootaxa</mods:title>
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<mods:part id="44A33773EC34B04E0569F3E76546C5F3">
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<mods:date id="11D431D4F275E0EB75C4718311A7D544">2016</mods:date>
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<mods:detail id="3CF7E5F07DB82759B57711E605EC2408" type="volume">
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<mods:number id="77C60B6A97FA49B46291F3326371B1EE">4088</mods:number>
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<mods:detail id="A0E038F2CB6B8329019852EE239887EB" type="issue">
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<mods:number id="FD305B75321D29BF2784C051BE4D1C17">2</mods:number>
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<mods:classification id="45E83CCF33C062BC8A66B0B2AB96F442">journal article</mods:classification>
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<mods:identifier id="7D868D7C67F73C58B15AB9804120B2D8" type="DOI">10.11646/zootaxa.4088.2.1</mods:identifier>
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<mods:identifier id="132CAAFE56843AB11DF0EB2DA2F53965" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8C90F4A496F661E47F79D9EA8D85AB61" type="Zenodo-Dep">258925</mods:identifier>
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<treatment id="655187F3051EB051F88A9C37FEF61CFA" ID-DOI="http://doi.org/10.5281/zenodo.6067116" ID-GBIF-Taxon="117196197" ID-Zenodo-Dep="6067116" LSID="urn:lsid:plazi:treatment:655187F3051EB051F88A9C37FEF61CFA" httpUri="http://treatment.plazi.org/id/655187F3051EB051F88A9C37FEF61CFA" lastPageId="17" lastPageNumber="168" pageId="14" pageNumber="165">
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<subSubSection id="A5E2656E051EB04EF88A9C37FEDD1BD4" pageId="14" pageNumber="165" type="nomenclature">
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<paragraph id="ED4736E5051EB04EF88A9C37FE5F1B99" blockId="14.[151,450,620,715]" box="[151,409,620,646]" pageId="14" pageNumber="165">
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<heading id="B60F8189051EB04EF88A9C37FE5F1B99" bold="true" box="[151,409,620,646]" fontSize="11" level="1" pageId="14" pageNumber="165" reason="1">
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<emphasis id="DF8CEAF7051EB04EF88A9C37FE5F1B99" bold="true" box="[151,409,620,646]" pageId="14" pageNumber="165">
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<taxonomicName id="2AF84D66051EB04EF88A9C37FEF21B99" box="[151,308,620,646]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
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<emphasis id="DF8CEAF7051EB04EF88A9C37FEF21B99" bold="true" box="[151,308,620,646]" italics="true" pageId="14" pageNumber="165">Oedura bella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C4BF578C051EB04EF9269C37FE5F1B99" box="[315,409,620,646]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="ED4736E5051EB04EF88A9CD5FEDD1BD4" blockId="14.[151,450,620,715]" pageId="14" pageNumber="165">
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Gulf marbled velvet gecko
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<figureCitation id="75C32A60051EB04EF88A9CE9FF221BD5" box="[151,228,690,715]" captionStart="FIGURE 5" captionStartId="8.[151,250,1847,1869]" captionTargetBox="[171,1436,205,1808]" captionTargetId="figure@8.[166,1451,193,1824]" captionTargetPageId="8" captionText="FIGURE 5. Species in the Oedura marmorata complex in life: A. O. cincta, Eastern ESU, Winton Region, Queensland; B. O. cincta, Eastern ESU, Danggalli Conservation Park, South Australia; C. O. cincta, central ESU, Simpson’s Gap, Northern Territory; D. O. cincta, central ESU, hatchling, Simpson’s Gap, Northern Territory; E. O. bella sp. nov., Doomadgee area, Queensland; F. O. bella sp. nov. holotype, 10 km south of Mt Isa, Queensland; G. O. fimbria sp. nov., Little Sandy Desert, Western Australia; H. O. fimbria sp. nov., Pannawonnica, Western Australia. Photographs: A, C – E. S. Macdonald; B. M. Hutchinson; F. P. M. Oliver; G. B. Maryan; H. R. Ellis." httpUri="https://zenodo.org/record/258930/files/figure.png" pageId="14" pageNumber="165">Figs. 5</figureCitation>
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,
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<figureCitation id="75C32A60051EB04EF8EC9CE8FEC71BD5" box="[241,257,691,714]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[194,1388,209,1363]" captionTargetId="figure@15.[183,1403,193,1378]" captionTargetPageId="15" captionText="FIGURE 7. Holotype of Oedura bella sp. nov. (QM J 94016) in dorsal and ventral views. Scale bar = 1 cm." httpUri="https://zenodo.org/record/258932/files/figure.png" pageId="14" pageNumber="165">7</figureCitation>
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,
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<figureCitation id="75C32A60051EB04EF9109CE8FEDD1BD4" box="[269,283,691,715]" captionStart="FIGURE 8" captionStartId="16.[151,250,1949,1971]" captionTargetBox="[194,1392,364,1926]" captionTargetId="figure@16.[181,1405,347,1943]" captionTargetPageId="16" captionText="FIGURE 8. Preserved specimens of Oedura bella sp. nov. showing range of variation in dorsal patterning. From top to bottom: SAMA R 35425; NTM R 7541; SAMA R 34188. Scale bar = 1 cm." httpUri="https://zenodo.org/record/258933/files/figure.png" pageId="14" pageNumber="165">8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E2656E051EB04EF88A9CA0FC101DDC" pageId="14" pageNumber="165" type="materials_examined">
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<paragraph id="ED4736E5051EB04EF88A9CA0FE4E1A44" blockId="14.[151,1437,762,1903]" pageId="14" pageNumber="165">
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<emphasis id="DF8CEAF7051EB04EF88A9CA0FECD1A0D" bold="true" box="[151,267,763,786]" pageId="14" pageNumber="165">Holotype.</emphasis>
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QM
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<accessionNumber id="F2ABAB06051EB04EF9509CA0FE581A0C" box="[333,414,763,787]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J94016" pageId="14" pageNumber="165" type="EnaNcbi">J94016</accessionNumber>
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(field number—OMAR #001), adult male with original tail, 10 km south of Mt Isa on Boulia Road, Queensland, Australia (20.8617°S, 139.4617°E), collected by P.M. Oliver, M. Vucko and M. Vickers, on 20 February 2007.
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</paragraph>
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<paragraph id="ED4736E5051EB04EF8DA9D3CFED71D64" blockId="14.[151,1437,762,1903]" pageId="14" pageNumber="165">
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<emphasis id="DF8CEAF7051EB04EF8DA9D3CFE801A61" bold="true" box="[199,326,871,894]" pageId="14" pageNumber="165">Paratypes.</emphasis>
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Northern Territory: AMS R53437–8, 37 km north of McArthur River base camp on Borroloola Road (16.10°S, 136.12°E); AMS R53467, Caranbirini Water Hole, 21 km north of McArthur River base camp (16.22°S, 136.15°E); AMS R53643, Glyde River, 10 km east of McArthur River base camp (16.43°S, 136.17°E); AMS R53782, 37 km north of McArthur River Camp (16.10°S, 136.12°E); SAMA R34188, McArthur River Station (16.67°S, 135.85°E). Queensland: NTM R21288, Musselbrook Reservoir (18.61°S, 138.08°E); SAMA R34208–9, SAMA R35425, Lawn Hill NP (18.58°S, 138.50°E); QM
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<accessionNumber id="F2ABAB06051EB04EFBB29A40FBC41D2C" box="[943,1026,1051,1075]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J52748" pageId="14" pageNumber="165" type="EnaNcbi">J52748</accessionNumber>
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, Lawn Hill Station, Century Project Site (18.75°S, 138.58°E); QM
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<accessionNumber id="F2ABAB06051EB04EFA1B9A64FD9C1D48" box="[518,602,1087,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74927" pageId="14" pageNumber="165" type="EnaNcbi">J74927</accessionNumber>
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, Hells Gate (17.47°S; 138.37°E); QM
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<accessionNumber id="F2ABAB06051EB04EFC309A64FB471D48" box="[1069,1153,1087,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J75207" pageId="14" pageNumber="165" type="EnaNcbi">J75207</accessionNumber>
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–8, Lawn Hill (18.71°S, 138.48°E).
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</paragraph>
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<paragraph id="ED4736E5051EB04EF8DA9ADDFC101DDC" blockId="14.[151,1437,762,1903]" pageId="14" pageNumber="165">
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<emphasis id="DF8CEAF7051EB04EF8DA9ADDFE631D80" bold="true" box="[199,421,1158,1183]" pageId="14" pageNumber="165">Referred material.</emphasis>
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AMS R138727–8, Groote Eylandt (13.83°S, 136.42°E); NTM R7494–5, Umbakumba Rd (13.88°S, 136.50°E); NTM R7541, Ayakamindadina (13.97°S, 136.60°E).
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E2656E051EB04EF8DA9A95FCE21C84" pageId="14" pageNumber="165" type="diagnosis">
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<paragraph id="ED4736E5051EB04EF8DA9A95FCE21C84" blockId="14.[151,1437,762,1903]" pageId="14" pageNumber="165">
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<emphasis id="DF8CEAF7051EB04EF8DA9A95FE871DF8" bold="true" box="[199,321,1230,1255]" pageId="14" pageNumber="165">Diagnosis.</emphasis>
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A medium-sized (SVL: mean 78 mm, max 92 mm) species in the
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<taxonomicName id="2AF84D66051EB04EFC5C9A94FB251DF9" box="[1089,1251,1231,1255]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="165" phylum="Chordata" rank="species" species="marmorata">
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<emphasis id="DF8CEAF7051EB04EFC5C9A94FB251DF9" box="[1089,1251,1231,1255]" italics="true" pageId="14" pageNumber="165">O. marmorata</emphasis>
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</taxonomicName>
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complex with a wide (HW/SVL 0.19–0.23) and moderately deep head (HD/SVL 0.10–0.12), short body (Trk/SVL 0.41–0.49), short original tail (TL/SVL 0.49–0.65) that is narrower than head and slightly depressed, rostral usually less than half divided, terminal lamellae moderately wide (ToeW/SVL 0.021–0.030), proximal subdigital lamellae of all fingers not wider than apical pair, 12–17 precloacal pores in males and base colouration dark purplish brown with 5 distinct to faint light dorsal bands from nape to hindlimbs.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E2656E051EB050F8DA9BFDFAAB1829" lastPageId="16" lastPageNumber="167" pageId="14" pageNumber="165" type="description">
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<paragraph id="ED4736E5051EB04EF8DA9BFDFEED1E70" blockId="14.[151,1437,762,1903]" pageId="14" pageNumber="165">
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<emphasis id="DF8CEAF7051EB04EF8DA9BFDFE9F1CA0" bold="true" box="[199,345,1446,1471]" pageId="14" pageNumber="165">Description.</emphasis>
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A medium-sized (to 92.0 mm SVL) and moderately robust
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<taxonomicName id="2AF84D66051EB04EFC199BFDFB9C1CA1" box="[1028,1114,1446,1470]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="165" phylum="Chordata" rank="genus">
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<emphasis id="DF8CEAF7051EB04EFC199BFDFB9C1CA1" box="[1028,1114,1446,1470]" italics="true" pageId="14" pageNumber="165">Oedura</emphasis>
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</taxonomicName>
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(Trk/SVL 0.41–0.49). Head moderately narrow (HW/SVL 0.19–0.23) and not strongly depressed (HD/SVL 0.10–0.12). Rostral 25–50% divided, bordered dorsally by two internasals, internasals bordered dorsally by two oblong moderately enlarged supranasals and 1–3 (mode 1) smaller intervening scales. Enlarged supralabials 9–10 to midpoint of eye, 11–13 in total; infralabials 9–13. Limbs moderately long (ArmL/SVL 0.12–0.15; LegL/SVL 0.14–0.16). Subdigital lamellae moderately expanded and prominent, 7–8 rows (mode 8) under finger three and 7–9 toe three, distal lamellae on all digits either divided or deeply notched, apical distinctly larger and generally wider than more proximal pairsespecially on fingers, proximal lamellae undivided. Enlarged postcloacal spurs 2 or 3, in a single cluster per side. Original tail relatively short (TL/SVL 0.49–0.65) and wide (TW/TL 0.18–0.28), gradually tapering to a point, slightly depressed in cross-section and slightly concave ventrally; moderately wide but shows considerable variation depending on body condition (TW/SVL 0.10–0.15), scalation relatively homogeneous with dorsal and ventral scales of similar size. Fully regrown tails shorter (TL/SVL 0.44–0.53) and wider (TW/TD 0.21–0.31) than original tails.
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</paragraph>
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<caption id="B987666D051FB04FF88A9B23FB1D1C91" box="[151,1243,1400,1422]" httpUri="https://zenodo.org/record/258932/files/figure.png" pageId="15" pageNumber="166" targetBox="[194,1388,209,1363]" targetPageId="15">
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<paragraph id="ED4736E5051FB04FF88A9B23FB1D1C91" blockId="15.[151,1243,1400,1422]" box="[151,1243,1400,1422]" pageId="15" pageNumber="166">
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<emphasis id="DF8CEAF7051FB04FF88A9B23FED41C91" bold="true" box="[151,274,1400,1422]" pageId="15" pageNumber="166">FIGURE 7.</emphasis>
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Holotype of
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<taxonomicName id="2AF84D66051FB04FF9849B23FDD91C91" box="[409,543,1400,1422]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="166" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
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<emphasis id="DF8CEAF7051FB04FF9849B23FDD91C91" box="[409,543,1400,1422]" italics="true" pageId="15" pageNumber="166">Oedura bella</emphasis>
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</taxonomicName>
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<emphasis id="DF8CEAF7051FB04FFA389B22FDB01C91" bold="true" box="[549,630,1401,1422]" pageId="15" pageNumber="166">
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<taxonomicNameLabel id="C4BF578C051FB04FFA389B22FDB01C91" box="[549,630,1401,1422]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(QM
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<accessionNumber id="F2ABAB06051FB04FFAA99B23FD391C91" box="[692,767,1400,1422]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J94016" pageId="15" pageNumber="166" type="EnaNcbi">J94016</accessionNumber>
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) in dorsal and ventral views. Scale bar = 1 cm.
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</paragraph>
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</caption>
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<paragraph id="ED4736E5051FB04FF8DA9B8EFF2A1E26" blockId="15.[151,1436,1492,2016]" pageId="15" pageNumber="166">In preservative, dorsal colouration of adults variable; base colouration dark purplish brown, with 5 distinct to very indistinct broad light bands from nape to hindlimbs, with a further 4–7 similar bands on the tail. Light bands with little to extensive dark brown flecking or blotches; on larger specimens dark brown pigmentation is extensive such that the broad light bands are reduced to two narrow indistinct and often broken bands. The anterior (nuchal) light band extends across the nape and joins or approaches a light lateral band extending from the labial scales and above the tympanum. The canthal stripe ranges from distinct to indistinct. Dark brown regions between light bands and elsewhere on the dorsal and lateral surfaces of the head, torso, limbs and tail, with extensive further light flecking and blotches. Venter relatively plain light buff with a very faint wash of light brown, particularly towards the lateral extremities and on the head and limbs, more extensive brown mottling also present along the ventrolateral edges of the tail. Regrown tails dark brown with varying amounts of light flecking, but no clear pattern.</paragraph>
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<paragraph id="ED4736E5051FB04FF8DA9919FD7B1EA0" blockId="15.[151,1436,1492,2016]" pageId="15" pageNumber="166">Hatchlings and small juveniles lack light flecking and generally have a very simple pattern of five sharplydefined light bands on the neck and body, and 5–7 light bands on the tail, separated by broad plain dark brown regions. As specimens grow, the sharp definition of light and dark regions decreases, and the amount of light flecking and blotching across the body increases.</paragraph>
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<paragraph id="ED4736E5051FB050F8DA9993FC5B19CD" blockId="15.[151,1436,1492,2016]" lastBlockId="16.[151,1437,151,310]" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">In life, the base colouration of adults is dark purplish brown during the day, washed out grey at night; light bands and flecks ranges tend to be bright yellow. The canthal stripe tends to be very strongly defined and a white labial stripe is also sometimes clearly apparent. Iris very dark brown.</paragraph>
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<paragraph id="ED4736E50500B050F8DA9E81FAAB1829" blockId="16.[151,1437,151,310]" pageId="16" pageNumber="167">
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<emphasis id="DF8CEAF70500B050F8DA9E81FE2719ED" bold="true" box="[199,481,218,243]" pageId="16" pageNumber="167">Particulars of holotype.</emphasis>
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QM
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<accessionNumber id="F2ABAB060500B050FA009E80FDA819EC" box="[541,622,219,243]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J94016" pageId="16" pageNumber="167" type="EnaNcbi">J94016</accessionNumber>
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(field tag number—OMAR#001), male with original tail (in mm): SVL 77.0, HW 14.6, HD 7.4, HL 19.6, EN 6.3, IN 2.8, IO 6.7, EYE 5.0, TrK 31.0, ArmL 10.1, LegL 10.9, TL 38.0, TW 8.3, TD 5.6, 3FW 1.8, 3TW 2.3. Scales: RC 25%, SuL 12(10), InF 11, CS, 3FL 7, 3TL 8, precloacal pores 16.
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</paragraph>
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</subSubSection>
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|
||
<paragraph id="ED4736E50500B050F88A99C6FCF71ECD" blockId="16.[151,1437,1949,2002]" pageId="16" pageNumber="167">
|
||
<emphasis id="DF8CEAF70500B050F88A99C6FED41EAC" bold="true" box="[151,274,1949,1971]" pageId="16" pageNumber="167">FIGURE 8.</emphasis>
|
||
Preserved specimens of
|
||
<taxonomicName id="2AF84D660500B050FA1699C6FD571EAC" box="[523,657,1949,1971]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
|
||
<emphasis id="DF8CEAF70500B050FA1699C6FD571EAC" box="[523,657,1949,1971]" italics="true" pageId="16" pageNumber="167">Oedura bella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DF8CEAF70500B050FA8A99C5FD211EAC" bold="true" box="[663,743,1950,1971]" pageId="16" pageNumber="167">
|
||
<taxonomicNameLabel id="C4BF578C0500B050FA8A99C5FD211EAC" box="[663,743,1950,1971]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showing range of variation in dorsal patterning. From top to bottom: SAMA R35425; NTM R7541; SAMA R34188. Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A5E2656E0501B051F8DA9ECCFDE618B2" pageId="17" pageNumber="168" type="distribution">
|
||
<paragraph id="ED4736E50501B051F8DA9ECCFBDA185F" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<emphasis id="DF8CEAF70501B051F8DA9ECCFE3819AF" bold="true" box="[199,510,151,176]" pageId="17" pageNumber="168">Distribution and habitat.</emphasis>
|
||
Restricted to the ranges around the south and western edges of the Gulf of Carpentaria; from the Selwyn Range around Mt Isa in the southeast, north to Riversleigh and Musselbrook regions in the Queensland-Northern Territory border, as far west as the McArthur River and Borroloola regions of Northern Territory, with a further apparently isolated insular population on Groote Eylandt to the north (see comments), and another isolated record from the northern edge of the Barkly Tablelands (
|
||
<figureCitation id="75C32A600501B051FBD79F7CFBC8185F" box="[970,1038,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1873,1895]" captionTargetBox="[159,1429,991,1845]" captionTargetId="figure@2.[151,1436,985,1852]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of three Oedura species resurrected or described in this paper: O. bella sp. nov. (blue, Gulf) from the Gulf Country, O. cincta (red, comprising Central and Eastern ESUs), and O. fimbria sp. nov. (yellow-brown, Western) from mid-western Australia. The distribution of the residual O. marmorata complex (green, comprising O. gemmata, true O. marmorata, and related lineages) in the Top End of northern Australia is also indicated. Darker regions indicate greater topographic complexity." httpUri="https://zenodo.org/record/258927/files/figure.png" pageId="17" pageNumber="168">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="ED4736E50501B051F8DA9F17FDE618B2" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">Field observations suggest this species is primarily saxicoline, using both horizontal screes and vertical faces (including road cuttings), however, it has also been recorded under bark around the base of trees (but in rocky country) (G. Bourke, pers. comm.).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E2656E0501B051F8DA9FE3FCCC1BD2" pageId="17" pageNumber="168" type="etymology">
|
||
<paragraph id="ED4736E50501B051F8DA9FE3FC371B07" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<emphasis id="DF8CEAF70501B051F8DA9FE3FE8818D0" bold="true" box="[199,334,440,463]" pageId="17" pageNumber="168">Etymology.</emphasis>
|
||
From the Latin masculine adjective
|
||
<emphasis id="DF8CEAF70501B051FAE89FECFC8718D0" box="[757,833,439,463]" italics="true" pageId="17" pageNumber="168">bellum</emphasis>
|
||
(used in its feminine form), meaning amongst other things pretty, handsome, charming, fine, lovely, neat, pleasant, agreeable, active, gallant or good. In reference to the very attractive contrasting yellow and dark colour pattern of this species.
|
||
</paragraph>
|
||
<paragraph id="ED4736E50501B051F8DA9C7FFCCC1BD2" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<emphasis id="DF8CEAF70501B051F8DA9C7FFEAA1B22" bold="true" box="[199,364,548,573]" pageId="17" pageNumber="168">Comparisons.</emphasis>
|
||
<taxonomicName id="2AF84D660501B051F9679C7FFDD21B23" box="[378,532,548,572]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051F9679C7FFDD21B23" box="[378,532,548,572]" italics="true" pageId="17" pageNumber="168">Oedura bella</emphasis>
|
||
</taxonomicName>
|
||
is similar to parapatric populations of
|
||
<taxonomicName id="2AF84D660501B051FBEF9C7FFB5E1B24" box="[1010,1176,548,572]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="DF8CEAF70501B051FBEF9C7FFB5E1B24" box="[1010,1176,548,572]" italics="true" pageId="17" pageNumber="168">O. marmorata</emphasis>
|
||
</taxonomicName>
|
||
in northern Australia (although it differs in both nuclear and mitochondrial loci). It differs in external morphology by possessing an original or regrown tail that is much narrower (TW/SVL 0.10–0.15
|
||
<emphasis id="DF8CEAF70501B051FB819C37FC221B9C" box="[924,996,620,643]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
0.19–0.24) and generally less than the width of head, and
|
||
<taxonomicName id="2AF84D660501B051F96D9CCBFE0D1BB8" box="[368,459,655,680]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051F96D9CCBFE0D1BB8" box="[368,459,655,680]" italics="true" pageId="17" pageNumber="168">O. bella</emphasis>
|
||
</taxonomicName>
|
||
also reaches a smaller maximum size, although adult sizes overlap (adult SVL usually 77–92 mm
|
||
<emphasis id="DF8CEAF70501B051F90B9CEFFE9B1BD4" box="[278,349,692,715]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
77–97 mm) and sample sizes are low.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E2656E0501B051F8DA9C8CFC361C68" pageId="17" pageNumber="168" type="description">
|
||
<paragraph id="ED4736E50501B051F8DA9C8CFB7B1A9F" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<taxonomicName id="2AF84D660501B051F8DA9C8CFE9A1BF0" box="[199,348,727,751]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051F8DA9C8CFE9A1BF0" box="[199,348,727,751]" italics="true" pageId="17" pageNumber="168">Oedura bella</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2AF84D660501B051F9EC9C83FD9F1BF0" box="[497,601,728,752]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="cincta">
|
||
<emphasis id="DF8CEAF70501B051F9EC9C83FD9F1BF0" box="[497,601,728,752]" italics="true" pageId="17" pageNumber="168">O. cincta</emphasis>
|
||
</taxonomicName>
|
||
and Western (see below) by the combination of its smaller maximum size (SVL 64–92 mm
|
||
<emphasis id="DF8CEAF70501B051F9419CA7FE651A0C" box="[348,419,764,787]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
77–106 mm), shorter original tail (TL/SVL 0.49–0.65
|
||
<emphasis id="DF8CEAF70501B051FC129CA7FB901A0C" box="[1039,1110,764,787]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
0.58–0.80), and its narrower terminal lamellae (0.21–0.30
|
||
<emphasis id="DF8CEAF70501B051F9E99D7BFDFD1A28" box="[500,571,800,823]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
0.23–0.36) and narrower lamellae series on the fingers (not wider than terminal lamellae
|
||
<emphasis id="DF8CEAF70501B051F9759D1FFE761A44" box="[360,432,836,859]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
wider on digits 3 and 4). It further differs from the geographically proximate
|
||
<taxonomicName id="2AF84D660501B051FD5B9D1FFF1A1A60" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="cincta">
|
||
<emphasis id="DF8CEAF70501B051FD5B9D1FFF1A1A60" italics="true" pageId="17" pageNumber="168">Oedura cincta</emphasis>
|
||
</taxonomicName>
|
||
by generally having a rostral partially divided by a crease (
|
||
<emphasis id="DF8CEAF70501B051FB689D33FC7B1A60" box="[885,957,872,895]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
usually fully divided).
|
||
</paragraph>
|
||
<paragraph id="ED4736E50501B051F8DA9DD7FC3D1D62" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<taxonomicName id="2AF84D660501B051F8DA9DD7FEA61ABB" box="[199,352,908,932]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051F8DA9DD7FEA61ABB" box="[199,352,908,932]" italics="true" pageId="17" pageNumber="168">Oedura bella</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the three species of
|
||
<taxonomicName id="2AF84D660501B051FB859DD7FC2B1ABB" box="[920,1005,908,932]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
|
||
<emphasis id="DF8CEAF70501B051FB859DD7FC2B1ABB" box="[920,1005,908,932]" italics="true" pageId="17" pageNumber="168">Oedura</emphasis>
|
||
</taxonomicName>
|
||
in the Kimberley region in Western Australia by having subdigital lamellae that are slightly expanded around the midpoint of the digit (
|
||
<emphasis id="DF8CEAF70501B051FCEF9DEBFAFC1AD8" box="[1266,1338,944,967]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
strongly tapering in
|
||
<taxonomicName id="2AF84D660501B051F9049D8FFDDA1AF3" authority="King, 1984" authorityName="King" authorityYear="1984" box="[281,540,980,1005]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="DF8CEAF70501B051F9049D8FFE571AF3" box="[281,401,980,1004]" italics="true" pageId="17" pageNumber="168">O. gracilis</emphasis>
|
||
King, 1984
|
||
</taxonomicName>
|
||
, or obviously flared and often as wider or wider than the apical lamellae in
|
||
<taxonomicName id="2AF84D660501B051FD9C9D8FFE4C1D0F" authority="King, 1984" authorityName="King" authorityYear="1984" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="filicipoda">
|
||
<emphasis id="DF8CEAF70501B051FD9C9D8FFEC51D10" italics="true" pageId="17" pageNumber="168">O. filicipoda</emphasis>
|
||
King, 1984
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AF84D660501B051F9DE9DA3FB8C1D0F" authority="Oliver, Laver, Melville & Doughty, 2014" authorityName="Oliver, Laver, Melville & Doughty" authorityYear="2014" box="[451,1098,1015,1040]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="murrumanu">
|
||
<emphasis id="DF8CEAF70501B051F9DE9DA3FDAD1D10" box="[451,619,1016,1040]" italics="true" pageId="17" pageNumber="168">O. murrumanu</emphasis>
|
||
Oliver, Laver, Melville & Doughty, 2014
|
||
</taxonomicName>
|
||
), and its moderately long and slightly swollen tail (
|
||
<emphasis id="DF8CEAF70501B051F9979A47FE141D2C" box="[394,466,1052,1075]" italics="true" pageId="17" pageNumber="168">versus</emphasis>
|
||
very long [approaching length of body] and tapering in
|
||
<taxonomicName id="2AF84D660501B051FC409A47FB751D2B" box="[1117,1203,1052,1076]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="DF8CEAF70501B051FC409A47FB751D2B" box="[1117,1203,1052,1076]" italics="true" pageId="17" pageNumber="168">gracilis</emphasis>
|
||
</taxonomicName>
|
||
, or greatly flattened and wider than body in
|
||
<taxonomicName id="2AF84D660501B051F9839A1BFDED1D48" box="[414,555,1087,1112]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="filicipoda">
|
||
<emphasis id="DF8CEAF70501B051F9839A1BFDED1D48" box="[414,555,1087,1112]" italics="true" pageId="17" pageNumber="168">O. filicipoda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AF84D660501B051FA7C9A1BFCC11D48" box="[609,775,1088,1112]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="murrumanu">
|
||
<emphasis id="DF8CEAF70501B051FA7C9A1BFCC11D48" box="[609,775,1088,1112]" italics="true" pageId="17" pageNumber="168">O. murrumanu</emphasis>
|
||
</taxonomicName>
|
||
). With a maximum SVL of 92 mm,
|
||
<taxonomicName id="2AF84D660501B051FC819A1BFB301D48" box="[1180,1270,1087,1112]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051FC819A1BFB301D48" box="[1180,1270,1087,1112]" italics="true" pageId="17" pageNumber="168">O. bella</emphasis>
|
||
</taxonomicName>
|
||
is also smaller than
|
||
<taxonomicName id="2AF84D660501B051F8D39A3FFE9D1D63" box="[206,347,1124,1148]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="filicipoda">
|
||
<emphasis id="DF8CEAF70501B051F8D39A3FFE9D1D63" box="[206,347,1124,1148]" italics="true" pageId="17" pageNumber="168">O. filicipoda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AF84D660501B051F98F9A3FFDFE1D64" box="[402,568,1124,1148]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="murrumanu">
|
||
<emphasis id="DF8CEAF70501B051F98F9A3FFDFE1D64" box="[402,568,1124,1148]" italics="true" pageId="17" pageNumber="168">O. murrumanu</emphasis>
|
||
</taxonomicName>
|
||
(which both regularly exceed 100 mm).
|
||
</paragraph>
|
||
<paragraph id="ED4736E50501B051F8DA9ADCFC361C68" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<taxonomicName id="2AF84D660501B051F8DA9ADCFEA61D80" box="[199,352,1159,1183]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="bella">
|
||
<emphasis id="DF8CEAF70501B051F8DA9ADCFEA61D80" box="[199,352,1159,1183]" italics="true" pageId="17" pageNumber="168">Oedura bella</emphasis>
|
||
</taxonomicName>
|
||
differs from all other
|
||
<taxonomicName id="2AF84D660501B051FA739ADCFD021D80" box="[622,708,1159,1183]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
|
||
<emphasis id="DF8CEAF70501B051FA739ADCFD021D80" box="[622,708,1159,1183]" italics="true" pageId="17" pageNumber="168">Oedura</emphasis>
|
||
</taxonomicName>
|
||
in eastern Australia by possessing more than one postcloacal tubercle and in having a base colouration of five relatively thin dorsal bands. The latter character distinguishes it from two other small saxicoline
|
||
<taxonomicName id="2AF84D660501B051FA1E9A94FD9E1DF8" box="[515,600,1231,1255]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
|
||
<emphasis id="DF8CEAF70501B051FA1E9A94FD9E1DF8" box="[515,600,1231,1255]" italics="true" pageId="17" pageNumber="168">Oedura</emphasis>
|
||
</taxonomicName>
|
||
in eastern Australia:
|
||
<taxonomicName id="2AF84D660501B051FB549A8BFC071DF8" box="[841,961,1232,1256]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="coggeri">
|
||
<emphasis id="DF8CEAF70501B051FB549A8BFC071DF8" box="[841,961,1232,1256]" italics="true" pageId="17" pageNumber="168">O. coggeri</emphasis>
|
||
</taxonomicName>
|
||
has large ocelli on limbs and torso and
|
||
<taxonomicName id="2AF84D660501B051FD9C9A8BFED31C13" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="jowalbinna">
|
||
<emphasis id="DF8CEAF70501B051FD9C9A8BFED31C13" italics="true" pageId="17" pageNumber="168">O. jowalbinna</emphasis>
|
||
</taxonomicName>
|
||
has a pale pinkish gray dorsum with distinct dark-edged bands across the neck and base of tail and a plain yellow original tail (Hoskin & Higgie 2008). All other
|
||
<taxonomicName id="2AF84D660501B051FB589B4CFC5D1C30" box="[837,923,1303,1327]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
|
||
<emphasis id="DF8CEAF70501B051FB589B4CFC5D1C30" box="[837,923,1303,1327]" italics="true" pageId="17" pageNumber="168">Oedura</emphasis>
|
||
</taxonomicName>
|
||
in eastern Australia tend to be larger (SVL> 90 mm) and also have dorsal patterns that do not feature thin light bands; specifically,
|
||
<taxonomicName id="2AF84D660501B051FC8F9B67FAED1C4B" box="[1170,1323,1340,1364]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="castelnaui">
|
||
<emphasis id="DF8CEAF70501B051FC8F9B67FAED1C4B" box="[1170,1323,1340,1364]" italics="true" pageId="17" pageNumber="168">O. castelnaui</emphasis>
|
||
</taxonomicName>
|
||
has wide bands,
|
||
<taxonomicName id="2AF84D660501B051F8FB9B3BFE9D1C68" box="[230,347,1375,1400]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="monilis">
|
||
<emphasis id="DF8CEAF70501B051F8FB9B3BFE9D1C68" box="[230,347,1375,1400]" italics="true" pageId="17" pageNumber="168">O. monilis</emphasis>
|
||
</taxonomicName>
|
||
has blotches or ocelli, and
|
||
<taxonomicName id="2AF84D660501B051FA919B3BFD371C68" box="[652,753,1376,1400]" class="Reptilia" family="Diplodactylidae" genus="Oedura" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tryoni">
|
||
<emphasis id="DF8CEAF70501B051FA919B3BFD371C68" box="[652,753,1376,1400]" italics="true" pageId="17" pageNumber="168">O. tryoni</emphasis>
|
||
</taxonomicName>
|
||
has dense small spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5E2656E0501B051F8DA9BDEFEF61CFA" pageId="17" pageNumber="168" type="discussion">
|
||
<paragraph id="ED4736E50501B051F8DA9BDEFEF61CFA" blockId="17.[151,1437,151,1509]" pageId="17" pageNumber="168">
|
||
<emphasis id="DF8CEAF70501B051F8DA9BDEFEFD1C83" bold="true" box="[199,315,1413,1436]" pageId="17" pageNumber="168">Remarks.</emphasis>
|
||
No genetic samples were available from the Groote Eylandt samples that we refer to this species. Given their geographic disjunction from all other material, the distinctiveness of this population warrants further investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |