209 lines
19 KiB
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209 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.63.8198" ID-GBIF-Dataset="bf51ef6c-3446-4fd4-bb71-fdddcf1f773b" ID-PMC="PMC4956932" ID-Pensoft-Pub="1314-2003-63-107" ID-Pensoft-UUID="FF82FFF8E373BE6BFFF3FF84BF08EE20" ID-PubMed="27489482" ID-Zenodo-Dep="121162" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2003-63-107" ModsDocOrigin="PhytoKeys 63" ModsDocTitle="A revision of PoasubsectionAphanelytrum (Poaceae, Pooideae, Poaeae, Poinae); and a new species, Poaauriculata" checkinTime="1465304805122" checkinUser="pensoft" docAuthor="Peterson, Paul M. & Soreng, Robert J." docDate="2016" docId="210594040E36B124ADC6C71CA87EF204" docLanguage="en" docName="PhytoKeys 63: 107-125" docOrigin="PhytoKeys 63" docSource="http://dx.doi.org/10.3897/phytokeys.63.8198" docTitle="Poa auriculata Soreng & P. M. Peterson 2016, sp. nov." docType="treatment" docVersion="6" id="FF82FFF8E373BE6BFFF3FF84BF08EE20" lastPageNumber="117" masterDocId="FF82FFF8E373BE6BFFF3FF84BF08EE20" masterDocTitle="A revision of Poa subsection Aphanelytrum (Poaceae, Pooideae, Poaeae, Poinae); and a new species, Poa auriculata" masterLastPageNumber="125" masterPageNumber="107" pageNumber="116" updateTime="1668141686291" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Poa subsection Aphanelytrum (Poaceae, Pooideae, Poaeae, Poinae); and a new species, Poa auriculata</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>63</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>125</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.63.8198</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.63.8198</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-63-107</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF82FFF8E373BE6BFFF3FF84BF08EE20</mods:identifier>
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<mods:identifier type="Zenodo-Dep">121162</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161677319" LSID="urn:lsid:plazi:treatment:210594040E36B124ADC6C71CA87EF204" httpUri="http://treatment.plazi.org/id/210594040E36B124ADC6C71CA87EF204" lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116">
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<subSubSection pageId="9" pageNumber="116" type="nomenclature">
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<paragraph pageId="9" pageNumber="116">
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<taxonomicName LSID="AD1DD940-5F49-55E7-B37E-EBAFC013B9A4" authority="Soreng & P. M. Peterson" authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata" status="sp. nov.">
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<pageBreakToken pageId="9" pageNumber="116" start="start">Poa</pageBreakToken>
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auriculata Soreng & P.M. Peterson
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</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="116">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Poa auriculata: A Habit B Sheath, auricles, ligule, and blade C Spikelet D Glumes E Lower glume F Floret G Lemma H Palea, lateral view I Stamens with palea J Pistil enclosed in palea K Lodicules, pistil, and rachilla. (Wurdack 1145, US- 2382274)" figureDoi="10.3897/phytokeys.63.8198.figure5" httpUri="https://binary.pensoft.net/fig/91501" pageId="9" pageNumber="116">Fig. 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="materials_examined">
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<paragraph pageId="9" pageNumber="116">Type.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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<normalizedToken originalValue="PERÚ">PERU</normalizedToken>
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. Departamento Amazonas, Provincia Chachapoyas, summit of Puma-urcu southeast of Chachapoyas, occasional on dry cliff face, 3100-3200 m, 3 Jul 1962,
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<emphasis italics="true" pageId="9" pageNumber="116">J.J. Wurdack 1145</emphasis>
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(holotype: US-2382274!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="diagnosis">
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<paragraph pageId="9" pageNumber="116">Diagnosis.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName authorityName="Tovar" authorityYear="1965" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa scabrivaginata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="scabrivaginata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa scabrivaginata</emphasis>
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</taxonomicName>
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Tovar in having 4-6-flowered spikelets, a glabrous callus, glumes 2-3 mm long, and lemmas 2.5-3.8 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="description">
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<paragraph pageId="9" pageNumber="116">Description.</paragraph>
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<paragraph pageId="9" pageNumber="116">Caespitose, annuals or short-lived perennials. Culms 40-72 cm tall, erect, scabrous, shiny, often weak; nodes 3-5. Leaf sheaths 2/3 to 4/5 as long the internodes, membranous, greenish to stramineous, scabrous, upper sheaths open for 1/3 the length, keeled, summit with prominent triangular auricles; ligules 3.3-5 mm long, membranous to hyaline, apex erose, often split down the center; blades 6-15 cm long (flag leaf 3-6 cm long), 3-6 mm wide, flat, thin, lax, linear, scabrous. Panicles 5-11 cm long, 2.5-5 cm wide, ovate, open; branches flexuous, effuse and spreading with numerous spikelets, scabrous, the lower branches capillary. Spikelets 5-7 mm long, 4-6-flowered, glabrous, ovate, greenish-yellow tinged with purple; rachilla 0.4-1.0 mm long; glumes 2-3 mm long, membranous, subequal; lower glumes 2-2.5 mm long, 1-veined, linear lanceolate, apex acuminate; upper glumes 2.4-3 mm long, 3-veined, the veins not conspicuous, lanceolate, apex acute; lemmas 2.5-3.8 mm long, 5-veined, lanceolate, membranous; apex acute, unawned; paleas 2.3-3.7 mm long, membranous, 2-keeled, the keels scabrous, apex minutely bifid; lodicules 0.4-0.5 mm long, ovate, membranous, glabrous; stamens 3; anthers 1.9-2.1 mm long, yellowish; ovaries glabrous with two styles and two stigmas. Caryopses not seen.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="phenology">
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<paragraph pageId="9" pageNumber="116">Phenology.</paragraph>
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<paragraph pageId="9" pageNumber="116">Flowering in June and July.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="distribution">
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<paragraph pageId="9" pageNumber="116">Distribution.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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is known only from the type locality in Cordillera de los Andes of Peru near Chachapoyas between 3100-3200 m growing on a dry cliff face.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="conservation status">
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<paragraph pageId="9" pageNumber="116">Conservation status.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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The species is rare, but its conservation status is data deficient (
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<bibRefCitation author="IUCN" journalOrPublisher="US Department of Agriculture Miscellaneous Publication" pageId="11" pageNumber="118" refId="B13" refString="IUCN, 2010. IUCN Red List of Threatened Species. Version 2010.3." title="IUCN Red List of Threatened Species. Version 2010.3" year="2010">IUCN 2010</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="116" type="etymology">
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<paragraph pageId="9" pageNumber="116">Etymology.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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The specific epithet refers to the triangular auricles that are found on the summit of the sheaths, a feature that is unique among species of
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<taxonomicName authorityName="subsect. Tovarochloa (Macfarl. & P. But) Soreng & P. M. Peterson" authorityYear="2016" baseAuthorityName="subsect. Tovarochloa (Macfarl. & P. But) Soreng & P. M. Peterson" baseAuthorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116" type="comments">
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<paragraph pageId="9" pageNumber="116">Comments.</paragraph>
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<paragraph pageId="9" pageNumber="116">
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Initially RJS considered
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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to be related to species in
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<taxonomicName authority="subsect. Aphanelytrum" authorityName="subsect. Aphanelytrum (Hack.) Soreng & P. M. Peterson" authorityYear="2016" baseAuthorityName="subsect. Tovarochloa (Macfarl. & P. But) Soreng & P. M. Peterson" baseAuthorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" infraspecific-rank="subsect." kingdom="Plantae" lsidName="Poa" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="genus" subsection="Aphanelytrum">Poa subsect. Aphanelytrum</taxonomicName>
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. We do not place the new species in
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<taxonomicName authority="subsect. Aphanelytrum" authorityName="subsect. Aphanelytrum (Hack.) Soreng & P. M. Peterson" authorityYear="2016" baseAuthorityName="subsect. Tovarochloa (Macfarl. & P. But) Soreng & P. M. Peterson" baseAuthorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" infraspecific-rank="subsect." kingdom="Plantae" lsidName="Poa" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="genus" subsection="Aphanelytrum">Poa subsect. Aphanelytrum</taxonomicName>
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because the spikelets have short rachillas and are 4-6-flowered, the habit is annual to short-lived perennials with erect culms, and the lemmas are unawned without mucros. However, aside from the auricles,
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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is morphologically consistent with the 300 or so species that reside within
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<taxonomicName authority="subg. Poa supersect. Homalopoa" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" infraspecific-rank="supersect." kingdom="Plantae" lsidName="Poa" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="genus" supersection="Homalopoa">Poa subg. Poa supersect. Homalopoa</taxonomicName>
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. We hope to include a sample of this species in upcoming molecular analyses.
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</paragraph>
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<paragraph lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116">
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<taxonomicName authorityName="Tovar" authorityYear="1965" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa scabrivaginata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="scabrivaginata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa scabrivaginata</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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in having 2-flowered spikelets (4-6 in
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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), a few cobwebby hairs on the callus (verses glabrous), glumes 3.6-4.3 mm long (verses 2-3 mm), and lemmas 4.2-4.5 mm long (verses 2.5-3.8 mm) [
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<bibRefCitation author="Tovar, O" journalOrPublisher="Memorias del Museo de Historia Natural " Javier Prado "" pageId="11" pageNumber="118" pagination="1 - 65" refId="B36" refString="Tovar, O, 1965. Revision de las especies Peruanas del genero Poa. Memorias del Museo de Historia Natural "Javier Prado" 15: 1 - 65" title="Revision de las especies Peruanas del genero Poa." volume="15" year="1965">Tovar 1965</bibRefCitation>
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].
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<taxonomicName authorityName="Hack" authorityYear="1902" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa aequatoriensis" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="aequatoriensis">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa aequatoriensis</emphasis>
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</taxonomicName>
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Hack., another species possible to confuse with
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="9" pageNumber="116">Poa auriculata</emphasis>
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</taxonomicName>
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,
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<pageBreakToken pageId="10" pageNumber="117" start="start">differs</pageBreakToken>
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in having 2- or 3-, rarely 4-flowered spikelets, a few cobwebby hairs on the callus, and lemmas 3.6-4 mm long (
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<bibRefCitation author="Tovar, O" journalOrPublisher="Memorias del Museo de Historia Natural " Javier Prado "" pageId="11" pageNumber="118" pagination="1 - 65" refId="B36" refString="Tovar, O, 1965. Revision de las especies Peruanas del genero Poa. Memorias del Museo de Historia Natural "Javier Prado" 15: 1 - 65" title="Revision de las especies Peruanas del genero Poa." volume="15" year="1965">Tovar 1965</bibRefCitation>
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).
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<taxonomicName authorityName="Hack" authorityYear="1902" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa aequatoriensis" order="Poales" pageId="10" pageNumber="117" phylum="Tracheophyta" rank="species" species="aequatoriensis">
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<emphasis italics="true" pageId="10" pageNumber="117">Poa aequatoriensis</emphasis>
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</taxonomicName>
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is more wide ranging and has been reported in Colombia, Ecuador, and Peru whereas
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<taxonomicName authorityName="Tovar" authorityYear="1965" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa scabrivaginata" order="Poales" pageId="10" pageNumber="117" phylum="Tracheophyta" rank="species" species="scabrivaginata">
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<emphasis italics="true" pageId="10" pageNumber="117">Poa scabrivaginata</emphasis>
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</taxonomicName>
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is known from the type (Depto. de
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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, Tambo de Vaca,
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<emphasis italics="true" pageId="10" pageNumber="117">J.F. Macbride 4354</emphasis>
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at US) and one other possible collection (Depto. de Cajamarca, Celendin, I.
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<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
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Vega 2668 at MO) [
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<bibRefCitation author="Sylvester, SP" journalOrPublisher="Contributions from the United States National Herbarium" pageId="11" pageNumber="118" refId="B34" refString="Sylvester, SP, Soreng, RJ, Peterson, PM, Sylvester, MDPV, in press. An updated checklist and key to the open-panicled species of Poa L. (Poaceae) in Peru including three new species, Poaramoniana, Poatayacajaensis, and Poaurubambensis. PhytoKeys." title="An updated checklist and key to the open-panicled species of Poa L. (Poaceae) in Peru including three new species, Poaramoniana, Poatayacajaensis, and Poaurubambensis. PhytoKeys." year="in press">Sylvester et al. in press</bibRefCitation>
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].
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</paragraph>
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<caption ID-Zenodo-Dep="979691" doi="10.3897/phytokeys.63.8198.figure5" httpUri="https://binary.pensoft.net/fig/91501" pageId="10" pageNumber="117" start="Figure 5" startId="F5">
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<paragraph pageId="10" pageNumber="117">
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<emphasis bold="true" pageId="10" pageNumber="117">Figure 5.</emphasis>
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<taxonomicName authorityName="Soreng & P. M. Peterson" authorityYear="2016" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa auriculata" order="Poales" pageId="10" pageNumber="117" phylum="Tracheophyta" rank="species" species="auriculata">
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<emphasis italics="true" pageId="10" pageNumber="117">Poa auriculata</emphasis>
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</taxonomicName>
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:
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<emphasis bold="true" pageId="10" pageNumber="117">A</emphasis>
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Habit
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<emphasis bold="true" pageId="10" pageNumber="117">B</emphasis>
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Sheath, auricles, ligule, and blade
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<emphasis bold="true" pageId="10" pageNumber="117">C</emphasis>
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Spikelet
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<emphasis bold="true" pageId="10" pageNumber="117">D</emphasis>
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Glumes
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<emphasis bold="true" pageId="10" pageNumber="117">E</emphasis>
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Lower glume
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<emphasis bold="true" pageId="10" pageNumber="117">F</emphasis>
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Floret
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<emphasis bold="true" pageId="10" pageNumber="117">G</emphasis>
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Lemma
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<emphasis bold="true" pageId="10" pageNumber="117">H</emphasis>
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Palea, lateral view
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<emphasis bold="true" pageId="10" pageNumber="117">I</emphasis>
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Stamens with palea
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<emphasis bold="true" pageId="10" pageNumber="117">J</emphasis>
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Pistil enclosed in palea
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<emphasis bold="true" pageId="10" pageNumber="117">K</emphasis>
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Lodicules, pistil, and rachilla. (
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<emphasis italics="true" pageId="10" pageNumber="117">Wurdack 1145</emphasis>
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, US-2382274)
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |