treatments-xml/data/65/04/B1/6504B17AFFBD654BFCF7C662F0732A64.xml
2024-06-21 12:38:42 +02:00

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<document id="80272746ECC03A199BE0D1714D25851C" ID-CLB-Dataset="8150" ID-DOI="10.5252/geodiversitas2021v43a3" ID-GBIF-Dataset="49326ee5-a67b-492e-b229-6f25dcd61f0e" ID-ISSN="1638-9395" ID-Zenodo-Dep="4548375" ID-ZooBank="urn:lsid:zoobank.org:pub:AA28648F-709F-44D6-A3CF-0C1D78B971CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613648752258" checkinUser="felipe" docAuthor="Pérez-García, Adán, Suarez-Hernando, Oier, Hernández, Jose M., García, Salvador &amp; Murelaga, Xabier" docDate="2021" docId="6504B17AFFBD654BFCF7C662F0732A64" docLanguage="en" docName="Geodiversitas.43.3.75-94.pdf" docOrigin="Geodiversitas 43 (3)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Ptychogaster (Temnoclemmys) bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia 1999" docType="treatment" docVersion="6" lastPageNumber="85" masterDocId="993DC902FFB86544FF8EC010F3742B56" masterDocTitle="The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin" masterLastPageNumber="94" masterPageNumber="75" pageNumber="78" updateTime="1698930251271" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="A84D9A8B47ECFE990ADB5E67CA580057">The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin</mods:title>
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<mods:namePart id="5E2CDAB1848C3B4EAF9CD2E57AED0C95">Pérez-García, Adán</mods:namePart>
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<mods:namePart id="C8FA064C9A85A857D0D134DA09AE3619">Suarez-Hernando, Oier</mods:namePart>
<mods:affiliation id="46987FDFF13788C2F276A53FD6594042">Universidad del País Vasco UPV / EHU, Facultad de Ciencia y Tecnología, Departamento de Estratigrafía y Paleontología, Bilbao Apartado 644, E- 48080 Bizkaia (Spain)</mods:affiliation>
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<mods:namePart id="A76AB2C7B00BC2D667650D999879BD7C">Hernández, Jose M.</mods:namePart>
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<mods:namePart id="4F03FB3FCC87348ED17A0FDC268E577C">García, Salvador</mods:namePart>
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<mods:namePart id="FBF4080613CF2216A3A813247E11FCDF">Murelaga, Xabier</mods:namePart>
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<treatment id="6504B17AFFBD654BFCF7C662F0732A64" ID-DOI="http://doi.org/10.5281/zenodo.4548640" ID-GBIF-Taxon="177423225" ID-Zenodo-Dep="4548640" LSID="urn:lsid:plazi:treatment:6504B17AFFBD654BFCF7C662F0732A64" httpUri="http://treatment.plazi.org/id/6504B17AFFBD654BFCF7C662F0732A64" lastPageId="15" lastPageNumber="85" pageId="5" pageNumber="78">
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<paragraph id="ED12006CFFBD6541FCF7C662F79E2D9A" blockId="5.[884,1384,1650,1741]" pageId="5" pageNumber="78">
<taxonomicName id="2AAD7BEFFFBD6541FCF7C662F79E2D9A" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="5" pageNumber="78" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFBD6541FCF7C662F08B2DDB" bold="true" box="[889,1023,1650,1677]" italics="true" pageId="5" pageNumber="78">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFBD6541FB81C662F7C12DDB" bold="true" box="[1039,1205,1650,1677]" italics="true" pageId="5" pageNumber="78">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFBD6541FB6BC662F6172DDB" bold="true" box="[1253,1379,1650,1677]" italics="true" pageId="5" pageNumber="78">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFBD6541FC46C682F79E2D9A" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="5" pageNumber="78" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
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<subSubSection id="A5B753E7FFBD6541FB7FC6A2F61C2D9B" box="[1265,1384,1714,1741]" pageId="5" pageNumber="78" type="description">
<paragraph id="ED12006CFFBD6541FB7FC6A2F61C2D9B" blockId="5.[884,1384,1650,1741]" box="[1265,1384,1714,1741]" pageId="5" pageNumber="78">
(
<figureCitation id="75961CE9FFBD6541FB74C6A2F62B2D9A" box="[1274,1375,1714,1741]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStartId-0="5.[132,143,1894,1911]" captionStartId-1="7.[132,143,1872,1889]" captionStartId-2="8.[132,143,1917,1934]" captionStartId-3="9.[132,143,1491,1508]" captionStartId-4="10.[132,143,1747,1764]" captionStartId-5="11.[132,143,1779,1796]" captionStartId-6="12.[132,143,1137,1154]" captionStartId-7="13.[132,143,1826,1843]" captionTargetBox-1="[134,1455,246,1846]" captionTargetBox-2="[136,1452,218,1848]" captionTargetBox-3="[136,1453,219,1454]" captionTargetBox-4="[135,1455,218,1710]" captionTargetBox-5="[136,1451,216,1737]" captionTargetBox-6="[134,1458,216,1095]" captionTargetBox-7="[135,1452,218,1736]" captionTargetId-0="figure-19@6.[132,1455,224,2010]" captionTargetId-1="figure-15@7.[132,1455,243,1849]" captionTargetId-2="figure-19@8.[132,1455,228,1844]" captionTargetId-3="figure-224@9.[132,1455,241,1471]" captionTargetId-4="figure-106@10.[132,1455,214,1715]" captionTargetId-5="figure-88@11.[132,1455,209,1739]" captionTargetId-6="figure-515@12.[129,1452,223,1098]" captionTargetId-7="figure-15@13.[132,1455,214,1742]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionText-0="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." captionText-1="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." captionText-2="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-3="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-4="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." captionText-5="FIG. 8. — Articulated elements of the posterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain). All of them are in ventral view: A, MP1a-1345; B, MP1a-1068b; C, MP1a-1423; D, MP1a-1421. Abbreviations for the plates (in lower case): hp, hypoplastron; xi, xiphiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; FE, femoral. Scale bar: 3 cm." captionText-6="FIG. 9. — Plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 2 (A-F) and Monte Pila 3 (G) (Lardero, La Rioja, Spain): A, B, articulated epiplastra, MP2- 306, in ventral (A) and dorsal (B) views; C, xiphiplastron, MP2-93, in ventral view; D, hyoplastra, MP2-308,in ventral view; E, hyoplastra, MP2-175,in ventral view; F, nuchal, MP2-307, in dorsal view; G, hyoplastra, MP3-1, in ventral view. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; CE, cervical; FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral. Scale bar: 3 cm." captionText-7="FIG. 10. — Elements of the shell of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Fuenmayor (La Rioja, Spain):A-E, selection of plates of the individual FM2-1; A, B, almost complete plastron, in ventral (A) and dorsal (B) views; C, D, neurals and proximal region of several adjacent costals, in dorsal view; E, posterior and right postero-lateral margin of the carapace, in dorsal view; F-N, disjointed and isolated plates of several individuals; F, partial nuchal, MCNA 16032, in dorsal view; G-H, epiplastron, FM1-2, in ventral (G) and dorsal (H) views; I, J, epiplastron, FM1-3, in dorsal (I) and ventral (J) views; K-L, anterior area of an anterior plastral lobe, FM1-4, in dorsal (K) and ventral (L) views; M, entoplastron, MCNA 16031, in ventral view; N, xiphiplastron, FM1-1, in ventral view. Abbreviations for the plates (in lower case): c, costal; en, entoplastron; ep, epiplastron; hp, hypoplastron; hy, hyoplastron; n, neural; nu, nuchal; p, peripheral; py, pygal; spy, suprapygal;xi, xiphiplastron. Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; CE, cervical;FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4548567" figureDoi-1="http://doi.org/10.5281/zenodo.4548381" figureDoi-2="http://doi.org/10.5281/zenodo.4548383" figureDoi-3="http://doi.org/10.5281/zenodo.4548385" figureDoi-4="http://doi.org/10.5281/zenodo.4548387" figureDoi-5="http://doi.org/10.5281/zenodo.4548391" figureDoi-6="http://doi.org/10.5281/zenodo.4548393" figureDoi-7="http://doi.org/10.5281/zenodo.4548395" httpUri-0="https://zenodo.org/record/4548567/files/figure.png" httpUri-1="https://zenodo.org/record/4548381/files/figure.png" httpUri-2="https://zenodo.org/record/4548383/files/figure.png" httpUri-3="https://zenodo.org/record/4548385/files/figure.png" httpUri-4="https://zenodo.org/record/4548387/files/figure.png" httpUri-5="https://zenodo.org/record/4548391/files/figure.png" httpUri-6="https://zenodo.org/record/4548393/files/figure.png" httpUri-7="https://zenodo.org/record/4548395/files/figure.png" pageId="5" pageNumber="78">Figs 3-10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5B753E7FFBD654DFCA3C6E9F6F72CBB" lastPageId="9" lastPageNumber="82" pageId="5" pageNumber="78" type="materials_examined">
<paragraph id="ED12006CFFBD654DFCA3C6E9F7072DA4" blockId="5.[812,1455,1785,1835]" lastBlockId="9.[820,1465,1648,1778]" lastPageId="9" lastPageNumber="82" pageId="5" pageNumber="78">
MATERIAL EXAMINED. —
<materialsCitation id="5DC50A31FFBD654DFBAAC6E9F7072DA4" ID-GBIF-Occurrence="3037926301" lastPageId="9" lastPageNumber="82" pageId="5" pageNumber="78">
Numerous disjointed and isolated plates, several partial carapace and plastra, and some relatively complete shells from Monte Pila (e.g.,
<collectionCode id="8BBC98A9FFB1654DFE19C660F2B42DDE" box="[407,448,1648,1672]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="82">MP</collectionCode>
1a-6,
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<figureCitation id="75961CE9FFB1654DFAB9C6B5F6F22DE8" box="[1335,1414,1701,1726]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStartId-0="5.[132,143,1894,1911]" captionStartId-1="7.[132,143,1872,1889]" captionStartId-2="8.[132,143,1917,1934]" captionStartId-3="9.[132,143,1491,1508]" captionStartId-4="10.[132,143,1747,1764]" captionStartId-5="11.[132,143,1779,1796]" captionStartId-6="12.[132,143,1137,1154]" captionTargetBox-1="[134,1455,246,1846]" captionTargetBox-2="[136,1452,218,1848]" captionTargetBox-3="[136,1453,219,1454]" captionTargetBox-4="[135,1455,218,1710]" captionTargetBox-5="[136,1451,216,1737]" captionTargetBox-6="[134,1458,216,1095]" captionTargetId-0="figure-19@6.[132,1455,224,2010]" captionTargetId-1="figure-15@7.[132,1455,243,1849]" captionTargetId-2="figure-19@8.[132,1455,228,1844]" captionTargetId-3="figure-224@9.[132,1455,241,1471]" captionTargetId-4="figure-106@10.[132,1455,214,1715]" captionTargetId-5="figure-88@11.[132,1455,209,1739]" captionTargetId-6="figure-515@12.[129,1452,223,1098]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." captionText-1="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." captionText-2="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-3="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-4="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." captionText-5="FIG. 8. — Articulated elements of the posterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain). All of them are in ventral view: A, MP1a-1345; B, MP1a-1068b; C, MP1a-1423; D, MP1a-1421. Abbreviations for the plates (in lower case): hp, hypoplastron; xi, xiphiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; FE, femoral. Scale bar: 3 cm." captionText-6="FIG. 9. — Plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 2 (A-F) and Monte Pila 3 (G) (Lardero, La Rioja, Spain): A, B, articulated epiplastra, MP2- 306, in ventral (A) and dorsal (B) views; C, xiphiplastron, MP2-93, in ventral view; D, hyoplastra, MP2-308,in ventral view; E, hyoplastra, MP2-175,in ventral view; F, nuchal, MP2-307, in dorsal view; G, hyoplastra, MP3-1, in ventral view. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; CE, cervical; FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4548567" figureDoi-1="http://doi.org/10.5281/zenodo.4548381" figureDoi-2="http://doi.org/10.5281/zenodo.4548383" figureDoi-3="http://doi.org/10.5281/zenodo.4548385" figureDoi-4="http://doi.org/10.5281/zenodo.4548387" figureDoi-5="http://doi.org/10.5281/zenodo.4548391" figureDoi-6="http://doi.org/10.5281/zenodo.4548393" httpUri-0="https://zenodo.org/record/4548567/files/figure.png" httpUri-1="https://zenodo.org/record/4548381/files/figure.png" httpUri-2="https://zenodo.org/record/4548383/files/figure.png" httpUri-3="https://zenodo.org/record/4548385/files/figure.png" httpUri-4="https://zenodo.org/record/4548387/files/figure.png" httpUri-5="https://zenodo.org/record/4548391/files/figure.png" httpUri-6="https://zenodo.org/record/4548393/files/figure.png" pageId="9" pageNumber="82">Figs 3-9</figureCitation>
) and Fuenmayor (e.g., FM1-1, FM1-2, FM1-3, FM1-4, FM2-1, MCNA 16032, MCNA 16031;
<figureCitation id="75961CE9FFB1654DFBAEC6CAF7102DA4" box="[1056,1124,1754,1778]" captionStart="FIG" captionStartId="13.[132,143,1826,1843]" captionTargetBox="[135,1452,218,1736]" captionTargetId="figure-15@13.[132,1455,214,1742]" captionTargetPageId="13" captionText="FIG. 10. — Elements of the shell of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Fuenmayor (La Rioja, Spain):A-E, selection of plates of the individual FM2-1; A, B, almost complete plastron, in ventral (A) and dorsal (B) views; C, D, neurals and proximal region of several adjacent costals, in dorsal view; E, posterior and right postero-lateral margin of the carapace, in dorsal view; F-N, disjointed and isolated plates of several individuals; F, partial nuchal, MCNA 16032, in dorsal view; G-H, epiplastron, FM1-2, in ventral (G) and dorsal (H) views; I, J, epiplastron, FM1-3, in dorsal (I) and ventral (J) views; K-L, anterior area of an anterior plastral lobe, FM1-4, in dorsal (K) and ventral (L) views; M, entoplastron, MCNA 16031, in ventral view; N, xiphiplastron, FM1-1, in ventral view. Abbreviations for the plates (in lower case): c, costal; en, entoplastron; ep, epiplastron; hp, hypoplastron; hy, hyoplastron; n, neural; nu, nuchal; p, peripheral; py, pygal; spy, suprapygal;xi, xiphiplastron. Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; CE, cervical;FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548395" httpUri="https://zenodo.org/record/4548395/files/figure.png" pageId="9" pageNumber="82">Fig. 10</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="B9D250E4FFBD6541FF0AC776F63C2CBF" ID-DOI="http://doi.org/10.5281/zenodo.4548567" ID-Zenodo-Dep="4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="5" pageNumber="78" targetBox="[136,1451,211,2026]" targetPageId="6">
<paragraph id="ED12006CFFBD6541FF0AC776F63C2CBF" blockId="5.[132,1457,1894,2025]" pageId="5" pageNumber="78">
FIG. 3. — Partial shells and carapaces of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFBD6541FD6AC776F0012C21" authorityName="De Stefano" authorityYear="1903" box="[740,885,1894,1911]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFBD6541FCF4C776F2852CD8" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="5" pageNumber="78" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFBD6541FCF4C776F09C2C21" box="[890,1000,1894,1911]" italics="true" pageId="5" pageNumber="78">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFBD6541FC7CC776F7022C21" box="[1010,1142,1894,1911]" italics="true" pageId="5" pageNumber="78">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFBD6541FB17C776F78A2C21" box="[1177,1278,1894,1911]" italics="true" pageId="5" pageNumber="78">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFBD6541FA8CC776F2852CD8" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="5" pageNumber="78" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):
<emphasis id="DFD9DC7EFFBD6541FBC1C76DF7052CD8" bold="true" box="[1103,1137,1917,1934]" pageId="5" pageNumber="78">A -D</emphasis>
, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (
<emphasis id="DFD9DC7EFFBD6541FE37C784F2B22CF3" bold="true" box="[441,454,1940,1957]" pageId="5" pageNumber="78">A</emphasis>
), ventral (
<emphasis id="DFD9DC7EFFBD6541FD99C784F1502CF3" bold="true" box="[535,548,1940,1957]" pageId="5" pageNumber="78">B</emphasis>
), left lateral (
<emphasis id="DFD9DC7EFFBD6541FD00C784F1E82CF3" bold="true" box="[654,668,1940,1957]" pageId="5" pageNumber="78">C</emphasis>
) and anterior (
<emphasis id="DFD9DC7EFFBD6541FC9AC784F0562CF3" bold="true" box="[788,802,1940,1957]" pageId="5" pageNumber="78">D</emphasis>
) views;
<emphasis id="DFD9DC7EFFBD6541FCEBC784F0F22CF3" bold="true" box="[869,902,1940,1957]" pageId="5" pageNumber="78">E -H</emphasis>
, carapace and disjointed posterior half of the plastron, MP1a-643;
<emphasis id="DFD9DC7EFFBD6541FF0AC7BAF3E42CED" bold="true" box="[132,144,1962,1979]" pageId="5" pageNumber="78">E</emphasis>
,
<emphasis id="DFD9DC7EFFBD6541FF15C7BAF3CB2CED" bold="true" box="[155,191,1962,1979]" pageId="5" pageNumber="78">G -H</emphasis>
, carapace, in right lateral (
<emphasis id="DFD9DC7EFFBD6541FE13C7BAF2DD2CED" bold="true" box="[413,425,1962,1979]" pageId="5" pageNumber="78">E</emphasis>
), dorsal (
<emphasis id="DFD9DC7EFFBD6541FE79C7BAF1712CED" bold="true" box="[503,517,1962,1979]" pageId="5" pageNumber="78">G</emphasis>
) and ventral (
<emphasis id="DFD9DC7EFFBD6541FDF9C7BAF1F12CED" bold="true" box="[631,645,1962,1979]" pageId="5" pageNumber="78">H</emphasis>
) views;
<emphasis id="DFD9DC7EFFBD6541FD47C7BAF1A02CED" bold="true" box="[713,724,1962,1979]" pageId="5" pageNumber="78">F</emphasis>
, posterior half of the plastron, in ventral view;
<emphasis id="DFD9DC7EFFBD6541FBD7C7BAF72B2CED" bold="true" box="[1113,1119,1962,1979]" pageId="5" pageNumber="78">I</emphasis>
, partial carapace, MP1a-1070, in dorsal view;
<emphasis id="DFD9DC7EFFBD6541FF3BC7D1F3CB2C84" bold="true" box="[181,191,1985,2002]" pageId="5" pageNumber="78">J</emphasis>
, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type):
<emphasis id="DFD9DC7EFFBD6541FBCEC7D1F73F2C84" bold="true" box="[1088,1099,1985,2002]" pageId="5" pageNumber="78">c</emphasis>
, costal;
<emphasis id="DFD9DC7EFFBD6541FB1DC7D1F7EA2C84" bold="true" box="[1171,1182,1985,2002]" pageId="5" pageNumber="78">n</emphasis>
, neural;
<emphasis id="DFD9DC7EFFBD6541FB69C7D1F7862C84" bold="true" box="[1255,1266,1985,2002]" pageId="5" pageNumber="78">p</emphasis>
, peripheral;
<emphasis id="DFD9DC7EFFBD6541FAD2C7D1F6052C84" bold="true" box="[1372,1393,1985,2002]" pageId="5" pageNumber="78">py</emphasis>
, pygal;
<emphasis id="DFD9DC7EFFBD6541FF0AC7C8F3D72CBF" bold="true" box="[132,163,2008,2025]" pageId="5" pageNumber="78">spy</emphasis>
, suprapygal. Abbreviations for the scutes (in upper case and in bold type):
<emphasis id="DFD9DC7EFFBD6541FC9BC7C8F05B2CBF" bold="true" box="[789,815,2008,2025]" pageId="5" pageNumber="78">CE</emphasis>
, cervical;
<emphasis id="DFD9DC7EFFBD6541FC0DC7C8F0E02CBF" bold="true" box="[899,916,2008,2025]" pageId="5" pageNumber="78">M</emphasis>
, marginal;
<emphasis id="DFD9DC7EFFBD6541FC7EC7C8F77C2CBF" bold="true" box="[1008,1032,2008,2025]" pageId="5" pageNumber="78">PL</emphasis>
, pleural;
<emphasis id="DFD9DC7EFFBD6541FBDBC7C8F7152CBF" bold="true" box="[1109,1121,2008,2025]" pageId="5" pageNumber="78">V</emphasis>
, vertebral. Scale bar: 5 cm.
</paragraph>
</caption>
<caption id="B9D250E4FFBF6543FF0AC740F0C02CBF" ID-DOI="http://doi.org/10.5281/zenodo.4548381" ID-Zenodo-Dep="4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="7" pageNumber="80" startId="7.[132,143,1872,1889]" targetBox="[134,1455,246,1846]" targetPageId="7">
<paragraph id="ED12006CFFBF6543FF0AC740F0C02CBF" blockId="7.[132,1457,1872,2025]" pageId="7" pageNumber="80">
FIG. 4. — Elements of the carapace of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFBF6543FD53C740F01A2C37" authorityName="De Stefano" authorityYear="1903" box="[733,878,1872,1889]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFBF6543FCFAC740F2802C21" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="7" pageNumber="80" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFBF6543FCFAC740F0962C37" box="[884,994,1872,1889]" italics="true" pageId="7" pageNumber="80">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFBF6543FC63C740F7052C37" box="[1005,1137,1872,1889]" italics="true" pageId="7" pageNumber="80">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFBF6543FB18C740F78F2C37" box="[1174,1275,1872,1889]" italics="true" pageId="7" pageNumber="80">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFBF6543FA8FC740F2802C21" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="7" pageNumber="80" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):
<emphasis id="DFD9DC7EFFBF6543FBD5C776F70E2C21" bold="true" box="[1115,1146,1894,1911]" pageId="7" pageNumber="80">A -L</emphasis>
, nuchals and adjacent plates, in dorsal view;
<emphasis id="DFD9DC7EFFBF6543FF5DC76DF3942CD8" bold="true" box="[211,224,1917,1934]" pageId="7" pageNumber="80">A</emphasis>
, MP1a-1570;
<emphasis id="DFD9DC7EFFBF6543FED6C76DF2112CD8" bold="true" box="[344,357,1917,1934]" pageId="7" pageNumber="80">B</emphasis>
, MP1a-1342;
<emphasis id="DFD9DC7EFFBF6543FE53C76DF29F2CD8" bold="true" box="[477,491,1917,1934]" pageId="7" pageNumber="80">C</emphasis>
, MP1a-1364;
<emphasis id="DFD9DC7EFFBF6543FDEDC76DF1052CD8" bold="true" box="[611,625,1917,1934]" pageId="7" pageNumber="80">D</emphasis>
, MP1a-638;
<emphasis id="DFD9DC7EFFBF6543FD51C76DF19F2CD8" bold="true" box="[735,747,1917,1934]" pageId="7" pageNumber="80">E</emphasis>
, MP1a-528;
<emphasis id="DFD9DC7EFFBF6543FCD7C76DF0102CD8" bold="true" box="[857,868,1917,1934]" pageId="7" pageNumber="80">F</emphasis>
, MP1a-228;
<emphasis id="DFD9DC7EFFBF6543FC5CC76DF0942CD8" bold="true" box="[978,992,1917,1934]" pageId="7" pageNumber="80">G</emphasis>
, MP1a-475;
<emphasis id="DFD9DC7EFFBF6543FBC0C76DF7282CD8" bold="true" box="[1102,1116,1917,1934]" pageId="7" pageNumber="80">H</emphasis>
, MP1a-341;
<emphasis id="DFD9DC7EFFBF6543FB44C76DF7A42CD8" bold="true" box="[1226,1232,1917,1934]" pageId="7" pageNumber="80">I</emphasis>
, MP1a-679;
<emphasis id="DFD9DC7EFFBF6543FAB3C76DF6332CD8" bold="true" box="[1341,1351,1917,1934]" pageId="7" pageNumber="80">J</emphasis>
, MP1a-918;
<emphasis id="DFD9DC7EFFBF6543FF0AC784F3E52CF3" bold="true" box="[132,145,1940,1957]" pageId="7" pageNumber="80">K</emphasis>
, MP1b-95;
<emphasis id="DFD9DC7EFFBF6543FF78C784F2752CF3" bold="true" box="[246,257,1940,1957]" pageId="7" pageNumber="80">L</emphasis>
, MP1a-14;
<emphasis id="DFD9DC7EFFBF6543FEEAC784F2FF2CF3" bold="true" box="[356,395,1940,1957]" pageId="7" pageNumber="80">M -O</emphasis>
, neurals and adjacent plates, in dorsal view;
<emphasis id="DFD9DC7EFFBF6543FC89C784F06C2CF3" bold="true" box="[775,792,1940,1957]" pageId="7" pageNumber="80">M</emphasis>
, MP1b-556;
<emphasis id="DFD9DC7EFFBF6543FC08C784F0E02CF3" bold="true" box="[902,916,1940,1957]" pageId="7" pageNumber="80">N</emphasis>
, MP1a-252;
<emphasis id="DFD9DC7EFFBF6543FB8FC784F7642CF3" bold="true" box="[1025,1040,1940,1957]" pageId="7" pageNumber="80">O</emphasis>
, MP1a-707;
<emphasis id="DFD9DC7EFFBF6543FBF3C784F7FD2CF3" bold="true" box="[1149,1161,1940,1957]" pageId="7" pageNumber="80">P</emphasis>
, two articulated costals, MP1a-48, in dorsal view;
<emphasis id="DFD9DC7EFFBF6543FE8AC7BAF2582CED" bold="true" box="[260,300,1962,1979]" pageId="7" pageNumber="80">Q -W</emphasis>
, suprapygals, in dorsal view;
<emphasis id="DFD9DC7EFFBF6543FDA6C7BAF1432CED" bold="true" box="[552,567,1962,1979]" pageId="7" pageNumber="80">Q</emphasis>
, MP1a-1305;
<emphasis id="DFD9DC7EFFBF6543FD3EC7BAF1C92CED" bold="true" box="[688,701,1962,1979]" pageId="7" pageNumber="80">R</emphasis>
, MP1a-1221;
<emphasis id="DFD9DC7EFFBF6543FCB8C7BAF0362CED" bold="true" box="[822,834,1962,1979]" pageId="7" pageNumber="80">S</emphasis>
, MP1b-369;
<emphasis id="DFD9DC7EFFBF6543FC3FC7BAF0C82CED" bold="true" box="[945,956,1962,1979]" pageId="7" pageNumber="80">T</emphasis>
, MP1a-1066;
<emphasis id="DFD9DC7EFFBF6543FBBBC7BAF7372CED" bold="true" box="[1077,1091,1962,1979]" pageId="7" pageNumber="80">U</emphasis>
, MP1a-1350;
<emphasis id="DFD9DC7EFFBF6543FB32C7BAF7BC2CED" bold="true" box="[1212,1224,1962,1979]" pageId="7" pageNumber="80">V</emphasis>
, MP1a-511;
<emphasis id="DFD9DC7EFFBF6543FAB8C7BAF63C2CED" bold="true" box="[1334,1352,1962,1979]" pageId="7" pageNumber="80">W</emphasis>
, MP1a-911;
<emphasis id="DFD9DC7EFFBF6543FF0AC7D1F3D02C84" bold="true" box="[132,164,1985,2002]" pageId="7" pageNumber="80">X -Z</emphasis>
, pygals, in dorsal view;
<emphasis id="DFD9DC7EFFBF6543FEE0C7D1F20E2C84" bold="true" box="[366,378,1985,2002]" pageId="7" pageNumber="80">X</emphasis>
, MP1a-757;
<emphasis id="DFD9DC7EFFBF6543FE69C7D1F2872C84" bold="true" box="[487,499,1985,2002]" pageId="7" pageNumber="80">Y</emphasis>
, MP1a-6;
<emphasis id="DFD9DC7EFFBF6543FDC2C7D1F12C2C84" bold="true" box="[588,600,1985,2002]" pageId="7" pageNumber="80">Z</emphasis>
, MP1a-879. Abbreviations for the plates (in lower case):
<emphasis id="DFD9DC7EFFBF6543FBBDC7D1F74A2C84" bold="true" box="[1075,1086,1985,2002]" pageId="7" pageNumber="80">c</emphasis>
, costal;
<emphasis id="DFD9DC7EFFBF6543FB08C7D1F7E52C84" bold="true" box="[1158,1169,1985,2002]" pageId="7" pageNumber="80">n</emphasis>
, neural;
<emphasis id="DFD9DC7EFFBF6543FB54C7D1F7842C84" bold="true" box="[1242,1264,1985,2002]" pageId="7" pageNumber="80">nu</emphasis>
, nuchal. Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFBF6543FE0FC7C8F2EF2CBF" bold="true" box="[385,411,2008,2025]" pageId="7" pageNumber="80">CE</emphasis>
, cervical;
<emphasis id="DFD9DC7EFFBF6543FE61C7C8F1742CBF" bold="true" box="[495,512,2008,2025]" pageId="7" pageNumber="80">M</emphasis>
, marginal;
<emphasis id="DFD9DC7EFFBF6543FDD3C7C8F1012CBF" bold="true" box="[605,629,2008,2025]" pageId="7" pageNumber="80">PL</emphasis>
, pleural;
<emphasis id="DFD9DC7EFFBF6543FD4FC7C8F1B92CBF" bold="true" box="[705,717,2008,2025]" pageId="7" pageNumber="80">V</emphasis>
, vertebral. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="B9D250E4FFB0654CFF0AC76DF2F52CBF" ID-DOI="http://doi.org/10.5281/zenodo.4548383" ID-Zenodo-Dep="4548383" httpUri="https://zenodo.org/record/4548383/files/figure.png" pageId="8" pageNumber="81" startId="8.[132,143,1917,1934]" targetBox="[136,1452,218,1848]" targetPageId="8">
<paragraph id="ED12006CFFB0654CFF0AC76DF2F52CBF" blockId="8.[132,1457,1917,2025]" pageId="8" pageNumber="81">
FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFB0654CFC1DC76DF7502CD8" authorityName="De Stefano" authorityYear="1903" box="[915,1060,1917,1934]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFB0654CFBA4C76DF1DF2CF3" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="8" pageNumber="81" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFB0654CFBA4C76DF7EC2CD8" box="[1066,1176,1917,1934]" italics="true" pageId="8" pageNumber="81">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFB0654CFB2CC76DF6522CD8" box="[1186,1318,1917,1934]" italics="true" pageId="8" pageNumber="81">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFB0654CFAC4C76DF6DB2CD8" box="[1354,1455,1917,1934]" italics="true" pageId="8" pageNumber="81">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFB0654CFF0AC784F1DF2CF3" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." box="[132,683,1940,1957]" pageId="8" pageNumber="81" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):
<emphasis id="DFD9DC7EFFB0654CFA97C784F6522CF3" bold="true" box="[1305,1318,1940,1957]" pageId="8" pageNumber="81">A</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFABEC784F6492CF3" bold="true" box="[1328,1341,1940,1957]" pageId="8" pageNumber="81">B</emphasis>
, MP1a-1242;
<emphasis id="DFD9DC7EFFB0654CFF0AC7BAF3E62CED" bold="true" box="[132,146,1962,1979]" pageId="8" pageNumber="81">C</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFF13C7BAF3DF2CED" bold="true" box="[157,171,1962,1979]" pageId="8" pageNumber="81">D</emphasis>
, MP1a-1422;
<emphasis id="DFD9DC7EFFB0654CFEADC7BAF25B2CED" bold="true" box="[291,303,1962,1979]" pageId="8" pageNumber="81">E</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFEB5C7BAF2322CED" bold="true" box="[315,326,1962,1979]" pageId="8" pageNumber="81">F</emphasis>
, MP1a-543;
<emphasis id="DFD9DC7EFFB0654CFE3DC7BAF2B52CED" bold="true" box="[435,449,1962,1979]" pageId="8" pageNumber="81">G</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFE43C7BAF2AF2CED" bold="true" box="[461,475,1962,1979]" pageId="8" pageNumber="81">H</emphasis>
, MP1a-277a;
<emphasis id="DFD9DC7EFFB0654CFDDCC7BAF12C2CED" bold="true" box="[594,600,1962,1979]" pageId="8" pageNumber="81">I</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFDEDC7BAF1192CED" bold="true" box="[611,621,1962,1979]" pageId="8" pageNumber="81">J</emphasis>
, MP1a-1594;
<emphasis id="DFD9DC7EFFB0654CFD6BC7BAF1862CED" bold="true" box="[741,754,1962,1979]" pageId="8" pageNumber="81">K</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFD70C7BAF07D2CED" bold="true" box="[766,777,1962,1979]" pageId="8" pageNumber="81">L</emphasis>
, MP1a-1417;
<emphasis id="DFD9DC7EFFB0654CFC0FC7BAF0E62CED" bold="true" box="[897,914,1962,1979]" pageId="8" pageNumber="81">M</emphasis>
,
<emphasis id="DFD9DC7EFFB0654CFC13C7BAF0DF2CED" bold="true" box="[925,939,1962,1979]" pageId="8" pageNumber="81">N</emphasis>
, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case):
<emphasis id="DFD9DC7EFFB0654CFE59C7D1F2992C84" bold="true" box="[471,493,1985,2002]" pageId="8" pageNumber="81">en</emphasis>
, entoplastron;
<emphasis id="DFD9DC7EFFB0654CFDE7C7D1F10B2C84" bold="true" box="[617,639,1985,2002]" pageId="8" pageNumber="81">ep</emphasis>
, epiplastron. Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFB0654CFBECC7D1F7082C84" bold="true" box="[1122,1148,1985,2002]" pageId="8" pageNumber="81">AB</emphasis>
, abdominal;
<emphasis id="DFD9DC7EFFB0654CFB66C7D1F6702C84" bold="true" box="[1256,1284,1985,2002]" pageId="8" pageNumber="81">GU</emphasis>
, gular;
<emphasis id="DFD9DC7EFFB0654CFACFC7D1F6292C84" bold="true" box="[1345,1373,1985,2002]" pageId="8" pageNumber="81">HU</emphasis>
, humeral;
<emphasis id="DFD9DC7EFFB0654CFF0AC7C8F3EA2CBF" bold="true" box="[132,158,2008,2025]" pageId="8" pageNumber="81">PC</emphasis>
, pectoral. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="B9D250E4FFB1654DFF0AC5C3F73D2D69" ID-DOI="http://doi.org/10.5281/zenodo.4548385" ID-Zenodo-Dep="4548385" httpUri="https://zenodo.org/record/4548385/files/figure.png" pageId="9" pageNumber="82" startId="9.[132,143,1491,1508]" targetBox="[136,1453,219,1454]" targetPageId="9">
<paragraph id="ED12006CFFB1654DFF0AC5C3F73D2D69" blockId="9.[132,1457,1491,1599]" pageId="9" pageNumber="82">
FIG. 6. — Plastral plates of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFB1654DFDEAC5C3F1802EB2" authorityName="De Stefano" authorityYear="1903" box="[612,756,1491,1508]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFB1654DFD76C5C3F2292EAD" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="9" pageNumber="82" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFB1654DFD76C5C3F0112EB2" box="[760,869,1491,1508]" italics="true" pageId="9" pageNumber="82">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFB1654DFCE3C5C3F0842EB2" box="[877,1008,1491,1508]" italics="true" pageId="9" pageNumber="82">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFB1654DFB9FC5C3F7012EB2" box="[1041,1141,1491,1508]" italics="true" pageId="9" pageNumber="82">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFB1654DFBF6C5C3F2292EAD" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="9" pageNumber="82" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999</bibRefCitation>
</taxonomicName>
,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):
<emphasis id="DFD9DC7EFFB1654DFC22C5FAF0B82EAD" bold="true" box="[940,972,1514,1531]" pageId="9" pageNumber="82">A -V</emphasis>
, epiplastra,in dorsal and ventral views;
<emphasis id="DFD9DC7EFFB1654DFA82C5FAF66D2EAD" bold="true" box="[1292,1305,1514,1531]" pageId="9" pageNumber="82">A</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFAACC5FAF65B2EAD" bold="true" box="[1314,1327,1514,1531]" pageId="9" pageNumber="82">B</emphasis>
, MP1b-sup1-a;
<emphasis id="DFD9DC7EFFB1654DFF0AC611F3E62D44" bold="true" box="[132,146,1537,1554]" pageId="9" pageNumber="82">C</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFF12C611F3DE2D44" bold="true" box="[156,170,1537,1554]" pageId="9" pageNumber="82">D</emphasis>
, MP1a-55;
<emphasis id="DFD9DC7EFFB1654DFE87C611F2612D44" bold="true" box="[265,277,1537,1554]" pageId="9" pageNumber="82">E</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFE91C611F25E2D44" bold="true" box="[287,298,1537,1554]" pageId="9" pageNumber="82">F</emphasis>
, MP1a-1210a;
<emphasis id="DFD9DC7EFFB1654DFE27C611F2C32D44" bold="true" box="[425,439,1537,1554]" pageId="9" pageNumber="82">G</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFE4EC611F2BA2D44" bold="true" box="[448,462,1537,1554]" pageId="9" pageNumber="82">H</emphasis>
, MP1a-975a;
<emphasis id="DFD9DC7EFFB1654DFDCCC611F13C2D44" bold="true" box="[578,584,1537,1554]" pageId="9" pageNumber="82">I</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFDDFC611F12F2D44" bold="true" box="[593,603,1537,1554]" pageId="9" pageNumber="82">J</emphasis>
, MP1a-1145a;
<emphasis id="DFD9DC7EFFB1654DFD54C611F1932D44" bold="true" box="[730,743,1537,1554]" pageId="9" pageNumber="82">K</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFD7FC611F1882D44" bold="true" box="[753,764,1537,1554]" pageId="9" pageNumber="82">L</emphasis>
, MP1a-1745;
<emphasis id="DFD9DC7EFFB1654DFCFFC611F0F62D44" bold="true" box="[881,898,1537,1554]" pageId="9" pageNumber="82">M</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFC05C611F0ED2D44" bold="true" box="[907,921,1537,1554]" pageId="9" pageNumber="82">N</emphasis>
, MP1b-481;
<emphasis id="DFD9DC7EFFB1654DFB8AC611F7672D44" bold="true" box="[1028,1043,1537,1554]" pageId="9" pageNumber="82">O</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFB92C611F75C2D44" bold="true" box="[1052,1064,1537,1554]" pageId="9" pageNumber="82">P</emphasis>
, MP1a-1679b;
<emphasis id="DFD9DC7EFFB1654DFB26C611F7C32D44" bold="true" box="[1192,1207,1537,1554]" pageId="9" pageNumber="82">Q</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFB4EC611F7B92D44" bold="true" box="[1216,1229,1537,1554]" pageId="9" pageNumber="82">R</emphasis>
, MP1a-1189a;
<emphasis id="DFD9DC7EFFB1654DFAC3C611F62D2D44" bold="true" box="[1357,1369,1537,1554]" pageId="9" pageNumber="82">S</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFAECC611F6192D44" bold="true" box="[1378,1389,1537,1554]" pageId="9" pageNumber="82">T</emphasis>
, MP1a- 1557;
<emphasis id="DFD9DC7EFFB1654DFF36C607F3B22D7E" bold="true" box="[184,198,1559,1576]" pageId="9" pageNumber="82">U</emphasis>
,
<emphasis id="DFD9DC7EFFB1654DFF5FC607F3A92D7E" bold="true" box="[209,221,1559,1576]" pageId="9" pageNumber="82">V</emphasis>
, MP1a-529b;
<emphasis id="DFD9DC7EFFB1654DFEDAC607F2FC2D7E" bold="true" box="[340,392,1559,1576]" pageId="9" pageNumber="82">W -AC</emphasis>
, entoplastral, in ventral view;
<emphasis id="DFD9DC7EFFB1654DFD0FC607F1E72D7E" bold="true" box="[641,659,1559,1576]" pageId="9" pageNumber="82">W</emphasis>
, MP1a-1394b;
<emphasis id="DFD9DC7EFFB1654DFC9AC607F0542D7E" bold="true" box="[788,800,1559,1576]" pageId="9" pageNumber="82">X</emphasis>
, MP1b-1597a;
<emphasis id="DFD9DC7EFFB1654DFC2DC607F0DB2D7E" bold="true" box="[931,943,1559,1576]" pageId="9" pageNumber="82">Y</emphasis>
, MP1a-347a;
<emphasis id="DFD9DC7EFFB1654DFBA8C607F7462D7E" bold="true" box="[1062,1074,1559,1576]" pageId="9" pageNumber="82">Z</emphasis>
, MP1a-672;
<emphasis id="DFD9DC7EFFB1654DFB10C607F7CC2D7E" bold="true" box="[1182,1208,1559,1576]" pageId="9" pageNumber="82">AA</emphasis>
, MP1a-449;
<emphasis id="DFD9DC7EFFB1654DFAAAC607F64A2D7E" bold="true" box="[1316,1342,1559,1576]" pageId="9" pageNumber="82">AB</emphasis>
, MP1a-180a;
<emphasis id="DFD9DC7EFFB1654DFF0AC63EF3EB2D69" bold="true" box="[132,159,1582,1599]" pageId="9" pageNumber="82">AC</emphasis>
, MP1a-409. Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFB1654DFDF1C63EF1EF2D69" bold="true" box="[639,667,1582,1599]" pageId="9" pageNumber="82">GU</emphasis>
, gular;
<emphasis id="DFD9DC7EFFB1654DFD57C63EF1812D69" bold="true" box="[729,757,1582,1599]" pageId="9" pageNumber="82">HU</emphasis>
, humeral;
<emphasis id="DFD9DC7EFFB1654DFCC3C63EF0132D69" bold="true" box="[845,871,1582,1599]" pageId="9" pageNumber="82">PC</emphasis>
, pectoral. Scale bar: 3 cm.
</paragraph>
</caption>
<paragraph id="ED12006CFFB1654DFCBBC70CF6F72CBB" blockId="9.[820,1465,1820,2029]" pageId="9" pageNumber="82">
LOCALITY AND HORIZON. —
<materialsCitation id="5DC50A31FFB1654DFBC7C70CF6F72CBB" ID-GBIF-Occurrence="3037926302" pageId="9" pageNumber="82">
Most specimens come from the Monte Pila sites, municipality of Lardero, Autonomous Community of
<collectingRegion id="2F69CE8EFFB1654DFA10C727F01F2C3F" country="Spain" name="La Rioja" pageId="9" pageNumber="82">La Rioja</collectingRegion>
,
<collectingCountry id="95BA40FCFFB1654DFCFBC741F0DB2C3F" box="[885,943,1873,1897]" name="Spain" pageId="9" pageNumber="82">Spain</collectingCountry>
. Levels
<collectionCode id="8BBC98A9FFB1654DFC77C741F7502C3F" box="[1017,1060,1873,1897]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="82">MP</collectionCode>
1,
<collectionCode id="8BBC98A9FFB1654DFBB6C741F72A2C3F" box="[1080,1118,1873,1897]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="82">MP</collectionCode>
2 and
<collectionCode id="8BBC98A9FFB1654DFB14C741F7B52C3F" box="[1178,1217,1873,1897]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="82">MP</collectionCode>
3 (
<figureCitation id="75961CE9FFB1654DFB55C741F65E2C3F" box="[1243,1322,1873,1897]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStartId-0="5.[132,143,1894,1911]" captionStartId-1="7.[132,143,1872,1889]" captionStartId-2="8.[132,143,1917,1934]" captionStartId-3="9.[132,143,1491,1508]" captionStartId-4="10.[132,143,1747,1764]" captionStartId-5="11.[132,143,1779,1796]" captionStartId-6="12.[132,143,1137,1154]" captionTargetBox-1="[134,1455,246,1846]" captionTargetBox-2="[136,1452,218,1848]" captionTargetBox-3="[136,1453,219,1454]" captionTargetBox-4="[135,1455,218,1710]" captionTargetBox-5="[136,1451,216,1737]" captionTargetBox-6="[134,1458,216,1095]" captionTargetId-0="figure-19@6.[132,1455,224,2010]" captionTargetId-1="figure-15@7.[132,1455,243,1849]" captionTargetId-2="figure-19@8.[132,1455,228,1844]" captionTargetId-3="figure-224@9.[132,1455,241,1471]" captionTargetId-4="figure-106@10.[132,1455,214,1715]" captionTargetId-5="figure-88@11.[132,1455,209,1739]" captionTargetId-6="figure-515@12.[129,1452,223,1098]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." captionText-1="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." captionText-2="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-3="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-4="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." captionText-5="FIG. 8. — Articulated elements of the posterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain). All of them are in ventral view: A, MP1a-1345; B, MP1a-1068b; C, MP1a-1423; D, MP1a-1421. Abbreviations for the plates (in lower case): hp, hypoplastron; xi, xiphiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; FE, femoral. Scale bar: 3 cm." captionText-6="FIG. 9. — Plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 2 (A-F) and Monte Pila 3 (G) (Lardero, La Rioja, Spain): A, B, articulated epiplastra, MP2- 306, in ventral (A) and dorsal (B) views; C, xiphiplastron, MP2-93, in ventral view; D, hyoplastra, MP2-308,in ventral view; E, hyoplastra, MP2-175,in ventral view; F, nuchal, MP2-307, in dorsal view; G, hyoplastra, MP3-1, in ventral view. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; CE, cervical; FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4548567" figureDoi-1="http://doi.org/10.5281/zenodo.4548381" figureDoi-2="http://doi.org/10.5281/zenodo.4548383" figureDoi-3="http://doi.org/10.5281/zenodo.4548385" figureDoi-4="http://doi.org/10.5281/zenodo.4548387" figureDoi-5="http://doi.org/10.5281/zenodo.4548391" figureDoi-6="http://doi.org/10.5281/zenodo.4548393" httpUri-0="https://zenodo.org/record/4548567/files/figure.png" httpUri-1="https://zenodo.org/record/4548381/files/figure.png" httpUri-2="https://zenodo.org/record/4548383/files/figure.png" httpUri-3="https://zenodo.org/record/4548385/files/figure.png" httpUri-4="https://zenodo.org/record/4548387/files/figure.png" httpUri-5="https://zenodo.org/record/4548391/files/figure.png" httpUri-6="https://zenodo.org/record/4548393/files/figure.png" pageId="9" pageNumber="82">Figs 3-9</figureCitation>
). Other specimens came from the site of Fuenmayor, municipality of Fuenmayor, Autonomous Community of
<collectingRegion id="2F69CE8EFFB1654DFBD2C796F7C72CC8" box="[1116,1203,1926,1950]" country="Spain" name="La Rioja" pageId="9" pageNumber="82">La Rioja</collectingRegion>
,
<collectingCountry id="95BA40FCFFB1654DFB4EC796F7832CC8" box="[1216,1271,1926,1950]" name="Spain" pageId="9" pageNumber="82">Spain</collectingCountry>
(
<figureCitation id="75961CE9FFB1654DFA87C796F6252CC8" box="[1289,1361,1926,1950]" captionStart="FIG" captionStartId="13.[132,143,1826,1843]" captionTargetBox="[135,1452,218,1736]" captionTargetId="figure-15@13.[132,1455,214,1742]" captionTargetPageId="13" captionText="FIG. 10. — Elements of the shell of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Fuenmayor (La Rioja, Spain):A-E, selection of plates of the individual FM2-1; A, B, almost complete plastron, in ventral (A) and dorsal (B) views; C, D, neurals and proximal region of several adjacent costals, in dorsal view; E, posterior and right postero-lateral margin of the carapace, in dorsal view; F-N, disjointed and isolated plates of several individuals; F, partial nuchal, MCNA 16032, in dorsal view; G-H, epiplastron, FM1-2, in ventral (G) and dorsal (H) views; I, J, epiplastron, FM1-3, in dorsal (I) and ventral (J) views; K-L, anterior area of an anterior plastral lobe, FM1-4, in dorsal (K) and ventral (L) views; M, entoplastron, MCNA 16031, in ventral view; N, xiphiplastron, FM1-1, in ventral view. Abbreviations for the plates (in lower case): c, costal; en, entoplastron; ep, epiplastron; hp, hypoplastron; hy, hyoplastron; n, neural; nu, nuchal; p, peripheral; py, pygal; spy, suprapygal;xi, xiphiplastron. Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; CE, cervical;FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548395" httpUri="https://zenodo.org/record/4548395/files/figure.png" pageId="9" pageNumber="82">Fig. 10</figureCitation>
). Western part of the Ebro Basin. Transitional facies between the Najera and Haro formations. Local zone Y, biozone
<collectionCode id="8BBC98A9FFB1654DFB35C7ABF79C2C85" box="[1211,1256,1979,2003]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="9" pageNumber="82">MN</collectionCode>
2, lower Miocene (see
<figureCitation id="75961CE9FFB1654DFCBBC7C5F01B2CBB" box="[821,879,2005,2029]" captionStart="FIG" captionStartId="4.[132,143,1173,1190]" captionTargetBox="[138,1453,213,1132]" captionTargetId="figure-433@4.[132,1455,213,1137]" captionTargetPageId="4" captionText="FIG. 1. — Geographic and geological position of the fossil areas from which the remains of turtles studied in this paper come. 1, Fuenmayor (La Rioja); 2, Monte Pila (La Rioja); 3, the Bardenas Reales of Navarre (Navarre). All of them are located in the Spanish Ebro Basin." figureDoi="http://doi.org/10.5281/zenodo.4548377" httpUri="https://zenodo.org/record/4548377/files/figure.png" pageId="9" pageNumber="82">Figs 1</figureCitation>
,
<figureCitation id="75961CE9FFB1654DFCF2C7C5F0FD2CBB" box="[892,905,2005,2029]" captionStart="FIG" captionStartId="5.[132,143,1499,1516]" captionTargetBox="[132,775,215,1463]" captionTargetId="graphics-531@5.[132,814,215,1464]" captionTargetPageId="5" captionText="FIG. 2. — Biostratigraphic diagram with the fossil areas studied in this paper situated in a stratigraphical table.The three outcrops indicated in the Bardenas Reales of Navarre correspond to Barranco de Tudela (BT), Nasa (N) and Rincón del Bu (RB). The chronological data are based on Larrasoaña et al. (2006) and Suarez-Hernando (2017)." figureDoi="http://doi.org/10.5281/zenodo.4548379" httpUri="https://zenodo.org/record/4548379/files/figure.png" pageId="9" pageNumber="82">2</figureCitation>
and the section Geographic and geological setting).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="B9D250E4FFB2654EFF0AC6C3F6C52C69" ID-DOI="http://doi.org/10.5281/zenodo.4548387" ID-Zenodo-Dep="4548387" httpUri="https://zenodo.org/record/4548387/files/figure.png" pageId="10" pageNumber="83" startId="10.[132,143,1747,1764]" targetBox="[135,1455,218,1710]" targetPageId="10">
<paragraph id="ED12006CFFB2654EFF0AC6C3F6C52C69" blockId="10.[132,1457,1747,1855]" pageId="10" pageNumber="83">
FIG. 7. — Plastral plates of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFB2654EFDEAC6C3F1802DB2" authorityName="De Stefano" authorityYear="1903" box="[612,756,1747,1764]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFB2654EFD76C6C3F2122DAD" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="10" pageNumber="83" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFB2654EFD76C6C3F0112DB2" box="[760,869,1747,1764]" italics="true" pageId="10" pageNumber="83">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFB2654EFCE3C6C3F0842DB2" box="[877,1008,1747,1764]" italics="true" pageId="10" pageNumber="83">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFB2654EFB9FC6C3F7012DB2" box="[1041,1141,1747,1764]" italics="true" pageId="10" pageNumber="83">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFB2654EFBF6C6C3F2122DAD" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="10" pageNumber="83" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):
<emphasis id="DFD9DC7EFFB2654EFC54C6FAF0802DAD" bold="true" box="[986,1012,1770,1787]" pageId="10" pageNumber="83">A -I</emphasis>
, hyoplastra, in ventral view;
<emphasis id="DFD9DC7EFFB2654EFB6AC6FAF7852DAD" bold="true" box="[1252,1265,1770,1787]" pageId="10" pageNumber="83">A</emphasis>
, MP1a-694;
<emphasis id="DFD9DC7EFFB2654EFAD1C6FAF6182DAD" bold="true" box="[1375,1388,1770,1787]" pageId="10" pageNumber="83">B</emphasis>
, MP1a- 385;
<emphasis id="DFD9DC7EFFB2654EFF20C710F3C82C47" bold="true" box="[174,188,1792,1809]" pageId="10" pageNumber="83">C</emphasis>
, MP1a-1333;
<emphasis id="DFD9DC7EFFB2654EFEBAC710F2362C47" bold="true" box="[308,322,1792,1809]" pageId="10" pageNumber="83">D</emphasis>
, MP1a-1025;
<emphasis id="DFD9DC7EFFB2654EFE37C710F2B12C47" bold="true" box="[441,453,1792,1809]" pageId="10" pageNumber="83">E</emphasis>
, MP1a-281;
<emphasis id="DFD9DC7EFFB2654EFDBCC710F1492C47" bold="true" box="[562,573,1792,1809]" pageId="10" pageNumber="83">F</emphasis>
, MP1a-1227;
<emphasis id="DFD9DC7EFFB2654EFD3AC710F1B62C47" bold="true" box="[692,706,1792,1809]" pageId="10" pageNumber="83">G</emphasis>
, MP1b-585;
<emphasis id="DFD9DC7EFFB2654EFCBFC710F04B2C47" bold="true" box="[817,831,1792,1809]" pageId="10" pageNumber="83">H</emphasis>
, MP1a-109;
<emphasis id="DFD9DC7EFFB2654EFC22C710F0C62C47" bold="true" box="[940,946,1792,1809]" pageId="10" pageNumber="83">I</emphasis>
, MP1b-169;
<emphasis id="DFD9DC7EFFB2654EFB91C710F7362C47" bold="true" box="[1055,1090,1792,1809]" pageId="10" pageNumber="83">J -M</emphasis>
, hypoplastra, in ventral view;
<emphasis id="DFD9DC7EFFB2654EFAB2C710F6322C47" bold="true" box="[1340,1350,1792,1809]" pageId="10" pageNumber="83">J</emphasis>
, MP1b-467;
<emphasis id="DFD9DC7EFFB2654EFF0AC707F3E52C7E" bold="true" box="[132,145,1815,1832]" pageId="10" pageNumber="83">K</emphasis>
, MP1a-1194;
<emphasis id="DFD9DC7EFFB2654EFE89C707F2662C7E" bold="true" box="[263,274,1815,1832]" pageId="10" pageNumber="83">L</emphasis>
, MP1b-456;
<emphasis id="DFD9DC7EFFB2654EFEF0C707F2FB2C7E" bold="true" box="[382,399,1815,1832]" pageId="10" pageNumber="83">M</emphasis>
, MP1a-367;
<emphasis id="DFD9DC7EFFB2654EFE74C707F1692C7E" bold="true" box="[506,541,1815,1832]" pageId="10" pageNumber="83">N -U</emphasis>
, xiphiplastra,in ventral view;
<emphasis id="DFD9DC7EFFB2654EFC9EC707F06A2C7E" bold="true" box="[784,798,1815,1832]" pageId="10" pageNumber="83">N</emphasis>
, MP1a-865;
<emphasis id="DFD9DC7EFFB2654EFC06C707F0E32C7E" bold="true" box="[904,919,1815,1832]" pageId="10" pageNumber="83">O</emphasis>
, MP1a-56;
<emphasis id="DFD9DC7EFFB2654EFC79C707F7772C7E" bold="true" box="[1015,1027,1815,1832]" pageId="10" pageNumber="83">P</emphasis>
, MP1a-659;
<emphasis id="DFD9DC7EFFB2654EFBE0C707F7092C7E" bold="true" box="[1134,1149,1815,1832]" pageId="10" pageNumber="83">Q</emphasis>
, MP1a-681;
<emphasis id="DFD9DC7EFFB2654EFB66C707F7812C7E" bold="true" box="[1256,1269,1815,1832]" pageId="10" pageNumber="83">R</emphasis>
, MP1a-474;
<emphasis id="DFD9DC7EFFB2654EFAEEC707F6182C7E" bold="true" box="[1376,1388,1815,1832]" pageId="10" pageNumber="83">S</emphasis>
, MP1b- 380b;
<emphasis id="DFD9DC7EFFB2654EFF36C73EF3B72C69" bold="true" box="[184,195,1838,1855]" pageId="10" pageNumber="83">T</emphasis>
, MP1a-81;
<emphasis id="DFD9DC7EFFB2654EFEADC73EF2452C69" bold="true" box="[291,305,1838,1855]" pageId="10" pageNumber="83">U</emphasis>
, MP1a-263.Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFB2654EFC87C73EF0572C69" bold="true" box="[777,803,1838,1855]" pageId="10" pageNumber="83">AB</emphasis>
, abdominal;
<emphasis id="DFD9DC7EFFB2654EFC00C73EF0DD2C69" bold="true" box="[910,937,1838,1855]" pageId="10" pageNumber="83">AN</emphasis>
, anal;
<emphasis id="DFD9DC7EFFB2654EFC50C73EF0812C69" bold="true" box="[990,1013,1838,1855]" pageId="10" pageNumber="83">FE</emphasis>
, femoral;
<emphasis id="DFD9DC7EFFB2654EFBC9C73EF7172C69" bold="true" box="[1095,1123,1838,1855]" pageId="10" pageNumber="83">HU</emphasis>
, humeral;
<emphasis id="DFD9DC7EFFB2654EFB36C73EF7A62C69" bold="true" box="[1208,1234,1838,1855]" pageId="10" pageNumber="83">PC</emphasis>
, pectoral. Scale bar:3 cm.
</paragraph>
</caption>
<subSubSection id="A5B753E7FFB2654BFF0AC762F0732A64" lastPageId="15" lastPageNumber="88" pageId="10" pageNumber="83" type="description">
<paragraph id="ED12006CFFB2654EFF0AC762F2612CDC" blockId="10.[130,776,1906,2028]" box="[132,277,1906,1932]" pageId="10" pageNumber="83">DESCRIPTION</paragraph>
<paragraph id="ED12006CFFB2654FFF0AC781F6C42CBD" blockId="10.[130,776,1906,2028]" lastBlockId="11.[810,1456,1937,2028]" lastPageId="11" lastPageNumber="84" pageId="10" pageNumber="83">
The maximum length of the shells of the larger specimens of the freshwater turtles identified at both Monte Pila and Fuenmayor is close to
<quantity id="2A55AD89FFB2654EFF69C7C1F25C2CBA" box="[231,296,2001,2028]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="83" unit="cm" value="20.0">20 cm</quantity>
(
<figureCitation id="75961CE9FFB2654EFEB9C7C1F2E22CBD" box="[311,406,2001,2028]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStartId-0="5.[132,143,1894,1911]" captionStartId-1="7.[132,143,1872,1889]" captionStartId-2="8.[132,143,1917,1934]" captionStartId-3="9.[132,143,1491,1508]" captionStartId-4="10.[132,143,1747,1764]" captionStartId-5="11.[132,143,1779,1796]" captionStartId-6="12.[132,143,1137,1154]" captionStartId-7="13.[132,143,1826,1843]" captionTargetBox-1="[134,1455,246,1846]" captionTargetBox-2="[136,1452,218,1848]" captionTargetBox-3="[136,1453,219,1454]" captionTargetBox-4="[135,1455,218,1710]" captionTargetBox-5="[136,1451,216,1737]" captionTargetBox-6="[134,1458,216,1095]" captionTargetBox-7="[135,1452,218,1736]" captionTargetId-0="figure-19@6.[132,1455,224,2010]" captionTargetId-1="figure-15@7.[132,1455,243,1849]" captionTargetId-2="figure-19@8.[132,1455,228,1844]" captionTargetId-3="figure-224@9.[132,1455,241,1471]" captionTargetId-4="figure-106@10.[132,1455,214,1715]" captionTargetId-5="figure-88@11.[132,1455,209,1739]" captionTargetId-6="figure-515@12.[129,1452,223,1098]" captionTargetId-7="figure-15@13.[132,1455,214,1742]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionText-0="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." captionText-1="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." captionText-2="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-3="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." captionText-4="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." captionText-5="FIG. 8. — Articulated elements of the posterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain). All of them are in ventral view: A, MP1a-1345; B, MP1a-1068b; C, MP1a-1423; D, MP1a-1421. Abbreviations for the plates (in lower case): hp, hypoplastron; xi, xiphiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; FE, femoral. Scale bar: 3 cm." captionText-6="FIG. 9. — Plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 2 (A-F) and Monte Pila 3 (G) (Lardero, La Rioja, Spain): A, B, articulated epiplastra, MP2- 306, in ventral (A) and dorsal (B) views; C, xiphiplastron, MP2-93, in ventral view; D, hyoplastra, MP2-308,in ventral view; E, hyoplastra, MP2-175,in ventral view; F, nuchal, MP2-307, in dorsal view; G, hyoplastra, MP3-1, in ventral view. Abbreviations for the scutes (in upper case): AB, abdominal; AN, anal; CE, cervical; FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral. Scale bar: 3 cm." captionText-7="FIG. 10. — Elements of the shell of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Fuenmayor (La Rioja, Spain):A-E, selection of plates of the individual FM2-1; A, B, almost complete plastron, in ventral (A) and dorsal (B) views; C, D, neurals and proximal region of several adjacent costals, in dorsal view; E, posterior and right postero-lateral margin of the carapace, in dorsal view; F-N, disjointed and isolated plates of several individuals; F, partial nuchal, MCNA 16032, in dorsal view; G-H, epiplastron, FM1-2, in ventral (G) and dorsal (H) views; I, J, epiplastron, FM1-3, in dorsal (I) and ventral (J) views; K-L, anterior area of an anterior plastral lobe, FM1-4, in dorsal (K) and ventral (L) views; M, entoplastron, MCNA 16031, in ventral view; N, xiphiplastron, FM1-1, in ventral view. Abbreviations for the plates (in lower case): c, costal; en, entoplastron; ep, epiplastron; hp, hypoplastron; hy, hyoplastron; n, neural; nu, nuchal; p, peripheral; py, pygal; spy, suprapygal;xi, xiphiplastron. Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; CE, cervical;FE, femoral; GU, gular; HU, humeral; M, marginal; PC, pectoral; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4548567" figureDoi-1="http://doi.org/10.5281/zenodo.4548381" figureDoi-2="http://doi.org/10.5281/zenodo.4548383" figureDoi-3="http://doi.org/10.5281/zenodo.4548385" figureDoi-4="http://doi.org/10.5281/zenodo.4548387" figureDoi-5="http://doi.org/10.5281/zenodo.4548391" figureDoi-6="http://doi.org/10.5281/zenodo.4548393" figureDoi-7="http://doi.org/10.5281/zenodo.4548395" httpUri-0="https://zenodo.org/record/4548567/files/figure.png" httpUri-1="https://zenodo.org/record/4548381/files/figure.png" httpUri-2="https://zenodo.org/record/4548383/files/figure.png" httpUri-3="https://zenodo.org/record/4548385/files/figure.png" httpUri-4="https://zenodo.org/record/4548387/files/figure.png" httpUri-5="https://zenodo.org/record/4548391/files/figure.png" httpUri-6="https://zenodo.org/record/4548393/files/figure.png" httpUri-7="https://zenodo.org/record/4548395/files/figure.png" pageId="10" pageNumber="83">Figs 3-10</figureCitation>
). However, elements corresponding to individuals of very varied sizes are identified, some of them with a length of less than half that of the largest specimens. Some of the most complete shells, as well as the articulated elements of other specimens recognized as adult individuals due to their size, show fusion of many of their sutures, so that the margins of each plate cannot be recognized in them (
<figureCitation id="75961CE9FFB3654FFD2FC7A1F1AA2C9D" box="[673,734,1969,1995]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="11" pageNumber="84">Figs 3</figureCitation>
;
<figureCitation id="75961CE9FFB3654FFD67C7A1F1832C9A" box="[745,759,1969,1996]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[136,1452,218,1848]" captionTargetId="figure-19@8.[132,1455,228,1844]" captionTargetPageId="8" captionText="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548383" httpUri="https://zenodo.org/record/4548383/files/figure.png" pageId="11" pageNumber="84">5</figureCitation>
). The thickness of the plates is greater in the larger individuals. Although the width/length ratio of the shell is variable, this form shows a relatively wide carapace, being recognized as slightly longer than wide in some cases (
<figureCitation id="75961CE9FFB3654FFBFEC7C1F7B42CBD" box="[1136,1216,2001,2027]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="11" pageNumber="84">Fig.3G</figureCitation>
). The carapaces are low.
</paragraph>
<caption id="B9D250E4FFB3654FFF0AC6E3F0B02C1E" ID-DOI="http://doi.org/10.5281/zenodo.4548391" ID-Zenodo-Dep="4548391" httpUri="https://zenodo.org/record/4548391/files/figure.png" pageId="11" pageNumber="84" startId="11.[132,143,1779,1796]" targetBox="[136,1451,216,1737]" targetPageId="11">
<paragraph id="ED12006CFFB3654FFF0AC6E3F0B02C1E" blockId="11.[132,1457,1779,1864]" pageId="11" pageNumber="84">
FIG. 8. — Articulated elements of the posterior plastral lobe of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFB3654FFC18C6E3F7532C52" authorityName="De Stefano" authorityYear="1903" box="[918,1063,1779,1796]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFB3654FFBA2C6E3F1D92C4D" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="11" pageNumber="84" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFB3654FFBA2C6E3F7EE2C52" box="[1068,1178,1779,1796]" italics="true" pageId="11" pageNumber="84">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFB3654FFB2DC6E3F6532C52" box="[1187,1319,1779,1796]" italics="true" pageId="11" pageNumber="84">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFB3654FFAC4C6E3F6DB2C52" box="[1354,1455,1779,1796]" italics="true" pageId="11" pageNumber="84">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFB3654FFF0AC71AF1D92C4D" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." box="[132,685,1802,1819]" pageId="11" pageNumber="84" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain). All of them are in ventral view:
<emphasis id="DFD9DC7EFFB3654FFF7AC730F2752C67" bold="true" box="[244,257,1824,1841]" pageId="11" pageNumber="84">A</emphasis>
, MP1a-1345;
<emphasis id="DFD9DC7EFFB3654FFEF7C730F2F22C67" bold="true" box="[377,390,1824,1841]" pageId="11" pageNumber="84">B</emphasis>
, MP1a-1068b;
<emphasis id="DFD9DC7EFFB3654FFD84C730F16C2C67" bold="true" box="[522,536,1824,1841]" pageId="11" pageNumber="84">C</emphasis>
, MP1a-1423;
<emphasis id="DFD9DC7EFFB3654FFD1EC730F1EA2C67" bold="true" box="[656,670,1824,1841]" pageId="11" pageNumber="84">D</emphasis>
, MP1a-1421. Abbreviations for the plates (in lower case):
<emphasis id="DFD9DC7EFFB3654FFB04C730F7D42C67" bold="true" box="[1162,1184,1824,1841]" pageId="11" pageNumber="84">hp</emphasis>
, hypoplastron;
<emphasis id="DFD9DC7EFFB3654FFAAAC730F6472C67" bold="true" box="[1316,1331,1824,1841]" pageId="11" pageNumber="84">xi</emphasis>
, xiphiplastron. Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFB3654FFE76C727F1662C1E" bold="true" box="[504,530,1847,1864]" pageId="11" pageNumber="84">AB</emphasis>
, abdominal;
<emphasis id="DFD9DC7EFFB3654FFDF0C727F1ED2C1E" bold="true" box="[638,665,1847,1864]" pageId="11" pageNumber="84">AN</emphasis>
, anal;
<emphasis id="DFD9DC7EFFB3654FFD5EC727F1932C1E" bold="true" box="[720,743,1847,1864]" pageId="11" pageNumber="84">FE</emphasis>
, femoral. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="B9D250E4FFB46548FF0AC461F0522F8A" ID-DOI="http://doi.org/10.5281/zenodo.4548393" ID-Zenodo-Dep="4548393" httpUri="https://zenodo.org/record/4548393/files/figure.png" pageId="12" pageNumber="85" startId="12.[132,143,1137,1154]" targetBox="[134,1458,216,1095]" targetPageId="12">
<paragraph id="ED12006CFFB46548FF0AC461F0522F8A" blockId="12.[132,1457,1137,1244]" pageId="12" pageNumber="85">
FIG. 9. — Plates of several individuals of freshwater
<taxonomicName id="2AAD7BEFFFB46548FDB1C461F1A52FD4" authorityName="De Stefano" authorityYear="1903" box="[575,721,1137,1154]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="78" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
<taxonomicName id="2AAD7BEFFFB46548FD56C461F2162FCE" authority="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999" authorityName="Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia" authorityYear="1999" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="12" pageNumber="85" phylum="Basidiomycota" rank="species" species="bardenensis" subGenus="Temnoclemmys">
<emphasis id="DFD9DC7EFFB46548FD56C461F0322FD4" box="[728,838,1137,1154]" italics="true" pageId="12" pageNumber="85">Ptychogaster</emphasis>
(
<emphasis id="DFD9DC7EFFB46548FCDCC461F0A22FD4" box="[850,982,1137,1154]" italics="true" pageId="12" pageNumber="85">Temnoclemmys</emphasis>
) cf.
<emphasis id="DFD9DC7EFFB46548FC73C461F7172FD4" box="[1021,1123,1137,1154]" italics="true" pageId="12" pageNumber="85">bardenensis</emphasis>
<bibRefCitation id="893C7D9DFFB46548FBE7C461F2162FCE" author="MURELAGA X. &amp; LAPPARENT DE BROIN F. DE &amp; PEREDA SUBERBIOLA X. &amp; ASTIBIA H." pageId="12" pageNumber="85" pagination="423 - 429" refId="ref13766" refString="MURELAGA X., LAPPARENT DE BROIN F. DE, PEREDA SUBERBIOLA X. &amp; ASTIBIA H. 1999. - Deux nouvelles especes de cheloniens dans le Miocene inferieur du bassin de l'Ebre (Bardenas Reales de Navarre). Comptes Rendus de l'Academie des Sciences, Series IIA, Earth and Planetary Science 328: 423 - 429. https: // doi. org / 10.1016 / S 1251 - 8050 (99) 80109 - 8" type="journal article" year="1999">Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999</bibRefCitation>
</taxonomicName>
, from the lower Miocene (MN2) of Monte Pila 2 (
<emphasis id="DFD9DC7EFFB46548FD63C497F0782FCE" bold="true" box="[749,780,1159,1176]" pageId="12" pageNumber="85">A -F</emphasis>
) and Monte Pila 3 (
<emphasis id="DFD9DC7EFFB46548FC24C497F0CC2FCE" bold="true" box="[938,952,1159,1176]" pageId="12" pageNumber="85">G</emphasis>
) (Lardero, La Rioja, Spain):
<emphasis id="DFD9DC7EFFB46548FB12C497F7DD2FCE" bold="true" box="[1180,1193,1159,1176]" pageId="12" pageNumber="85">A</emphasis>
,
<emphasis id="DFD9DC7EFFB46548FB3CC497F7CB2FCE" bold="true" box="[1202,1215,1159,1176]" pageId="12" pageNumber="85">B</emphasis>
, articulated epiplastra, MP2- 306, in ventral (
<emphasis id="DFD9DC7EFFB46548FE8FC48EF27A2FF9" bold="true" box="[257,270,1182,1199]" pageId="12" pageNumber="85">A</emphasis>
) and dorsal (
<emphasis id="DFD9DC7EFFB46548FEF9C48EF2F02FF9" bold="true" box="[375,388,1182,1199]" pageId="12" pageNumber="85">B</emphasis>
) views;
<emphasis id="DFD9DC7EFFB46548FE48C48EF2A02FF9" bold="true" box="[454,468,1182,1199]" pageId="12" pageNumber="85">C</emphasis>
, xiphiplastron, MP2-93, in ventral view;
<emphasis id="DFD9DC7EFFB46548FC92C48EF05E2FF9" bold="true" box="[796,810,1182,1199]" pageId="12" pageNumber="85">D</emphasis>
, hyoplastra, MP2-308,in ventral view;
<emphasis id="DFD9DC7EFFB46548FBE7C48EF7012FF9" bold="true" box="[1129,1141,1182,1199]" pageId="12" pageNumber="85">E</emphasis>
, hyoplastra, MP2-175,in ventral view;
<emphasis id="DFD9DC7EFFB46548FF0AC4A5F3FB2F90" bold="true" box="[132,143,1205,1222]" pageId="12" pageNumber="85">F</emphasis>
, nuchal, MP2-307, in dorsal view;
<emphasis id="DFD9DC7EFFB46548FE3CC4A5F2B42F90" bold="true" box="[434,448,1205,1222]" pageId="12" pageNumber="85">G</emphasis>
, hyoplastra, MP3-1, in ventral view. Abbreviations for the scutes (in upper case):
<emphasis id="DFD9DC7EFFB46548FBE5C4A5F7F12F90" bold="true" box="[1131,1157,1205,1222]" pageId="12" pageNumber="85">AB</emphasis>
, abdominal;
<emphasis id="DFD9DC7EFFB46548FB7CC4A5F6792F90" bold="true" box="[1266,1293,1205,1222]" pageId="12" pageNumber="85">AN</emphasis>
, anal;
<emphasis id="DFD9DC7EFFB46548FACBC4A5F62B2F90" bold="true" box="[1349,1375,1205,1222]" pageId="12" pageNumber="85">CE</emphasis>
, cervical;
<emphasis id="DFD9DC7EFFB46548FF0AC4DBF3EF2F8A" bold="true" box="[132,155,1227,1244]" pageId="12" pageNumber="85">FE</emphasis>
, femoral;
<emphasis id="DFD9DC7EFFB46548FF61C4DBF27F2F8A" bold="true" box="[239,267,1227,1244]" pageId="12" pageNumber="85">GU</emphasis>
, gular;
<emphasis id="DFD9DC7EFFB46548FEC7C4DBF2112F8A" bold="true" box="[329,357,1227,1244]" pageId="12" pageNumber="85">HU</emphasis>
, humeral;
<emphasis id="DFD9DC7EFFB46548FE33C4DBF2BA2F8A" bold="true" box="[445,462,1227,1244]" pageId="12" pageNumber="85">M</emphasis>
, marginal;
<emphasis id="DFD9DC7EFFB46548FDA4C4DBF1302F8A" bold="true" box="[554,580,1227,1244]" pageId="12" pageNumber="85">PC</emphasis>
, pectoral. Scale bar: 3 cm.
</paragraph>
</caption>
<paragraph id="ED12006CFFB46548FF15C503F0732DFB" blockId="12.[130,777,1299,2028]" pageId="12" pageNumber="85">
The abundance of specimens allows assessment of the variability in morphology and dimensions of several elements. For example, the relatively wide nuchal plates are almost as long as wide in some individuals, but slightly wider than long in others (
<figureCitation id="75961CE9FFB46548FF7BC583F2402EFB" box="[245,308,1427,1453]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
A-L). All of them show a thickening in the visceral region. The neural series anteriorly contacts the nuchal plate, and posteriorly the suprapygals. The neurals are wide in relation to their length (
<figureCitation id="75961CE9FFB46548FE0BC5E3F2B62D5B" box="[389,450,1523,1549]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
M-O; 10C, D). The morphology of these plates is variable. However, most of them are hexagonal, with the shortest margins being postero-laterally located. Although the anterior suprapygal is subquadrangular, the posterior is hexagonal, noticeably wider than long (
<figureCitation id="75961CE9FFB46548FF03C682F3BE2DFA" box="[141,202,1682,1708]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
Q-W). The pygal is also subquadrangular (
<figureCitation id="75961CE9FFB46548FD03C682F1BE2DFA" box="[653,714,1682,1708]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
X-Z).
</paragraph>
<paragraph id="ED12006CFFB46548FF15C6A2F7212E38" blockId="12.[130,777,1299,2028]" lastBlockId="12.[811,1458,1299,2028]" pageId="12" pageNumber="85">
This taxon has a single cervical scute (
<figureCitation id="75961CE9FFB46548FDBAC6A2F1FC2D9A" box="[564,648,1714,1740]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="12" pageNumber="85">Figs 3A</figureCitation>
;
<figureCitation id="75961CE9FFB46548FD1AC6A2F1D62D9A" box="[660,674,1714,1740]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">4</figureCitation>
A-L; 9F). Its lateral margins are subrounded. Its width/length ratio shows a wide range of variability, being slightly wider than long in some specimens, but almost twice as long as wide in others. The overlap of the cervical on the visceral surface of the nuchal is relatively long. The vertebral scutes are hexagonal, almost as wide as they are long (
<figureCitation id="75961CE9FFB46548FE58C761F1662CDD" box="[470,530,1905,1931]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="12" pageNumber="85">Fig. 3</figureCitation>
A-G). The first and last ones are slightly wider than the second to fourth vertebrals, which are all similar in width. However, the vertebral series is narrow, the first scute being narrower than the nuchal in many cases, not overlapping the postero-lateral margins of this plate. The fifth vertebral overlaps the anterior region of the pygal plate (
<figureCitation id="75961CE9FFB46548FC54C543F76C2E3B" box="[986,1048,1363,1389]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[134,1455,246,1846]" captionTargetId="figure-15@7.[132,1455,243,1849]" captionTargetPageId="7" captionText="FIG. 4. — Elements of the carapace of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-L, nuchals and adjacent plates, in dorsal view; A, MP1a-1570; B, MP1a-1342; C, MP1a-1364; D, MP1a-638; E, MP1a-528; F, MP1a-228; G, MP1a-475; H, MP1a-341; I, MP1a-679; J, MP1a-918; K, MP1b-95; L, MP1a-14; M-O, neurals and adjacent plates, in dorsal view; M, MP1b-556; N, MP1a-252; O, MP1a-707; P, two articulated costals, MP1a-48, in dorsal view; Q-W, suprapygals, in dorsal view; Q, MP1a-1305; R, MP1a-1221; S, MP1b-369; T, MP1a-1066; U, MP1a-1350; V, MP1a-511; W, MP1a-911; X-Z, pygals, in dorsal view; X, MP1a-757; Y, MP1a-6; Z, MP1a-879. Abbreviations for the plates (in lower case): c, costal; n, neural; nu, nuchal. Abbreviations for the scutes (in upper case): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548381" httpUri="https://zenodo.org/record/4548381/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
X-Z).
</paragraph>
<paragraph id="ED12006CFFB46548FCCDC563F7C92CDD" blockId="12.[811,1458,1299,2028]" pageId="12" pageNumber="85">
The contact between the hyoplastra and the hypoplastra, and that between the hypoplastra and the peripheral series, corresponds to a ligamentous junction, so that a hinge is developed between these plates, allowing mobility of the posterior plastral region (
<figureCitation id="75961CE9FFB46548FC0FC5E3F0A62D5B" box="[897,978,1523,1549]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="12" pageNumber="85">Figs 3F</figureCitation>
;
<figureCitation id="75961CE9FFB46548FC50C5E2F0982D5B" box="[990,1004,1522,1549]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[136,1452,218,1848]" captionTargetId="figure-19@8.[132,1455,228,1844]" captionTargetPageId="8" captionText="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548383" httpUri="https://zenodo.org/record/4548383/files/figure.png" pageId="12" pageNumber="85">5</figureCitation>
A-F; 7B-M; 8; 9D-E; 10A, B). The lateral margins of both plastral lobes are subrounded. The anterior lobe is very wide. Its anterior margin varies from subrounded to almost straight. Although the posterior lobe is clearly expanded in some specimens, it is much narrower in others. Thus, the development of the anal notch is variable in width and length, as is the shape of its margins (
<figureCitation id="75961CE9FFB46548FB60C6A2F64B2D9B" box="[1262,1343,1714,1741]" captionStart="FIG" captionStartId="5.[132,143,1894,1911]" captionTargetId="figure-19@6.[132,1455,224,2010]" captionTargetPageId="6" captionText="FIG. 3. — Partial shells and carapaces of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain):A-D, carapace and articulated anterior half of the plastron, MP1a-169, in dorsal (A), ventral (B), left lateral (C) and anterior (D) views; E-H, carapace and disjointed posterior half of the plastron, MP1a-643; E, G-H, carapace, in right lateral (E), dorsal (G) and ventral (H) views; F, posterior half of the plastron, in ventral view; I, partial carapace, MP1a-1070, in dorsal view; J, partial carapace, MP1b-415, in dorsal view. Abbreviations for the plates (in lower case and normal type): c, costal; n, neural; p, peripheral; py, pygal; spy, suprapygal. Abbreviations for the scutes (in upper case and in bold type): CE, cervical; M, marginal; PL, pleural; V, vertebral. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4548567" httpUri="https://zenodo.org/record/4548567/files/figure.png" pageId="12" pageNumber="85">Figs 3F</figureCitation>
;
<figureCitation id="75961CE9FFB46548FAC5C6A3F62D2D9B" box="[1355,1369,1715,1741]" captionStart="FIG" captionStartId="10.[132,143,1747,1764]" captionTargetBox="[135,1455,218,1710]" captionTargetId="figure-106@10.[132,1455,214,1715]" captionTargetPageId="10" captionText="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548387" httpUri="https://zenodo.org/record/4548387/files/figure.png" pageId="12" pageNumber="85">7</figureCitation>
N-U; 8; 10A, B). The epiplastral lip is concave (
<figureCitation id="75961CE9FFB46548FB5CC6C2F6602DBB" box="[1234,1300,1746,1773]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[136,1452,218,1848]" captionTargetId="figure-19@8.[132,1455,228,1844]" captionTargetPageId="8" captionText="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548383" httpUri="https://zenodo.org/record/4548383/files/figure.png" pageId="12" pageNumber="85">Figs 5</figureCitation>
;
<figureCitation id="75961CE9FFB46548FAACC6C2F6442DBA" box="[1314,1328,1746,1772]" captionStart="FIG" captionStartId="9.[132,143,1491,1508]" captionTargetBox="[136,1453,219,1454]" captionTargetId="figure-224@9.[132,1455,241,1471]" captionTargetPageId="9" captionText="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548385" httpUri="https://zenodo.org/record/4548385/files/figure.png" pageId="12" pageNumber="85">6</figureCitation>
A-V; 9A, B; 10A, B, G-L). Both the length and the thickness of this lip are observed to be highly variable. The dorsal thickening of this lip shows a pair of anterior processes, which exceed the margin of the anterior plastral lobe, at the level of the contacts between the gulars and the humerals.
</paragraph>
<paragraph id="ED12006CFFB4654BFCCDC781F0732A64" blockId="12.[811,1458,1299,2028]" lastBlockId="15.[132,775,215,306]" lastPageId="15" lastPageNumber="88" pageId="12" pageNumber="85">
The gular scutes overlap the anterior region of the entoplastron in some specimens, but not in others (
<figureCitation id="75961CE9FFB46548FB72C7A1F64D2C9D" box="[1276,1337,1969,1995]" captionStart="FIG" captionStartId="9.[132,143,1491,1508]" captionTargetBox="[136,1453,219,1454]" captionTargetId="figure-224@9.[132,1455,241,1471]" captionTargetPageId="9" captionText="FIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia,1999,from the lower Miocene (MN2) of Monte Pila 1 (Lardero,La Rioja,Spain):A-V, epiplastra,in dorsal and ventral views;A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b;Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548385" httpUri="https://zenodo.org/record/4548385/files/figure.png" pageId="12" pageNumber="85">Fig. 6</figureCitation>
W-AC). In the same way, the humero-pectoral sulci may be located next to the posterior margin of the entoplastron in some specimens, but overlap this plate in others, ranging from a very short overlap of the pectorals on the entoplastron in some individu- als, to a relatively long overlap, greater than a quarter of the length of this plate, in others. The abdominal scutes overlap the posterior region of the hyoplastra (
<figureCitation id="75961CE9FFB6654AFB32C7C1F78E2CBA" box="[1212,1274,2001,2028]" captionStart="FIG" captionStartId="8.[132,143,1917,1934]" captionTargetBox="[136,1452,218,1848]" captionTargetId="figure-19@8.[132,1455,228,1844]" captionTargetPageId="8" captionText="FIG. 5. — Articulated elements of the anterior plastral lobe of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A, B, MP1a-1242; C, D, MP1a-1422; E, F, MP1a-543; G, H, MP1a-277a; I, J, MP1a-1594; K, L, MP1a-1417; M, N, MP1a-1724. All of them are in dorsal and ventral views. Abbreviations for the plates (in lower case): en, entoplastron; ep, epiplastron. Abbreviations for the scutes (in upper case): AB, abdominal; GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548383" httpUri="https://zenodo.org/record/4548383/files/figure.png" pageId="14" pageNumber="87">Figs 5</figureCitation>
A-F; 7B-I; 9D-E; 10A). The anal scutes are restricted to the xiphiplastra, their anterior margin being well away from the suture between those plates and the hypoplastra (
<figureCitation id="75961CE9FFB7654BFE6DC107F1552A64" box="[483,545,279,306]" captionStart="FIG" captionStartId="10.[132,143,1747,1764]" captionTargetBox="[135,1455,218,1710]" captionTargetId="figure-106@10.[132,1455,214,1715]" captionTargetPageId="10" captionText="FIG. 7. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga,Lapparent de Broin,Pereda Suberbiola &amp; Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-I, hyoplastra, in ventral view; A, MP1a-694; B, MP1a- 385; C, MP1a-1333; D, MP1a-1025; E, MP1a-281; F, MP1a-1227; G, MP1b-585; H, MP1a-109; I, MP1b-169; J-M, hypoplastra, in ventral view; J, MP1b-467; K, MP1a-1194; L, MP1b-456; M, MP1a-367; N-U, xiphiplastra,in ventral view; N, MP1a-865; O, MP1a-56; P, MP1a-659; Q, MP1a-681; R, MP1a-474; S, MP1b- 380b; T, MP1a-81; U, MP1a-263.Abbreviations for the scutes (in upper case):AB, abdominal; AN, anal; FE, femoral; HU, humeral;PC, pectoral. Scale bar:3 cm." figureDoi="http://doi.org/10.5281/zenodo.4548387" httpUri="https://zenodo.org/record/4548387/files/figure.png" pageId="15" pageNumber="88">Figs 7</figureCitation>
N-U; 8; 9C; 10A, N).
</paragraph>
</subSubSection>
</treatment>
</document>