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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.466.8047" ID-GBIF-Dataset="470bc8d8-7432-47cb-bae3-88617b4e91a9" ID-PMC="PMC4296484" ID-Pensoft-Pub="1313-2970-466-77" ID-PubMed="25610337" ID-ZBK="19C20324B7A548EBBBA45D849697EA23" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-466-77" ModsDocOrigin="ZooKeys 466" ModsDocTitle="Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China" checkinTime="1451244919976" checkinUser="pensoft" docAuthor="Liu, Sulin, Shih, Chungkun &amp; Ren, Dong" docDate="2014" docId="C061F1F83EB638BD426A98B54F6C4072" docLanguage="en" docName="ZooKeys 466: 77-94" docOrigin="ZooKeys 466" docSource="http://dx.doi.org/10.3897/zookeys.466.8047" docTitle="Exilibittacus plagioneurus Liu, Shih &amp; Ren, 2014, sp. n." docType="treatment" docUuid="CCEF6E6E-B54D-43CB-8E24-F289597FB4C1" docUuidSource="ZooBank" docVersion="4" lastPageNumber="86" masterDocId="FF84FF8DFFDAFFD6F05B8E6A350B1F27" masterDocTitle="Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China" masterLastPageNumber="94" masterPageNumber="77" pageNumber="84" updateTime="1668159766707" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Sulin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shih, Chungkun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Dong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>466</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.466.8047</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.466.8047</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-466-77</mods:identifier>
<mods:identifier type="ZBK">19C20324B7A548EBBBA45D849697EA23</mods:identifier>
<mods:identifier type="ZooBank">19C20324B7A548EBBBA45D849697EA23</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152057236" LSID="urn:lsid:zoobank.org:act:CCEF6E6E-B54D-43CB-8E24-F289597FB4C1" httpUri="http://treatment.plazi.org/id/C061F1F83EB638BD426A98B54F6C4072" lastPageId="9" lastPageNumber="86" pageId="7" pageNumber="84">
<subSubSection pageId="7" pageNumber="84" type="multiple">
<paragraph pageId="7" pageNumber="84">Taxon classification Animalia Mecoptera Bittacidae</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="84" type="nomenclature">
<paragraph pageId="7" pageNumber="84">
<taxonomicName LSID="http://zoobank.org/CCEF6E6E-B54D-43CB-8E24-F289597FB4C1" class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus plagioneurus" order="Mecoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="plagioneurus">Exilibittacus plagioneurus</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="84">sp. n.</taxonomicNameLabel>
Figs 8, 9
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="etymology">
<paragraph pageId="8" pageNumber="85">
<pageBreakToken pageId="8" pageNumber="85" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="85">
From Greek
<normalizedToken originalValue="“plagios”">&quot;plagios&quot;</normalizedToken>
(oblique) and
<normalizedToken originalValue="“neuron”">&quot;neuron&quot;</normalizedToken>
(vein), referring to oblique crossveins of the wings.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="holotype">
<paragraph pageId="8" pageNumber="85">Holotype.</paragraph>
<paragraph pageId="8" pageNumber="85">Female, CNU-MEC-NN2013013 P/C, in dorsal view. Abdomen length 8.3 mm, forewing length 9.3 mm with a maximal width of 2.3 mm; hind wing length 8.4 mm with a maximal width of 2.2 mm.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="horizon and locality">
<paragraph pageId="8" pageNumber="85">Horizon and locality.</paragraph>
<paragraph pageId="8" pageNumber="85">Jiulongshan Formation, late Middle Jurassic, Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="diagnosis">
<paragraph pageId="8" pageNumber="85">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="85">Forewing Sc terminating at the anterior margin at the same level of the R4+5 forking; Vein 1A terminating at the posterior margin distad of the origination of Rs from R1.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="86" pageId="8" pageNumber="85" type="description">
<paragraph pageId="8" pageNumber="85">Description.</paragraph>
<paragraph pageId="8" pageNumber="85">
Female, small-sized, head not preserved but mesothorax and metathorax preserved. Legs partially preserved, one hind leg with five tarsomeres present but the pretarsal claw not preserved, the fifth tarsomere folded against the fourth; the first and second tarsomeres with several spines. (Fig. 8
<normalizedToken originalValue="AD">A-D</normalizedToken>
)
</paragraph>
<caption pageId="8" pageNumber="85">
<paragraph pageId="8" pageNumber="85">
Figure 8.
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus plagioneurus" order="Mecoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="plagioneurus">Exilibittacus plagioneurus</taxonomicName>
sp. n., holotype, CNU-MEC-NN2013013 P/C, dorsal view. A photo of part B photo of counterpart C line drawing of part D photo of a hind leg, under alcohol. Scale bars represent 3 mm in
<normalizedToken originalValue="AC">A-C</normalizedToken>
, 1 mm in D.
</paragraph>
</caption>
<paragraph lastPageId="9" lastPageNumber="86" pageId="8" pageNumber="85">
Forewing. Wing narrow basally with obviously dark pterostigma. Sc long, reaching the anterior margin at the same level of the R4+5 forking; R1 not forking, one subcostal crossvein (Scv) between Sc and R1, Scv about 1/6 as long as the Sc length between Scv and the apex of Sc; one pterostigmal crossvein (Pcv) and one crossvein between R1 and R2+3; Rs with four branches, one crossvein between R3 and R4, one between R2+3 and R4 and one between R4 and R5; M with four branches, M4 base bending sharply; the 'bittacid
<normalizedToken originalValue="cross">cross'</normalizedToken>
aligned, the posterior part of the 'bittacid
<normalizedToken originalValue="cross">cross'</normalizedToken>
reaching M3 distad of the M3+4 forking point; two crossveins between R5 and M1, one between M1 and M2 and one between M2 and M3; Cu1 ending before the forking of R4+5, one crossvein between M4 and Cu1, one between Cu1 and Cu2; one short
<pageBreakToken pageId="9" pageNumber="86" start="start">crossvein</pageBreakToken>
between Cu2 and 1A; 1A terminating at the posterior margin distad of the origination of Rs; 2A ending proximad of the originations of Rs and M, one crossvein between 1A and 2A (Fig. 9A, C).
</paragraph>
<caption pageId="9" pageNumber="86">
<paragraph pageId="9" pageNumber="86">
Figure 9.
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus plagioneurus" order="Mecoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="plagioneurus">Exilibittacus plagioneurus</taxonomicName>
sp. n., holotype. Line drawings of part. A right forewing of part B right hind wing of part C left forewing of part D left hind wing of part. Scale bars represent 1 mm in
<normalizedToken originalValue="AD">A-D</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="86">
Hind wing. Sc short, reaching the anterior margin before the forking of R4+5, one crossvein (Scv) between Sc and R1; R1 smooth and not sagging through the pterostigmal area; one pterostigmal crossvein (Pcv) present. Rs with three branches; one crossvein between R1 and R2, one between R2 and R3 and one between R3 and R4; M with three branches; two crossveins between R4 and M1, one between M1 and M2 and one between M2 and M3; the 'bittacid
<normalizedToken originalValue="cross">cross'</normalizedToken>
not aligned; one between M3 and Cu1 and one between Cu1 and Cu2. Vein 1A terminating at the posterior margin distad of the origination of Rs, one crossvein between Cu2 and 1A (Fig. 9B, D).
</paragraph>
<paragraph pageId="9" pageNumber="86">
Abdomen. Ten segments visible, genital segments not preserved (Fig. 8
<normalizedToken originalValue="AC">A-C</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="86" type="remarks">
<paragraph pageId="9" pageNumber="86">Remarks.</paragraph>
<paragraph pageId="9" pageNumber="86">
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus plagioneurus" order="Mecoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="plagioneurus">Exilibittacus plagioneurus</taxonomicName>
sp. n. (Figs 8, 9) is assigned to
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus" order="Mecoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="genus">Exilibittacus</taxonomicName>
Yang, Ren &amp; Shih, 2012 based on the following generic diagnostic characters: in forewing, Sc reaching the anterior margin at the same level of the forking of R4+5 and the 'bittacid
<normalizedToken originalValue="cross">cross'</normalizedToken>
aligned, and in hind wing, M with three branches. The new species is differentiated from
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus lii" order="Mecoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="lii">Exilibittacus lii</taxonomicName>
and
<taxonomicName class="Insecta" family="Bittacidae" genus="Exilibittacus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Exilibittacus foliaceus" order="Mecoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="foliaceus">Exilibittacus foliaceus</taxonomicName>
sp. n. by characters shown in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>