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<document id="72514C59802645DCED5B348B090474AF" ID-CLB-Dataset="53501" ID-DOI="10.5852/ejt.2023.870.2123" ID-GBIF-Dataset="bac760e2-c211-419a-a1ca-5f5af426268f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7986633" ID-ZooBank="508560B7-BD89-472A-B3FD-F48851B024C3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1685458305949" checkinUser="felipe" docAuthor="Nagy, Hajnalka, Dózsa-Farkas, Klára &amp; Felföldi, Tamás" docDate="2023" docId="B857879AFFD9FFBEFDD35844FB44FDC7" docLanguage="en" docName="ejt-870_107-145.pdf" docOrigin="European Journal of Taxonomy 870" docSource="http://dx.doi.org/10.5852/ejt.2023.870.2123" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enchytraeus adrianensis Nagy &amp; Dózsa-Farkas &amp; Felföldi 2023, sp. nov." docType="treatment" docUuid="B49B8279-3D90-408F-B45C-BF89ACFC3C2B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="13" masterDocId="446EFFE2FFDFFFB2FF825D44FFFEFFDC" masterDocTitle="New insights into the Enchytraeus albidus complex (Annelida, Enchytraeidae), with the description of three new species from seashores in Italy and Croatia" masterLastPageNumber="145" masterPageNumber="107" pageNumber="7" updateTime="1699511372322" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4A684B4FA70FC321471F8CEADADEE180">New insights into the Enchytraeus albidus complex (Annelida, Enchytraeidae), with the description of three new species from seashores in Italy and Croatia</mods:title>
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<mods:namePart id="4243AD278E7AD4952DE26C8C845DA65D">Nagy, Hajnalka</mods:namePart>
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<mods:affiliation id="DB5810BF5708B6FB0E2EBA63283DF3A1">Department of Microbiology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Hungarian Natural History Museum, Baross utca 13, H- 1088 Budapest, Hungary. Department of Systematic Zoology and Ecology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Institute of Aquatic Ecology, Centre for Ecological Research, Karolina út 29, H- 1113 Budapest, Hungary.</mods:affiliation>
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<mods:namePart id="2B13767902E7AC5F37389BD650435568">Dózsa-Farkas, Klára</mods:namePart>
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<mods:namePart id="5544AA494B67F14F67D886B22BD91766">Felföldi, Tamás</mods:namePart>
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<mods:affiliation id="8DA68FB1F4A975248ABBB734ECA0E87C">Department of Microbiology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Hungarian Natural History Museum, Baross utca 13, H- 1088 Budapest, Hungary. Department of Systematic Zoology and Ecology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Institute of Aquatic Ecology, Centre for Ecological Research, Karolina út 29, H- 1113 Budapest, Hungary.</mods:affiliation>
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<treatment id="B857879AFFD9FFBEFDD35844FB44FDC7" ID-DOI="http://doi.org/10.5281/zenodo.7987789" ID-GBIF-Taxon="209663766" ID-Zenodo-Dep="7987789" LSID="urn:lsid:zoobank.org:act:B49B8279-3D90-408F-B45C-BF89ACFC3C2B" httpUri="http://treatment.plazi.org/id/B857879AFFD9FFBEFDD35844FB44FDC7" lastPageId="12" lastPageNumber="13" pageId="6" pageNumber="7">
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<taxonomicName id="F7FE4D0FFFD9FFB4FDD35844FC7FFAC7" ID-CoL="f3ef1a07-d0f8-4411-a92b-bdc65316bab4" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[593,897,1280,1307]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
<emphasis id="028AEA9EFFD9FFB4FDD35844FC7FFAC7" bold="true" box="[593,897,1280,1307]" italics="true" pageId="6" pageNumber="7">Enchytraeus adrianensis</emphasis>
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<subSubSection id="78E46507FFD9FFB4FD055803FC53FABD" box="[647,941,1351,1377]" pageId="6" pageNumber="7" type="description">
<paragraph id="3041368CFFD9FFB4FD055803FC53FABD" blockId="6.[362,1225,1280,1377]" box="[647,941,1351,1377]" pageId="6" pageNumber="7">
<figureCitation id="A8C52A09FFD9FFB4FD055803FD19FABD" box="[647,743,1351,1377]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[346,1228,954,1877]" captionTargetId="figure-231@7.[313,1275,942,1892]" captionTargetPageId="7" captionText="Fig. 1. Schematic drawing of spermathecaeof the specimens studied.A. Enchytraeus adrianensissp. nov. B. E. krumbachi (Čejka, 1913). C–D. E. andrasi sp. nov. E. E. andrasiformis sp. nov. F. E. albidus Henle, 1837. G. E. irregularis Nielsen &amp; Christensen, 1961." figureDoi="http://doi.org/10.5281/zenodo.7986635" httpUri="https://zenodo.org/record/7986635/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
, 
<figureCitation id="A8C52A09FFD9FFB4FD715803FCDCFABD" box="[755,802,1351,1377]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,232,1681,1707]" captionStartId-1="11.[189,232,1787,1813]" captionTargetBox-0="[189,1398,461,1643]" captionTargetBox-1="[207,1365,612,1734]" captionTargetId-0="figure-48@9.[189,1398,461,1646]" captionTargetId-1="figure-129@11.[207,1380,612,1747]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." captionText-1="Fig. 3. Micrograph of Enchytraeus adrianensis sp. nov. A, C, G–H. In vivo. B, D–F, I. Fixed, stained. A. Sperm funnels. B. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). C–F. Multiple male glands (E–F primary bulb marked with arrow). D. Paratype (slide 2748, P.143.3, ELTE). E. Paratype (slide 2748, P.143.3, ELTE). F. Holotype (En.2. slide 2842, ELTE). I. Paratype (slide 2827, P. 143.8, ELTE). F. Holotype (En.2. slide 2842, ELTE). G–I. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.7986637" figureDoi-1="http://doi.org/10.5281/zenodo.7986639" httpUri-0="https://zenodo.org/record/7986637/files/figure.png" httpUri-1="https://zenodo.org/record/7986639/files/figure.png" pageId="6" pageNumber="7">2–3</figureCitation>
, 
<tableCitation id="7D7C0337FFD9FFB4FCAF5803FC53FABD" box="[813,941,1351,1377]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="3.[137,207,517,545]" captionStartId-1="13.[137,207,545,573]" captionTargetPageId-0="3" captionTargetPageId-1="13" captionText-0="Table 1 (continued on next page). List of specimens used for molecular taxonomic analyses with collection data and GenBank accession numbers. Sequences determined in this study appear in bold. Holotypes and paratypes of new species are indicated with H and P in parentheses.Abbreviation: n. d. = no data." captionText-1="Table 2 (continued on next two pages). Comparison of morphological characters (measurements in fixed specimens). The data are derived from this study and from Erséus et al. (2019), Arslan et al. (2018) and according to the author of original species description (marked with*). The data given by us marked with bold, dimensions in μm." httpUri-0="http://table.plazi.org/id/64816604FFDCFFB1FF0B5F41FEC7FDBB" httpUri-1="http://table.plazi.org/id/64816604FFD2FFBFFF0B5F65FD9FFD5F" pageId="6" pageNumber="7" tableUuid-0="64816604FFDCFFB1FF0B5F41FEC7FDBB" tableUuid-1="64816604FFD2FFBFFF0B5F65FD9FFD5F">Tables 1–2</tableCitation>
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<heading id="6B0981E0FFD9FFB4FF3F58CCFEC9FA7E" bold="true" box="[189,311,1416,1442]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="028AEA9EFFD9FFB4FF3F58CCFEC9FA7E" bold="true" box="[189,311,1416,1442]" pageId="6" pageNumber="7">Diagnosis</emphasis>
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(1) Body length 
<quantity id="F7069B69FFD9FFB4FE0858F3FDF2FA0D" box="[394,524,1463,1489]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="3.0" metricValueMin="1.4" pageId="6" pageNumber="7" unit="mm" value="22.0" valueMax="30.0" valueMin="14.0">14–30 mm</quantity>
(in vivo), segment number 51–84; (2) chaetae maximum 4 per bundle, straight with ental hook; (3) clitellum in XII–XIII, hyalocytes and granulocytes in dense transverse rows dorsally and laterally (or in a reticulate pattern), mostly only granulocytes between the male pores; (4) epidermal gland cells in 3–4 rows/segments, often brownish; (5) four pairs of nephridia preclitellarly; (6) pharyngeal glands widely or slightly connected dorsally and with ventral lobes; (7) dorsal blood vessel origin from XIV–XV, blood colourless; (8) sperm funnel cylindrical, 350–690 μm long, 2–4 × as long as wide in vivo; (9) vasa deferentia tripartite, extending into XV–XIX, the slender part ciliated, the thickened middle section not ciliated; (10) male glands multiple: one large rounded primary bulb (60– 125 μm long and 50–95 μm wide), and 6–9 smaller secondary glands, ventral recess between the male copulatory organs absent; (11) spermathecal ectal duct (90–160 μm long in vivo) covered with gland cells, ampulla oval or spherical (70–115 μm wide) with wide wall and without diverticula, connecting with oesophagus; (12) 2–7 mature eggs present at a time.
</paragraph>
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<subSubSection id="78E46507FFD9FFB4FF3F5A38FCFFF818" box="[189,769,1916,1989]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="3041368CFFD9FFB4FF3F5A38FEBBF84A" blockId="6.[189,325,1916,1942]" box="[189,325,1916,1942]" pageId="6" pageNumber="7">
<heading id="6B0981E0FFD9FFB4FF3F5A38FEBBF84A" bold="true" box="[189,325,1916,1942]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="028AEA9EFFD9FFB4FF3F5A38FEBBF84A" bold="true" box="[189,325,1916,1942]" pageId="6" pageNumber="7">Etymology</emphasis>
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<paragraph id="3041368CFFD9FFB4FF3F5AEEFCFFF818" blockId="6.[189,769,1962,1989]" box="[189,769,1962,1989]" pageId="6" pageNumber="7">Named after the Adriatic Sea where it was found.</paragraph>
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<paragraph id="3041368CFFD8FFB5FF3F5C4CFE53FEFE" blockId="7.[189,429,264,290]" box="[189,429,264,290]" pageId="7" pageNumber="8">
<heading id="6B0981E0FFD8FFB5FF3F5C4CFE53FEFE" bold="true" box="[189,429,264,290]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="028AEA9EFFD8FFB5FF3F5C4CFE53FEFE" bold="true" box="[189,429,264,290]" pageId="7" pageNumber="8">Material examined</emphasis>
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<paragraph id="3041368CFFD8FFB5FF685C73FEA3FE8D" blockId="7.[189,1399,311,446]" box="[234,349,311,337]" pageId="7" pageNumber="8">
<emphasis id="028AEA9EFFD8FFB5FF685C73FEA3FE8D" bold="true" box="[234,349,311,337]" pageId="7" pageNumber="8">
<typeStatus id="EF45882EFFD8FFB5FF685C73FEA3FE8D" box="[234,349,311,337]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
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<paragraph id="3041368CFFD8FFB5FF3F5C1AFEF4FE62" blockId="7.[189,1399,311,446]" pageId="7" pageNumber="8">
<materialsCitation id="80963CD1FFD8FFB5FF3F5C1AFEF4FE62" ID-GBIF-Occurrence="4120752309" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok" country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Adriatic Sea, Kale Cove, seashore, decaying Zostera debris" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea, Kale Cove, seashore, decaying Zostera debris" pageId="7" pageNumber="8" specimenCode="slide 2842" specimenCount="1" stateProvince="Istria" typeStatus="holotype">
<collectingCountry id="48E9761CFFD8FFB5FF3F5C1AFEC4FEA4" box="[189,314,350,376]" name="Croatia" pageId="7" pageNumber="8">CROATIA</collectingCountry>
• 
<collectingRegion id="F23AF86EFFD8FFB5FED45C19FE6BFEAB" box="[342,405,349,375]" country="Croatia" name="Istarska" pageId="7" pageNumber="8">Istria</collectingRegion>
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<collectingMunicipality id="D025ACF6FFD8FFB5FD215C19FA8CFEAB" box="[675,1394,347,376]" country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" name="Adriatic Sea, Kale Cove, seashore, decaying Zostera debris" pageId="7" pageNumber="8" stateProvince="Istria">
Adriatic Sea, Kale Cove, seashore, decaying 
<taxonomicName id="F7FE4D0FFFD8FFB5FB475C1AFADEFEAB" box="[1221,1312,350,375]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="028AEA9EFFD8FFB5FB475C1AFADEFEAB" box="[1221,1312,350,375]" italics="true" pageId="7" pageNumber="8">Zostera</emphasis>
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debris
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<geoCoordinate id="55CA504BFFD8FFB5FEF05C3BFDE4FE47" box="[370,538,383,411]" degrees="13" direction="east" minutes="58" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="50.5" value="13.980694">13°58′50.5″ E</geoCoordinate>
; 
<date id="4440104CFFD8FFB5FDA45CC5FD4EFE47" box="[550,688,385,411]" pageId="7" pageNumber="8" value="2019-04-03">
<collectingDate id="5404E9A4FFD8FFB5FDA45CC5FD4EFE47" box="[550,688,385,411]" pageId="7" pageNumber="8" value="2019-04-03">3 Apr. 2019</collectingDate>
</date>
; 
<collectorName id="9D0B535AFFD8FFB5FD3E5CC5FCBFFE47" box="[700,833,385,411]" pageId="7" pageNumber="8">Júlia Török</collectorName>
leg., from culture: 
<date id="4440104CFFD8FFB5FB9D5CC4FB58FE47" box="[1055,1190,384,411]" pageId="7" pageNumber="8" value="2019-10-15">
<collectingDate id="5404E9A4FFD8FFB5FB9D5CC4FB58FE47" box="[1055,1190,384,411]" pageId="7" pageNumber="8" value="2019-10-15">15.10.2019</collectingDate>
</date>
, En.2. 
<specimenCode id="60589EF7FFD8FFB5FB775CC5FA8CFE47" box="[1269,1394,385,411]" pageId="7" pageNumber="8">slide 2842</specimenCode>
; 
<collectionCode id="56EFAE49FFD8FFB5FF3F5CE0FEF8FE62" box="[189,262,420,446]" pageId="7" pageNumber="8">ELTE</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="3041368CFFD8FFB5FF685CA1FE17FE23" blockId="7.[188,1401,485,864]" box="[234,489,485,511]" pageId="7" pageNumber="8">
<emphasis id="028AEA9EFFD8FFB5FF685CA1FE97FE23" bold="true" box="[234,361,485,511]" pageId="7" pageNumber="8">
<typeStatus id="EF45882EFFD8FFB5FF685CA1FE97FE23" box="[234,361,485,511]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
(21 specs)
</paragraph>
<paragraph id="3041368CFFD8FFBAFF3F5F4DFE7FFDBC" blockId="7.[188,1401,485,864]" lastBlockId="8.[189,1401,265,608]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<materialsCitation id="80963CD1FFD8FFB5FF3F5F4DFB2FFDFF" ID-GBIF-Occurrence="4120752333" box="[189,1233,520,547]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="CROATIA" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2841a,b" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<collectingCountry id="48E9761CFFD8FFB5FF3F5F4DFEC4FDFF" box="[189,314,521,547]" name="Croatia" pageId="7" pageNumber="8">CROATIA</collectingCountry>
• 
<specimenCount id="26F8FD05FFD8FFB5FED45F4DFE55FDFF" box="[342,427,521,547]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FCF45F4CFBE6FDFE" box="[886,1048,520,546]" pageId="7" pageNumber="8">slide 2841a,b</specimenCode>
; P.143.1; 
<collectionCode id="56EFAE49FFD8FFB5FB0E5F4DFB2FFDFF" box="[1164,1233,521,547]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FB6F5F4DFE0DFDB5" ID-GBIF-Occurrence="4120752312" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2717" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FB6F5F4DFABCFDFF" box="[1261,1346,521,547]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
, last 19 segments, 
<quantity id="F7069B69FFD8FFB5FED95F68FE60FD9A" box="[347,414,556,582]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="mm" value="3.0">3 mm</quantity>
used for DNA analysis (No. 1265, ID number); same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FF3E5F0BFEC5FDB5" box="[188,315,591,617]" pageId="7" pageNumber="8">slide 2717</specimenCode>
; P.143.2; 
<collectionCode id="56EFAE49FFD8FFB5FE2C5F0BFE0DFDB5" box="[430,499,591,617]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FD925F0BFA9BFDB5" ID-GBIF-Occurrence="4120752304" box="[528,1381,591,617]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2748" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FD925F0BFD98FDB5" box="[528,614,591,617]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FBAC5F0BFB52FDB5" box="[1070,1196,591,617]" pageId="7" pageNumber="8">slide 2748</specimenCode>
; P.143.3; 
<collectionCode id="56EFAE49FFD8FFB5FAA25F0BFA9BFDB5" box="[1312,1381,591,617]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FF3C5F36FD41FD6C" ID-GBIF-Occurrence="4120752334" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2750" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FF3C5F36FEEDFD51" box="[190,275,626,653]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
, last 18 segments, 
<quantity id="F7069B69FFD8FFB5FE725F36FDB5FD51" box="[496,587,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="8" unit="mm" value="2.3">2.3 mm</quantity>
used for DNA analysis (No. 1324, ID number); same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FE015FD1FDFDFD6C" box="[387,515,661,688]" pageId="7" pageNumber="8">slide 2750</specimenCode>
; P.143.4; 
<collectionCode id="56EFAE49FFD8FFB5FDF85FD2FD41FD6C" box="[634,703,662,688]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FD5C5FD1FB3CFD0F" ID-GBIF-Occurrence="4120752307" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2751" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FD5C5FD1FCCAFD6C" box="[734,820,661,688]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
, last 15 segments, 
<quantity id="F7069B69FFD8FFB5FB9B5FD1FBA1FD6C" box="[1049,1119,661,688]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="mm" value="3.0">3 mm</quantity>
used for DNA analysis (No. 1266, ID number); same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FC0F5FFCFBF5FD0F" box="[909,1035,696,723]" pageId="7" pageNumber="8">slide 2751</specimenCode>
; P.143.5; 
<collectionCode id="56EFAE49FFD8FFB5FBFF5FFDFB3CFD0F" box="[1149,1218,697,723]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FB585FFDFCE2FD2A" ID-GBIF-Occurrence="4120752317" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="2760" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FB585FFDFAD0FD0F" box="[1242,1326,697,723]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FDAB5F98FD99FD2A" box="[553,615,732,758]" pageId="7" pageNumber="8">2760</specimenCode>
; P.143.6; 
<collectionCode id="56EFAE49FFD8FFB5FD555F98FCE2FD2A" box="[727,796,732,758]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FCB35F98FE24FCC5" ID-GBIF-Occurrence="4120752322" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2822a-c" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FCB35F98FC7BFD2A" box="[817,901,732,758]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FABC5F98FEDCFCC5" pageId="7" pageNumber="8">slide 2822a–c</specimenCode>
; P.143.7; 
<collectionCode id="56EFAE49FFD8FFB5FE175FBBFE24FCC5" box="[405,474,767,793]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FE765FBBFA9BFCC5" ID-GBIF-Occurrence="4120752303" box="[500,1381,767,793]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2827a,b" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FE765FBBFDB7FCC5" box="[500,585,767,793]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FB895FBBFB53FCC5" box="[1035,1197,767,793]" pageId="7" pageNumber="8">slide 2827a,b</specimenCode>
; P.143.8; 
<collectionCode id="56EFAE49FFD8FFB5FAA25FBBFA9BFCC5" box="[1312,1381,767,793]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FF3F5E66FBF2FCE1" ID-GBIF-Occurrence="4120752329" box="[189,1036,802,829]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2850" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FF3F5E66FEECFCE1" box="[189,274,802,829]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FD555E66FCAAFCE1" box="[727,852,802,829]" pageId="7" pageNumber="8">slide 2850</specimenCode>
; P.143.9; 
<collectionCode id="56EFAE49FFD8FFB5FC455E67FBF2FCE1" box="[967,1036,803,829]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFB5FBA75E66FD0DFCBC" ID-GBIF-Occurrence="4120752330" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2851a,b" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FBA75E66FB84FCE1" box="[1061,1146,802,829]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFB5FE075E01FDD6FCBC" box="[389,552,837,864]" pageId="7" pageNumber="8">slide 2851a,b</specimenCode>
; P.143.10; 
<collectionCode id="56EFAE49FFD8FFB5FD2C5E02FD0DFCBC" box="[686,755,838,864]" pageId="7" pageNumber="8">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD8FFBAFC935E01FE1BFEFF" ID-GBIF-Occurrence="4120752337" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" lastPageId="8" lastPageNumber="9" latitude="44.85361" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="7" pageNumber="8" specimenCode="slide 2852a,b" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD8FFB5FC935E01FC97FCBC" box="[785,873,837,864]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD8FFBAFABD5E01FEE0FEF8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">slide 2852a,b</specimenCode>
; P.143.11; 
<collectionCode id="56EFAE49FFD7FFBAFE225C4DFE1BFEFF" box="[416,485,265,291]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFD835C4DFA9BFEFF" ID-GBIF-Occurrence="4120752331" box="[513,1381,265,292]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2853" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFD835C4DFDA8FEFF" box="[513,598,265,291]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFB9D5C4DFB63FEF8" box="[1055,1181,265,292]" pageId="8" pageNumber="9">slide 2853</specimenCode>
; P.143.12; 
<collectionCode id="56EFAE49FFD7FFBAFAA25C4DFA9BFEFF" box="[1312,1381,265,291]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFF3C5C68FD3FFEB6" ID-GBIF-Occurrence="4120752302" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2854" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFF3C5C68FEF1FE9B" box="[190,271,300,327]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
[last 16 segments, 
<quantity id="F7069B69FFD7FFBAFE6D5C68FDB6FE9B" box="[495,584,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="9" unit="mm" value="2.2">2.2 mm</quantity>
used for DNA analysis (
<accessionNumber id="2FADAB6FFFD7FFBAFCEA5C68FC27FE9A" box="[872,985,300,326]" pageId="8" pageNumber="9">No. 1379</accessionNumber>
, ID number)]; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFEFF5C0BFE04FEB6" box="[381,506,335,362]" pageId="8" pageNumber="9">slide 2854</specimenCode>
; P.143.13; 
<collectionCode id="56EFAE49FFD7FFBAFDFE5C14FD3FFEB6" box="[636,705,336,362]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFD595C0BFE84FE51" ID-GBIF-Occurrence="4120752326" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2855" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFD595C0BFCCEFEB6" box="[731,816,335,362]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFB775C0BFA8CFEB6" box="[1269,1394,335,362]" pageId="8" pageNumber="9">slide 2855</specimenCode>
; P.143.14; 
<collectionCode id="56EFAE49FFD7FFBAFEB75C37FE84FE51" box="[309,378,371,397]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFE155C37FC2BFE6C" ID-GBIF-Occurrence="4120752301" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2856" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFE155C37FE10FE51" box="[407,494,371,397]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
, last 22 segments, 
<quantity id="F7069B69FFD7FFBAFD575C37FCCCFE51" box="[725,818,371,397]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="9" unit="mm" value="2.7">2.7 mm</quantity>
used for DNA analysis (
<accessionNumber id="2FADAB6FFFD7FFBAFBE25C37FB2DFE51" box="[1120,1235,371,397]" pageId="8" pageNumber="9">No. 1380</accessionNumber>
, ID number); same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFD0A5CD2FCF6FE6D" box="[648,776,406,433]" pageId="8" pageNumber="9">slide 2856</specimenCode>
; P.143.15; 
<collectionCode id="56EFAE49FFD7FFBAFC125CD2FC2BFE6C" box="[912,981,406,432]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFC775CD2FD55FE0F" ID-GBIF-Occurrence="4120752306" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2857" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFC775CD2FBB3FE6C" box="[1013,1101,406,432]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFEE35CFDFE1EFE08" box="[353,480,441,468]" pageId="8" pageNumber="9">slide 2857</specimenCode>
; P.143.16; 
<collectionCode id="56EFAE49FFD7FFBAFDE45CFDFD55FE0F" box="[614,683,441,467]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFD495CFDFE85FE2B" ID-GBIF-Occurrence="4120752323" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2858" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFD495CFDFCDCFE0F" box="[715,802,441,467]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFB775CFDFA8AFE08" box="[1269,1396,441,468]" pageId="8" pageNumber="9">slide 2858</specimenCode>
; P.143.17; 
<collectionCode id="56EFAE49FFD7FFBAFEB45C99FE85FE2B" box="[310,379,477,503]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFE0F5C98FAFEFE2B" ID-GBIF-Occurrence="4120752308" box="[397,1280,476,503]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2859" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFE0F5C98FE1AFE2B" box="[397,484,476,503]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFC355C98FBC8FE2B" box="[951,1078,476,503]" pageId="8" pageNumber="9">slide 2859</specimenCode>
; P.143.18; 
<collectionCode id="56EFAE49FFD7FFBAFB395C99FAFEFE2B" box="[1211,1280,477,503]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFA9F5C98FC29FDC6" ID-GBIF-Occurrence="4120752310" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 2861" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFA9F5C98FA8AFE2B" box="[1309,1396,476,503]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFD085CBBFCF4FDC5" box="[650,778,511,537]" pageId="8" pageNumber="9">slide 2861</specimenCode>
; P.143.19; 
<collectionCode id="56EFAE49FFD7FFBAFC105F44FC29FDC6" box="[914,983,512,538]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFC775CBBFD55FDE1" ID-GBIF-Occurrence="4120752324" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 3010" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFC775CBBFBB3FDC6" box="[1013,1101,511,538]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFEE35F66FE1FFDE1" box="[353,481,546,573]" pageId="8" pageNumber="9">slide 3010</specimenCode>
; P.143.20; 
<collectionCode id="56EFAE49FFD7FFBAFDE45F67FD55FDE1" box="[614,683,547,573]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFD4E5F67FE83FDBC" ID-GBIF-Occurrence="4120752305" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCode="slide 3011" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<specimenCount id="26F8FD05FFD7FFBAFD4E5F67FCDDFDE1" box="[716,803,547,573]" pageId="8" pageNumber="9" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="60589EF7FFD7FFBAFB745F66FA8AFDE1" box="[1270,1396,546,573]" pageId="8" pageNumber="9">slide 3011</specimenCode>
; P.143.21; 
<collectionCode id="56EFAE49FFD7FFBAFEB65F02FE83FDBC" box="[308,381,582,608]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="64816604FFD8FFB5FF3F5ACCFC7BF834" ID-DOI="http://doi.org/10.5281/zenodo.7986635" ID-Zenodo-Dep="7986635" httpUri="https://zenodo.org/record/7986635/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1928,1954]" targetBox="[346,1228,954,1877]" targetPageId="7">
<paragraph id="3041368CFFD8FFB5FF3F5ACCFC7BF834" blockId="7.[189,1398,1928,2025]" pageId="7" pageNumber="8">
<emphasis id="028AEA9EFFD8FFB5FF3F5ACCFEF6F87E" bold="true" box="[189,264,1928,1954]" pageId="7" pageNumber="8">Fig. 1.</emphasis>
Schematic drawing of spermathecae of the specimens studied. 
<emphasis id="028AEA9EFFD8FFB5FC535ACCFC18F87E" bold="true" box="[977,998,1928,1954]" pageId="7" pageNumber="8">A</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FC745ACCFAE6F87E" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[1014,1304,1928,1954]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
<emphasis id="028AEA9EFFD8FFB5FC745ACCFAE6F87E" box="[1014,1304,1928,1954]" italics="true" pageId="7" pageNumber="8">Enchytraeus adrianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B957E5FFD8FFB5FA985ACCFA88F87E" box="[1306,1398,1928,1954]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="028AEA9EFFD8FFB5FF3F5AEFFF2FF819" bold="true" box="[189,209,1963,1989]" pageId="7" pageNumber="8">B</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FF605AE8FDCEF819" authority="(Cejka, 1913)" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[226,560,1963,1992]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD8FFB5FF605AE8FE7AF819" box="[226,388,1963,1989]" italics="true" pageId="7" pageNumber="8">E. krumbachi</emphasis>
(
<bibRefCitation id="546F4B7DFFD8FFB5FE155AE8FDD6F819" author="Cejka B." box="[407,552,1963,1992]" pageId="7" pageNumber="8" pagination="145 - 151" refId="ref20885" refString="Cejka B. 1913. Litorea krumbachi n. spec. n. gen. Ein Beitrag zur Systematik der Enchytraeiden. Zoologischer Anzeiger 17: 145 - 151." type="journal article" year="1913">Čejka, 1913</bibRefCitation>
)
</taxonomicName>
. 
<emphasis id="028AEA9EFFD8FFB5FDC05AEFFD85F819" bold="true" box="[578,635,1963,1989]" pageId="7" pageNumber="8">C–D</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FD0E5AE8FCF4F819" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[652,778,1963,1989]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="andrasi" status="sp. nov.">
<emphasis id="028AEA9EFFD8FFB5FD0E5AE8FCF4F819" box="[652,778,1963,1989]" italics="true" pageId="7" pageNumber="8">E. andrasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B957E5FFD8FFB5FC945AE8FC8FF819" box="[790,881,1964,1990]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="028AEA9EFFD8FFB5FCF95AEFFC71F819" bold="true" box="[891,911,1963,1989]" pageId="7" pageNumber="8">E</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FC225AE8FB97F819" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[928,1129,1963,1989]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="andrasiformis" status="sp. nov.">
<emphasis id="028AEA9EFFD8FFB5FC225AE8FB97F819" box="[928,1129,1963,1989]" italics="true" pageId="7" pageNumber="8">E. andrasiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B957E5FFD8FFB5FBF45AE8FB2CF81A" box="[1142,1234,1964,1990]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="028AEA9EFFD8FFB5FB595AEFFB15F819" bold="true" box="[1243,1259,1963,1989]" pageId="7" pageNumber="8">F</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FB7E5AE8FEB0F834" authority="Henle, 1837" authorityName="Henle" authorityYear="1837" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="albidus">
<emphasis id="028AEA9EFFD8FFB5FB7E5AE8FA88F819" box="[1276,1398,1963,1989]" italics="true" pageId="7" pageNumber="8">E. albidus</emphasis>
<bibRefCitation id="546F4B7DFFD8FFB5FF3D5A8AFEB0F834" author="Henle F. G. J." box="[191,334,1998,2024]" pageId="7" pageNumber="8" pagination="74 - 90" refId="ref21665" refString="Henle F. G. J. 1837. Ueber Enchytraeus, eine neue Anneliden-Gattung. Archiv fur Anatomie, Physiologie und Wissenschaftliche Medicin 74 - 90." type="book chapter" year="1837">Henle, 1837</bibRefCitation>
</taxonomicName>
. 
<emphasis id="028AEA9EFFD8FFB5FEDF5A8AFE8AF834" bold="true" box="[349,372,1998,2024]" pageId="7" pageNumber="8">G</emphasis>
. 
<taxonomicName id="F7FE4D0FFFD8FFB5FE015A8BFC83F834" authority="Nielsen &amp; Christensen, 1961" authorityName="Nielsen &amp; Christensen" authorityYear="1961" box="[387,893,1998,2024]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="irregularis">
<emphasis id="028AEA9EFFD8FFB5FE015A8BFDDAF834" box="[387,548,1998,2024]" italics="true" pageId="7" pageNumber="8">E. irregularis</emphasis>
<bibRefCitation id="546F4B7DFFD8FFB5FDA95A8AFC83F834" author="Nielsen C. O. &amp; Christensen B." box="[555,893,1998,2024]" pageId="7" pageNumber="8" pagination="1 - 23" refId="ref22135" refString="Nielsen C. O. &amp; Christensen B. 1961. The Enchytraeidae. Critical revision and taxonomy of European species. Supplement 1. Natura Jutlandica 10: 1 - 23." type="journal article" year="1961">Nielsen &amp; Christensen, 1961</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3041368CFFD7FFBAFF685FC3FE56FD7D" blockId="8.[189,1399,647,744]" box="[234,424,647,673]" pageId="8" pageNumber="9">
<emphasis id="028AEA9EFFD7FFBAFF685FC3FE56FD7D" bold="true" box="[234,424,647,673]" pageId="8" pageNumber="9">Other material</emphasis>
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5FEFFD3CFD34" blockId="8.[189,1399,647,744]" pageId="8" pageNumber="9">
<materialsCitation id="80963CD1FFD7FFBAFF3F5FEFFC2CFD19" ID-GBIF-Occurrence="4120752332" box="[189,978,682,709]" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="CROATIA" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCount="15" stateProvince="Istarska">
<collectingCountry id="48E9761CFFD7FFBAFF3F5FEFFEC4FD19" box="[189,314,683,709]" name="Croatia" pageId="8" pageNumber="9">CROATIA</collectingCountry>
• 
<specimenCount id="26F8FD05FFD7FFBAFED45FEEFE3EFD19" box="[342,448,682,709]" pageId="8" pageNumber="9">15 specs</specimenCount>
; same collection data as for holotype; 
<collectionCode id="56EFAE49FFD7FFBAFC0F5FEFFC2CFD19" box="[909,978,683,709]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
• 
<materialsCitation id="80963CD1FFD7FFBAFC6C5FEFFD40FD34" ID-GBIF-Occurrence="4120752338" collectingDate="2019-04-03" collectingDateMax="2019-10-15" collectingDateMin="2019-04-03" collectionCode="ELTE" collectorName="Julia Torok &amp; En." country="Croatia" county="Kamenjak Peninsula" latitude="44.85361" location="Kale Cove" longLatPrecision="1" longitude="13.980694" municipality="Adriatic Sea" pageId="8" pageNumber="9" specimenCount="3" stateProvince="Istarska">
<specimenCount id="26F8FD05FFD7FFBAFC6C5FEFFBBBFD19" box="[1006,1093,683,709]" pageId="8" pageNumber="9">3 specs</specimenCount>
(only for DNA analysis); same collection data as for holotype; 
<collectionCode id="56EFAE49FFD7FFBAFDF75F8AFD40FD34" box="[629,702,718,744]" pageId="8" pageNumber="9">ELTE</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78E46507FFD7FFB8FF3F5E4BFB2EFB80" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="3041368CFFD7FFBAFF3F5E4BFEB1FCF5" blockId="8.[189,335,783,809]" box="[189,335,783,809]" pageId="8" pageNumber="9">
<heading id="6B0981E0FFD7FFBAFF3F5E4BFEB1FCF5" bold="true" box="[189,335,783,809]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="028AEA9EFFD7FFBAFF3F5E4BFEB1FCF5" bold="true" box="[189,335,783,809]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5E7AFD4CFC38" blockId="8.[189,1399,829,997]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5E7AFE90FC8A" baselines="849,849" box="[189,366,830,856]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="8" pageNumber="9">MEASUREMENTS</smallCapsWord>
. Large specimens. 
<typeStatus id="EF45882EFFD7FFBAFDCF5E79FD44FC8B" box="[589,698,829,855]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
<quantity id="F7069B69FFD7FFBAFD425E7AFCD9FC84" box="[704,807,830,856]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="8" pageNumber="9" unit="mm" value="22.8">22.8 mm</quantity>
long, 570 μm wide at VIII and 770 μm at clitellum, fixed, segment number 60. Body length of 
<typeStatus id="EF45882EFFD7FFBAFD3F5E25FCD3FCA7" box="[701,813,865,891]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
<quantity id="F7069B69FFD7FFBAFCB45E25FC47FCA7" box="[822,953,865,891]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="3.0" metricValueMin="1.4" pageId="8" pageNumber="9" unit="mm" value="22.0" valueMax="30.0" valueMin="14.0">14–30 mm</quantity>
, width 450–780 μm at VIII and 550– 1000 μm at clitellum, in vivo. Length of fixed specimens 
<quantity id="F7069B69FFD7FFBAFCD15EC0FC13FC43" box="[851,1005,900,927]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" metricValueMax="2.4899999999999998" metricValueMin="0.77" pageId="8" pageNumber="9" unit="mm" value="16.3" valueMax="24.9" valueMin="7.7">7.7–24.9 mm</quantity>
(in 
<specimenCount id="26F8FD05FFD7FFBAFB985EC1FB47FC42" box="[1050,1209,900,927]" pageId="8" pageNumber="9" type="generic">one specimen</specimenCount>
<quantity id="F7069B69FFD7FFBAFB3D5EC0FAD4FC43" box="[1215,1322,900,927]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="8" pageNumber="9" unit="mm" value="28.2">28.2 mm</quantity>
, width 410–730 μm at VIII and 490–950 μm at clitellum). Length of the first 12 segments 
<quantity id="F7069B69FFD7FFBAFB205EE3FACAFC1E" box="[1186,1332,935,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.8" metricValueMin="2.2" pageId="8" pageNumber="9" unit="mm" value="3.5" valueMax="4.8" valueMin="2.2">2.2–4.8 mm</quantity>
, after fixation. Segment number 51–84 (N = 34).
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5948FA87FB6F" blockId="8.[189,1401,1036,1203]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5948FEDCFBF8" baselines="1055,1055" box="[189,290,1036,1062]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Chaetae" pageId="8" pageNumber="9">CHAETAE</smallCapsWord>
. Chaetal formula according to 
<bibRefCitation id="546F4B7DFFD7FFBAFD1B5948FC1FFBFA" author="Schmelz R. M. &amp; Collado R." box="[665,993,1036,1062]" pageId="8" pageNumber="9" pagination="1 - 176" refId="ref22440" refString="Schmelz R. M. &amp; Collado R. 2010. A guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta). Soil Organisms 82: 1 - 176." type="journal article" year="2010">Schmelz &amp; Collado (2010)</bibRefCitation>
: 2,3–2,3:3,4–2,3. Chaetae straight with ental hook, about equal in size within bundle, 72–112μm × 5–7.5 μm preclitellarly and 70–100 μm × 5–7.5 μm posteriorly. In ventral preclitellar bundles of some specimens 3 chaetae, and only one or two bundles with 4 chaetae, in other specimens 4 chaetae in most of all other bundles. Often 2–3 surplus chaetae near the bundles (
<figureCitation id="A8C52A09FFD7FFBAFE6E59DDFDB6FB6F" box="[492,584,1177,1203]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2B</figureCitation>
). Chaetae in XII absent ventrally, but 2–3 per bundle present laterally.
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F599FFE6DFB28" blockId="8.[189,403,1242,1269]" box="[189,403,1242,1269]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F599FFF02FB2F" baselines="1262,1262" box="[189,252,1243,1269]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="8" pageNumber="9">HEAD</smallCapsWord>
<smallCapsWord id="36A7A050FFD7FFBAFE8659A4FEC7FB2F" baselines="1262" box="[260,313,1248,1267]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pore" pageId="8" pageNumber="9">PORE</smallCapsWord>
. At 0/I.
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5858FC20FAEA" blockId="8.[189,990,1307,1334]" box="[189,990,1307,1334]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5858FEC7FAE8" baselines="1327,1327" box="[189,313,1308,1334]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Epidermal" pageId="8" pageNumber="9">EPIDERMAL</smallCapsWord>
<smallCapsWord id="36A7A050FFD7FFBAFEC35865FE6BFAE8" baselines="1327" box="[321,405,1313,1332]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="glands" pageId="8" pageNumber="9">GLANDS</smallCapsWord>
. Often brown, arranged in 3–4 rows per segment.
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5819FC8EFA61" blockId="8.[188,1399,1373,1470]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5819FEC7FAAA" baselines="1392,1393" box="[189,313,1373,1399]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Clitellum" pageId="8" pageNumber="9">CLITELLUM</smallCapsWord>
. Girdle-shaped, in XII–XIII, hyalocytes and granulocytes in dense transverse rows dorsally and laterally (
<figureCitation id="A8C52A09FFD7FFBAFEDC58C4FE47FA46" box="[350,441,1408,1434]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
), but in well-developed sexual condition gland cells in reticulate pattern (
<figureCitation id="A8C52A09FFD7FFBAFA8F58C4FA94FA46" box="[1293,1386,1408,1434]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2D</figureCitation>
). Mostly only granulocytes between male pores (
<figureCitation id="A8C52A09FFD7FFBAFD6A58E7FC9AFA61" box="[744,868,1443,1469]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2E–F</figureCitation>
).
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F58A1FCD9F9FE" blockId="8.[189,1398,1509,1570]" pageId="8" pageNumber="9">
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. About 1.3–1.8 × as long as wide, rounded posteriorly, sides slightly merging anteriad often with 2 small aggregations of refractile globules (
<figureCitation id="A8C52A09FFD7FFBAFD3E5B4CFCE4F9FE" box="[700,794,1544,1570]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5B0EFEECF911" blockId="8.[189,1399,1609,1741]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5B0EFEA4F9BE" baselines="1629,1629" box="[189,346,1610,1636]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Oesophageal" pageId="8" pageNumber="9">OESOPHAGEAL</smallCapsWord>
<smallCapsWord id="36A7A050FFD7FFBAFEE75B0BFE12F9BE" baselines="1629" box="[357,492,1615,1634]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="appendages" pageId="8" pageNumber="9">APPENDAGES</smallCapsWord>
. Pair of blind-ending tubes in III / IV, with common root inserting dorsally behind pharyngeal pad. Mostly all primary pharyngeal glands widely or slightly connected dorsally with ventral lobes (
<figureCitation id="A8C52A09FFD7FFBAFE225BD4FE01F976" box="[416,511,1680,1706]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2H</figureCitation>
). In some specimens first pair separate (
<figureCitation id="A8C52A09FFD7FFBAFC575BD4FBCAF976" box="[981,1076,1680,1706]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2G</figureCitation>
). Ventral lobes of third pair largest.
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5BB1FAE9F8D2" blockId="8.[189,1303,1780,1807]" box="[189,1303,1780,1807]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5BB1FEE8F8D1" baselines="1800,1800" box="[189,278,1781,1807]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dorsal" pageId="8" pageNumber="9">DORSAL</smallCapsWord>
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<smallCapsWord id="36A7A050FFD7FFBAFEEF5BBEFE47F8D1" baselines="1800" box="[365,441,1786,1805]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="vessel" pageId="8" pageNumber="9">VESSEL</smallCapsWord>
. From XIV–XV, blood colourless. Anterior bifurcation near prostomium.
</paragraph>
<paragraph id="3041368CFFD7FFBAFF3F5A72FCE2F8AF" blockId="8.[189,1398,1846,1907]" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5A72FECDF892" baselines="1865,1865" box="[189,307,1846,1872]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Nephridia" pageId="8" pageNumber="9">NEPHRIDIA</smallCapsWord>
. Four pairs of preclitellar nephridia from 6/ 7–9 /10, anteseptale funnel only, postseptale bulged, efferent duct short, origin postero-ventrally.
</paragraph>
<paragraph id="3041368CFFD7FFBBFF3F5ADFFB59FE9A" blockId="8.[189,1399,1946,2008]" lastBlockId="9.[189,1398,265,327]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<smallCapsWord id="36A7A050FFD7FFBAFF3F5ADFFE95F86F" baselines="1966,1966" box="[189,363,1947,1973]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coelomocytes" pageId="8" pageNumber="9">COELOMOCYTES</smallCapsWord>
. Oval or narrowed at one end, texture granulated, about 24–40 μm long, in vivo (
<figureCitation id="A8C52A09FFD7FFBAFA9B5ADEFF37F804" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2I– J</figureCitation>
) [15–30 μm, fixed (
<figureCitation id="A8C52A09FFD7FFBAFE445AFAFDD5F804" box="[454,555,1982,2008]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="8" pageNumber="9">Fig. 2K</figureCitation>
)]. In addition, many shining, hyaline, round or tetragonal corpuscles [diameter (10–19 μm)] also present (
<figureCitation id="A8C52A09FFD6FFBBFDE95C4DFD41FEFF" box="[619,703,265,291]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="9" pageNumber="10">Fig. 2J</figureCitation>
), which at lower magnification shine like grains of sand. In young specimens corpuscles always fewer. Note, corpuscles not visible after fixation.
</paragraph>
<paragraph id="3041368CFFD6FFBBFF3F5C2AFDFCFE54" blockId="9.[189,514,366,392]" box="[189,514,366,392]" pageId="9" pageNumber="10">
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. Absent.
</paragraph>
<caption id="64816604FFD6FFBBFF3F5BD5FE16F835" ID-DOI="http://doi.org/10.5281/zenodo.7986637" ID-Zenodo-Dep="7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1681,1707]" targetBox="[189,1398,461,1643]" targetPageId="9">
<paragraph id="3041368CFFD6FFBBFF3F5BD5FE16F835" blockId="9.[188,1399,1681,2025]" pageId="9" pageNumber="10">
<emphasis id="028AEA9EFFD6FFBBFF3F5BD5FEF2F977" bold="true" box="[189,268,1681,1707]" pageId="9" pageNumber="10">Fig. 2.</emphasis>
Micrograph of 
<taxonomicName id="F7FE4D0FFFD6FFBBFE445BD5FD12F977" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[454,748,1681,1707]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
<emphasis id="028AEA9EFFD6FFBBFE445BD5FD12F977" box="[454,748,1681,1707]" italics="true" pageId="9" pageNumber="10">Enchytraeus adrianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B957E5FFD6FFBBFD765BD6FCB1F970" box="[756,847,1682,1708]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="028AEA9EFFD6FFBBFCD55BD5FC39F970" bold="true" box="[855,967,1681,1708]" pageId="9" pageNumber="10">A–D, I–J</emphasis>
. In vivo. 
<emphasis id="028AEA9EFFD6FFBBFBB85BD6FB45F970" bold="true" box="[1082,1211,1682,1708]" pageId="9" pageNumber="10">E–H, K–L</emphasis>
. Fixed, stained. 
<emphasis id="028AEA9EFFD6FFBBFF3F5BF0FF2CF912" bold="true" box="[189,210,1716,1742]" pageId="9" pageNumber="10">A</emphasis>
. Brain (small aggregations of refractile globules marked with arrows). 
<emphasis id="028AEA9EFFD6FFBBFB8E5BF1FBDEF913" bold="true" box="[1036,1056,1717,1743]" pageId="9" pageNumber="10">B</emphasis>
. Chaetae in a ventral bundles with 3 surplus chaetae. 
<emphasis id="028AEA9EFFD6FFBBFE635B9CFE08F92E" bold="true" box="[481,502,1752,1778]" pageId="9" pageNumber="10">C</emphasis>
. Clitellar glands in regular rows. 
<emphasis id="028AEA9EFFD6FFBBFC1A5B9CFC53F92E" bold="true" box="[920,941,1752,1778]" pageId="9" pageNumber="10">D</emphasis>
. Clitellar glands in reticulate pattern. 
<emphasis id="028AEA9EFFD6FFBBFF3F5BBFFF2FF8C9" bold="true" box="[189,209,1787,1813]" pageId="9" pageNumber="10">E</emphasis>
. Paratype (slide 2748, P.143.3, ELTE). 
<emphasis id="028AEA9EFFD6FFBBFD305BBFFD3AF8C9" bold="true" box="[690,708,1787,1813]" pageId="9" pageNumber="10">F</emphasis>
. Slide 2684 (ELTE). 
<emphasis id="028AEA9EFFD6FFBBFC485BBFFC1FF8C9" bold="true" box="[970,993,1787,1813]" pageId="9" pageNumber="10">H</emphasis>
. Slide 2761 (ELTE). 
<emphasis id="028AEA9EFFD6FFBBFB6A5BBFFB01F8C9" bold="true" box="[1256,1279,1787,1813]" pageId="9" pageNumber="10">K</emphasis>
. Paratype (slide 2851, P.143.10, ELTE). 
<emphasis id="028AEA9EFFD6FFBBFDAC5A5AFDBCF8E4" bold="true" box="[558,578,1822,1848]" pageId="9" pageNumber="10">L</emphasis>
. Paratype (slide 2760, P.143.6). 
<emphasis id="028AEA9EFFD6FFBBFC525A5AFBFBF8E4" bold="true" box="[976,1029,1822,1848]" pageId="9" pageNumber="10">E–F</emphasis>
. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). 
<emphasis id="028AEA9EFFD6FFBBFEF05A20FE79F8A2" bold="true" box="[370,391,1892,1918]" pageId="9" pageNumber="10">G</emphasis>
. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. 
<emphasis id="028AEA9EFFD6FFBBFAA95A21FABCF8A3" bold="true" box="[1323,1346,1893,1919]" pageId="9" pageNumber="10">H</emphasis>
. All pharyngeal glands connected dorsally (spermathecae marked with arrows). 
<emphasis id="028AEA9EFFD6FFBBFBA95ACCFBA2F87E" bold="true" box="[1067,1116,1928,1954]" pageId="9" pageNumber="10">I–K</emphasis>
. Coelomocytes (in J the small hyaline, refracting corpuscules also present). 
<emphasis id="028AEA9EFFD6FFBBFC935AEFFCDBF819" bold="true" box="[785,805,1963,1989]" pageId="9" pageNumber="10">L</emphasis>
. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm.
</paragraph>
</caption>
<paragraph id="3041368CFFD5FFB8FF3F5C4DFC08FE9B" blockId="10.[189,1399,265,327]" pageId="10" pageNumber="11">
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<smallCapsWord id="36A7A050FFD5FFB8FE8E5C4AFEBEFEFD" baselines="284" box="[268,320,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="sacs" pageId="10" pageNumber="11">SACS</smallCapsWord>
. Two or three paired lobes of sperm sacs, very large, filling the coelom of IX /X–XI (
<figureCitation id="A8C52A09FFD5FFB8FA925C4DFA94FEFF" box="[1296,1386,265,291]" captionStart="Fig" captionStartId="9.[189,232,1681,1707]" captionTargetBox="[189,1398,461,1643]" captionTargetId="figure-48@9.[189,1398,461,1646]" captionTargetPageId="9" captionText="Fig. 2. Micrograph of Enchytraeus adrianensis sp. nov. A–D, I–J. In vivo. E–H, K–L. Fixed, stained. A. Brain (small aggregations of refractile globules marked with arrows). B. Chaetae in a ventral bundles with 3 surplus chaetae. C. Clitellar glands in regular rows. D. Clitellar glands in reticulate pattern. E. Paratype (slide 2748, P.143.3, ELTE). F. Slide 2684 (ELTE). H. Slide 2761 (ELTE). K. Paratype (slide 2851, P.143.10, ELTE). L. Paratype (slide 2760, P.143.6). E–F. Clitellar glands, ventral view (male copulatory organs marked with black arrow, between these organs the granulocytes marked with white arrows). G. Paratype (slide 2856, P.143.15, ELTE), first pharyngeal glands free dorsally. H. All pharyngeal glands connected dorsally (spermathecae marked with arrows). I–K. Coelomocytes (in J the small hyaline, refracting corpuscules also present). L. Large paired lobes of sperm sacs. Scale bars: A–I, K–L = 50 μm; J = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7986637" httpUri="https://zenodo.org/record/7986637/files/figure.png" pageId="10" pageNumber="11">Fig. 2L</figureCitation>
). Testes and sperm funnels in XI, ovaries, male pores and glands in XII.
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F5C2AFB2AFE2D" blockId="10.[188,1399,365,498]" pageId="10" pageNumber="11">
<smallCapsWord id="36A7A050FFD5FFB8FF3F5C2AFEFBFE5A" baselines="385,385" box="[189,261,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sperm" pageId="10" pageNumber="11">SPERM</smallCapsWord>
<smallCapsWord id="36A7A050FFD5FFB8FE8F5C37FE94FE5A" baselines="385" box="[269,362,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="funnels" pageId="10" pageNumber="11">FUNNELS</smallCapsWord>
. 350–690 μm long in vivo (300–500 μm, fixed), 1.7–4 × as long as wide, collar narrower than funnel body (
<figureCitation id="A8C52A09FFD5FFB8FE125CD5FE13FE77" box="[400,493,401,427]" captionStart="Fig" captionStartId="11.[189,232,1787,1813]" captionTargetBox="[207,1365,612,1734]" captionTargetId="figure-129@11.[207,1380,612,1747]" captionTargetPageId="11" captionText="Fig. 3. Micrograph of Enchytraeus adrianensis sp. nov. A, C, G–H. In vivo. B, D–F, I. Fixed, stained. A. Sperm funnels. B. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). C–F. Multiple male glands (E–F primary bulb marked with arrow). D. Paratype (slide 2748, P.143.3, ELTE). E. Paratype (slide 2748, P.143.3, ELTE). F. Holotype (En.2. slide 2842, ELTE). I. Paratype (slide 2827, P. 143.8, ELTE). F. Holotype (En.2. slide 2842, ELTE). G–I. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7986639" httpUri="https://zenodo.org/record/7986639/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
). Vasa deferentia distinctly tripartite, long, extending into segments XV–XIX. After sperm funnel and before male opening, vasa deferentia slender and ciliated (20–40 μm wide in vivo and fixed equally), middle part thickened (50–95 μm wide) and unciliated (
<figureCitation id="A8C52A09FFD5FFB8FBE85C93FB38FE2D" box="[1130,1222,471,497]" captionStart="Fig" captionStartId="11.[189,232,1787,1813]" captionTargetBox="[207,1365,612,1734]" captionTargetId="figure-129@11.[207,1380,612,1747]" captionTargetPageId="11" captionText="Fig. 3. Micrograph of Enchytraeus adrianensis sp. nov. A, C, G–H. In vivo. B, D–F, I. Fixed, stained. A. Sperm funnels. B. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). C–F. Multiple male glands (E–F primary bulb marked with arrow). D. Paratype (slide 2748, P.143.3, ELTE). E. Paratype (slide 2748, P.143.3, ELTE). F. Holotype (En.2. slide 2842, ELTE). I. Paratype (slide 2827, P. 143.8, ELTE). F. Holotype (En.2. slide 2842, ELTE). G–I. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7986639" httpUri="https://zenodo.org/record/7986639/files/figure.png" pageId="10" pageNumber="11">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F5F5DFCB4FDEF" blockId="10.[189,842,536,563]" box="[189,842,536,563]" pageId="10" pageNumber="11">
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. 75–100 μm long, heads 20–30 μm, fixed.
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F5F1FFCA5FC9B" blockId="10.[189,1399,602,840]" pageId="10" pageNumber="11">
<smallCapsWord id="36A7A050FFD5FFB8FF3F5F1FFF01FDAF" baselines="622,622" box="[189,255,603,629]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="10" pageNumber="11">MALE</smallCapsWord>
<smallCapsWord id="36A7A050FFD5FFB8FE8A5F24FE6EFDAF" baselines="622" box="[264,400,608,627]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="copulatory" pageId="10" pageNumber="11">COPULATORY</smallCapsWord>
<smallCapsWord id="36A7A050FFD5FFB8FE1B5F24FE10FDAF" baselines="622" box="[409,494,608,627]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="10" pageNumber="11">ORGANS</smallCapsWord>
. Male glands multiple: one large rounded primary bulb near male pore, and 6–9 smaller secondary glands, mostly 8 (usually 4–5 on each side, but sometimes 
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one side and only 3 at other). Besides, near male opening in middle 2–3 smaller additional glands. Glands arranged roughly in semicircle around male pore and primary bulb (diameter 200–350 μm, fixed) (
<figureCitation id="A8C52A09FFD5FFB8FB695F80FA94FD02" box="[1259,1386,708,734]" captionStart="Fig" captionStartId="11.[189,232,1787,1813]" captionTargetBox="[207,1365,612,1734]" captionTargetId="figure-129@11.[207,1380,612,1747]" captionTargetPageId="11" captionText="Fig. 3. Micrograph of Enchytraeus adrianensis sp. nov. A, C, G–H. In vivo. B, D–F, I. Fixed, stained. A. Sperm funnels. B. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). C–F. Multiple male glands (E–F primary bulb marked with arrow). D. Paratype (slide 2748, P.143.3, ELTE). E. Paratype (slide 2748, P.143.3, ELTE). F. Holotype (En.2. slide 2842, ELTE). I. Paratype (slide 2827, P. 143.8, ELTE). F. Holotype (En.2. slide 2842, ELTE). G–I. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7986639" httpUri="https://zenodo.org/record/7986639/files/figure.png" pageId="10" pageNumber="11">Fig. 3C–F</figureCitation>
). Primary bulb 60–125 μm long and 50–95 μm wide, secondary glands near to primary glands longer than outside ones (40–100 μm long and 20–50 μm wide). Male pores covered by lip-like folds (100–160 μm wide), ventral recess absent (all data in fixed specimens).
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F5E2BFB2EFB80" blockId="10.[189,1399,879,1116]" pageId="10" pageNumber="11">
<smallCapsWord id="36A7A050FFD5FFB8FF3F5E2BFE9AFC5B" baselines="898,898" box="[189,356,879,905]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermathecae" pageId="10" pageNumber="11">SPERMATHECAE</smallCapsWord>
(
<figureCitation id="A8C52A09FFD5FFB8FEFB5E2BFE24FC55" box="[377,474,879,905]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[346,1228,954,1877]" captionTargetId="figure-231@7.[313,1275,942,1892]" captionTargetPageId="7" captionText="Fig. 1. Schematic drawing of spermathecaeof the specimens studied.A. Enchytraeus adrianensissp. nov. B. E. krumbachi (Čejka, 1913). C–D. E. andrasi sp. nov. E. E. andrasiformis sp. nov. F. E. albidus Henle, 1837. G. E. irregularis Nielsen &amp; Christensen, 1961." figureDoi="http://doi.org/10.5281/zenodo.7986635" httpUri="https://zenodo.org/record/7986635/files/figure.png" pageId="10" pageNumber="11">Figs 1A</figureCitation>
, 
<figureCitation id="A8C52A09FFD5FFB8FE6F5E2BFDD4FC55" box="[493,554,879,905]" captionStart="Fig" captionStartId="11.[189,232,1787,1813]" captionTargetBox="[207,1365,612,1734]" captionTargetId="figure-129@11.[207,1380,612,1747]" captionTargetPageId="11" captionText="Fig. 3. Micrograph of Enchytraeus adrianensis sp. nov. A, C, G–H. In vivo. B, D–F, I. Fixed, stained. A. Sperm funnels. B. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). C–F. Multiple male glands (E–F primary bulb marked with arrow). D. Paratype (slide 2748, P.143.3, ELTE). E. Paratype (slide 2748, P.143.3, ELTE). F. Holotype (En.2. slide 2842, ELTE). I. Paratype (slide 2827, P. 143.8, ELTE). F. Holotype (En.2. slide 2842, ELTE). G–I. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7986639" httpUri="https://zenodo.org/record/7986639/files/figure.png" pageId="10" pageNumber="11">3G–I</figureCitation>
). With ectal duct, ampulla without diverticula, separate openings into oesophagus. Spermathecal ectal pore at 
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/ 
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. Ectal duct with different length (90–160 μm in vivo, 70– 160 μm, fixed), covered with gland cells (70–117 μm widely in vivo and 60–100 μm widely, fixed). In most specimens, ectal duct near ampulla without glands for short stretch, length about 7–12 μm. Canal of ectal duct 6–9 μm wide, widening entad. Ampulla oval or spherical, about 70–115 μm wide and 75–150 μm long, with distinct, about 13–20 μm wide walls, lumen with masses of spermatozoa, ental duct very short, connecting laterally with oesophagus. 2–7 mature eggs present at a time.
</paragraph>
</subSubSection>
<subSubSection id="78E46507FFD5FFBEFF3F59C7FB39FE77" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="3041368CFFD5FFB8FF3F59C7FE32FB41" blockId="10.[189,460,1155,1181]" box="[189,460,1155,1181]" pageId="10" pageNumber="11">
<heading id="6B0981E0FFD5FFB8FF3F59C7FE32FB41" bold="true" box="[189,460,1155,1181]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="028AEA9EFFD5FFB8FF3F59C7FE32FB41" bold="true" box="[189,460,1155,1181]" pageId="10" pageNumber="11">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F59F6FBA2FAA0" blockId="10.[188,1399,1202,1404]" pageId="10" pageNumber="11">
In the 
<taxonomicName id="F7FE4D0FFFD5FFB8FE8E59F6FDFEFB10" authorityName="Henle" authorityYear="1837" box="[268,512,1202,1228]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="albidus">
<emphasis id="028AEA9EFFD5FFB8FE8E59F6FDFEFB10" box="[268,512,1202,1228]" italics="true" pageId="10" pageNumber="11">Enchytraeus albidus</emphasis>
</taxonomicName>
species complex as circumscribed in 
<bibRefCitation id="546F4B7DFFD5FFB8FC4059F6FB50FB10" author="Erseus C. &amp; Klinth M. J. &amp; Rota E. &amp; Wit P. D. &amp; Gustafsson D. R. &amp; Martinsson S." box="[962,1198,1202,1228]" pageId="10" pageNumber="11" pagination="105 - 133" refId="ref21493" refString="Erseus C., Klinth M. J., Rota E., Wit P. D., Gustafsson D. R. &amp; Martinsson S. 2019. The popular model annelid Enchytraeus albidus is only one species in a complex of seashore white worms (Clitellata, Enchytraeidae). Organisms Diversity &amp; Evolution 19: 105 - 133. https: // doi. org / 10.1007 / s 13127 - 019 - 00402 - 6" type="journal article" year="2019">
Erséus 
<emphasis id="028AEA9EFFD5FFB8FB9859F7FBA8FB10" box="[1050,1110,1202,1228]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2019)
</bibRefCitation>
, the new species and two other species of which sequences are available (
<taxonomicName id="F7FE4D0FFFD5FFB8FCE65992FC1BFB33" authorityName="Klinth, Erseus &amp; Rota" authorityYear="2019" box="[868,997,1237,1263]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="albellus">
<emphasis id="028AEA9EFFD5FFB8FCE65992FC1BFB33" box="[868,997,1237,1263]" italics="true" pageId="10" pageNumber="11">E. albellus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7FE4D0FFFD5FFB8FC745992FB40FB33" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[1014,1214,1237,1263]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" isUncertain="true" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FC745992FBF1FB33" box="[1014,1039,1238,1263]" italics="true" pageId="10" pageNumber="11">E.</emphasis>
cf. 
<emphasis id="028AEA9EFFD5FFB8FBBD5991FB40FB33" box="[1087,1214,1237,1263]" italics="true" pageId="10" pageNumber="11">krumbachi</emphasis>
</taxonomicName>
) have tripartite vasa deferentia. In 
<taxonomicName id="F7FE4D0FFFD5FFB8FE2059BDFDDFFACE" authorityName="Klinth, Erseus &amp; Rota" authorityYear="2019" box="[418,545,1272,1298]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="albellus">
<emphasis id="028AEA9EFFD5FFB8FE2059BDFDDFFACE" box="[418,545,1272,1298]" italics="true" pageId="10" pageNumber="11">E. albellus</emphasis>
</taxonomicName>
, the ductus is ciliated in its full length, whereas in the new species and in 
<taxonomicName id="F7FE4D0FFFD5FFB8FF595858FE85FAE9" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[219,379,1307,1333]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FF595858FE85FAE9" box="[219,379,1307,1333]" italics="true" pageId="10" pageNumber="11">E. krumbachi</emphasis>
</taxonomicName>
the thickened middle part is not ciliated. Moreover, 
<taxonomicName id="F7FE4D0FFFD5FFB8FC605858FB9FFAE9" authorityName="Klinth, Erseus &amp; Rota" authorityYear="2019" box="[994,1121,1307,1333]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="albellus">
<emphasis id="028AEA9EFFD5FFB8FC605858FB9FFAE9" box="[994,1121,1307,1333]" italics="true" pageId="10" pageNumber="11">E. albellus</emphasis>
</taxonomicName>
is different, because its spermathecal ampulla is irregular sac-like with one smaller diverticulum dorsally, but in the new species the ampulla is oval or spherical with a well developed wall without diverticula.
</paragraph>
<paragraph id="3041368CFFD5FFB8FF3F58E7FEC2F811" blockId="10.[188,1400,1443,1997]" pageId="10" pageNumber="11">
The new species is closely related to 
<taxonomicName id="F7FE4D0FFFD5FFB8FD1E58E0FCBCFA61" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[668,834,1443,1469]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FD1E58E0FCBCFA61" box="[668,834,1443,1469]" italics="true" pageId="10" pageNumber="11">E. krumbachi</emphasis>
</taxonomicName>
based on morphological and molecular data. Morphological similarity is primarily observable in the form of the spermatheca and the absence of the ventral recess of the clitellum. In both species the ampullae are without diverticula, but in 
<taxonomicName id="F7FE4D0FFFD5FFB8FB5558AEFA88F9D8" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[1239,1398,1514,1540]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FB5558AEFA88F9D8" box="[1239,1398,1514,1540]" italics="true" pageId="10" pageNumber="11">E. krumbachi</emphasis>
</taxonomicName>
round and the diameter is slightly larger (70–140 μm in the Ligurian Sea specimens of 
<taxonomicName id="F7FE4D0FFFD5FFB8FB575B4AFA89F9FB" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[1237,1399,1549,1575]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FB575B4AFA89F9FB" box="[1237,1399,1549,1575]" italics="true" pageId="10" pageNumber="11">E. krumbachi</emphasis>
</taxonomicName>
collected in this study but in the new species often oval and only 70–115 μm). Furthermore, the new species is bigger (segment number 51–84, body length 
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and 490–950 μm wide at clitellum vs segment number 40–66, body length 
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and 500–860 μm wide at clitellum, in vivo). No doubt, that the ventral recess absent in both species, but in the male copulatory organ the primary bulb about the same size in both species. In the new species mostly 8 long, narrower secondary glands are in a semicircle, around male pore and primary bulb, but in 
<taxonomicName id="F7FE4D0FFFD5FFB8FCDD5BA5FBFEF926" authorityName="CE" authorityYear="1689" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[863,1024,1760,1786]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krumbachi">
<emphasis id="028AEA9EFFD5FFB8FCDD5BA5FBFEF926" box="[863,1024,1760,1786]" italics="true" pageId="10" pageNumber="11">E. krumbachi</emphasis>
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the primary bulb is surrounded with very much smaller secondary glands. Although the molecular data also showed a close relationship between the two species, they still supported a certain species-level separation. 
<taxonomicName id="F7FE4D0FFFD5FFB8FBD25A62FA8BF89C" authority="Arslan &amp; Timm, 2018" authorityName="Arslan &amp; Timm, CE" authorityYear="2018" box="[1104,1397,1830,1856]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="polatdemiri">
<emphasis id="028AEA9EFFD5FFB8FBD25A62FA8BF89C" box="[1104,1397,1830,1856]" italics="true" pageId="10" pageNumber="11">Enchytraeus polatdemiri</emphasis>
</taxonomicName>
differs from the new species by its smaller size (segment number 40–55, length 
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and 410– 580 μm wide at clitellum), and the sperm funnel is 5 × as long as wide (vs 1.7–4 × as long as in the new species) and the vasa deferentia are not tripartite, ciliated throughout, and mostly confined to XII (vs XV–XIX).
</paragraph>
<paragraph id="3041368CFFD4FFB9FF3F5C4DFAA4FE50" blockId="11.[189,1399,265,397]" pageId="11" pageNumber="12">
<taxonomicName id="F7FE4D0FFFD4FFB9FF3F5C4DFE54FEFF" authorityName="CE" authorityYear="1687" baseAuthorityName="Michaelsen" baseAuthorityYear="1885" box="[189,426,265,291]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="moebii">
<emphasis id="028AEA9EFFD4FFB9FF3F5C4DFE54FEFF" box="[189,426,265,291]" italics="true" pageId="11" pageNumber="12">Enchytraeus moebii</emphasis>
</taxonomicName>
is similar to the new species in the form of the male apparatus (large primary bulb and smaller accessory glands especially on the basis of figure of 
<bibRefCitation id="546F4B7DFFD4FFB9FC325C68FB76FE9A" author="Michaelsen W." box="[944,1160,300,326]" pageId="11" pageNumber="12" refId="ref22019" refString="Michaelsen W. 1886. Untersuchungen uber Enchytraeus Mobii Mich. und andere Enchytraeiden. Lipsius &amp; Tischer, Kiel." type="book" year="1886">Michaelsen (1886</bibRefCitation>
: fig. 3 pl. II), but the principal differences are that the vasa deferentia are not tripartite, ciliated throughout and the collar of sperm funnel is wider than the funnel body (vs collar narrower than funnel body in the new species).
</paragraph>
<paragraph id="3041368CFFD4FFBEFF3F5CF0FD14FEB5" blockId="11.[189,1399,436,533]" lastBlockId="12.[189,1399,265,362]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<taxonomicName id="F7FE4D0FFFD4FFB9FF3F5CF0FE27FE12" authorityName="Nielsen &amp; Christensen" authorityYear="1961" box="[189,473,436,462]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="irregularis">
<emphasis id="028AEA9EFFD4FFB9FF3F5CF0FE27FE12" box="[189,473,436,462]" italics="true" pageId="11" pageNumber="12">Enchytraeus irregularis</emphasis>
</taxonomicName>
differs principally from the new species by the well developed diverticulum of the spermatheca, by the smaller primary bulb of male apparatus (diameter 40–80 μm vs 60–125 μm in the new species), and the vasa deferentia of 
<taxonomicName id="F7FE4D0FFFD4FFB9FD3F5CBFFCA3FDC8" authorityName="Nielsen &amp; Christensen" authorityYear="1961" box="[701,861,506,532]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="irregularis">
<emphasis id="028AEA9EFFD4FFB9FD3F5CBFFCA3FDC8" box="[701,861,506,532]" italics="true" pageId="11" pageNumber="12">E. irregularis</emphasis>
</taxonomicName>
are uniform, ciliated throughout and confined to XII. Very characteristic trait for 
<taxonomicName id="F7FE4D0FFFD3FFBEFDEB5C4EFCF0FEFF" authorityName="Nielsen &amp; Christensen" authorityYear="1961" box="[617,782,265,291]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="12" pageNumber="13" phylum="Annelida" rank="species" species="irregularis">
<emphasis id="028AEA9EFFD3FFBEFDEB5C4EFCF0FEFF" box="[617,782,265,291]" italics="true" pageId="12" pageNumber="13">E. irregularis</emphasis>
</taxonomicName>
is the oval or squarish glandular structure without external orifice (‘accessory sexual glands’) after or before male copulatory organs in XIII, XII or XI but in some specimens these organs may be absent.
</paragraph>
<caption id="64816604FFD4FFB9FF3F5BBFFC54F835" ID-DOI="http://doi.org/10.5281/zenodo.7986639" ID-Zenodo-Dep="7986639" httpUri="https://zenodo.org/record/7986639/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1787,1813]" targetBox="[207,1365,612,1734]" targetPageId="11">
<paragraph id="3041368CFFD4FFB9FF3F5BBFFC54F835" blockId="11.[189,1399,1787,2025]" pageId="11" pageNumber="12">
<emphasis id="028AEA9EFFD4FFB9FF3F5BBFFEF2F8C9" bold="true" box="[189,268,1787,1813]" pageId="11" pageNumber="12">Fig. 3.</emphasis>
Micrograph of 
<taxonomicName id="F7FE4D0FFFD4FFB9FE445BBFFD12F8C9" authority="Nagy &amp; Dózsa-Farkas &amp; Felföldi, 2023" authorityName="Nagy &amp; Dózsa-Farkas &amp; Felföldi" authorityYear="2023" box="[454,748,1787,1813]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
<emphasis id="028AEA9EFFD4FFB9FE445BBFFD12F8C9" box="[454,748,1787,1813]" italics="true" pageId="11" pageNumber="12">Enchytraeus adrianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B957E5FFD4FFB9FD765BB8FCB1F8CA" box="[756,847,1788,1814]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="028AEA9EFFD4FFB9FCD65BBFFC27F8C9" bold="true" box="[852,985,1787,1813]" pageId="11" pageNumber="12">A, C, G–H</emphasis>
. In vivo. 
<emphasis id="028AEA9EFFD4FFB9FBC95BBFFB45F8C9" bold="true" box="[1099,1211,1787,1813]" pageId="11" pageNumber="12">B, D–F, I</emphasis>
. Fixed, stained. 
<emphasis id="028AEA9EFFD4FFB9FF3F5A5AFF2CF8E4" bold="true" box="[189,210,1822,1848]" pageId="11" pageNumber="12">A</emphasis>
. Sperm funnels. 
<emphasis id="028AEA9EFFD4FFB9FE215A5AFE49F8E4" bold="true" box="[419,439,1822,1848]" pageId="11" pageNumber="12">B</emphasis>
. Paratype (slide 2748, P. 143.3, ELTE), sperm ducts (slender part marked with white arrow, wider part with black arrows). 
<emphasis id="028AEA9EFFD4FFB9FD3C5A05FD0AF887" bold="true" box="[702,756,1857,1883]" pageId="11" pageNumber="12">C–F</emphasis>
. Multiple male glands (E–F primary bulb marked with arrow). 
<emphasis id="028AEA9EFFD4FFB9FE985A21FED1F8A3" bold="true" box="[282,303,1893,1919]" pageId="11" pageNumber="12">D</emphasis>
. Paratype (slide 2748, P.143.3, ELTE). 
<emphasis id="028AEA9EFFD4FFB9FC805A21FCEBF8A3" bold="true" box="[770,789,1893,1919]" pageId="11" pageNumber="12">E</emphasis>
. Paratype (slide 2748, P.143.3, ELTE). 
<emphasis id="028AEA9EFFD4FFB9FB6B5A21FB05F8A3" bold="true" box="[1257,1275,1893,1919]" pageId="11" pageNumber="12">F</emphasis>
. Holotype (En.2. slide 2842, ELTE). 
<emphasis id="028AEA9EFFD4FFB9FD835ACCFDF2F87E" bold="true" box="[513,524,1928,1954]" pageId="11" pageNumber="12">I</emphasis>
. Paratype (slide 2827, P. 143.8, ELTE). 
<emphasis id="028AEA9EFFD4FFB9FB835ACCFBEDF87E" bold="true" box="[1025,1043,1928,1954]" pageId="11" pageNumber="12">F</emphasis>
. Holotype (En.2. slide 2842, ELTE). 
<emphasis id="028AEA9EFFD4FFB9FE995AEFFEB2F819" bold="true" box="[283,332,1963,1989]" pageId="11" pageNumber="12">G–I</emphasis>
. Spermathecae (ectal duct covered with glands marked with white arrows, ampullae filled with sperm in it marked with black arrows).Scale bars = 50 μm.
</paragraph>
</caption>
<paragraph id="3041368CFFD3FFBEFF3F5CD5FB39FE77" blockId="12.[189,1223,401,427]" box="[189,1223,401,427]" pageId="12" pageNumber="13">
For similarities and differences of the species of 
<taxonomicName id="F7FE4D0FFFD3FFBEFD7A5CD5FC72FE77" authorityName="Henle" authorityYear="1837" box="[760,908,401,427]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="12" pageNumber="13" phylum="Annelida" rank="genus">
<emphasis id="028AEA9EFFD3FFBEFD7A5CD5FC72FE77" box="[760,908,401,427]" italics="true" pageId="12" pageNumber="13">Enchytraeus</emphasis>
</taxonomicName>
studied by us, see 
<tableCitation id="7D7C0337FFD3FFBEFBE85CD5FB3AFE77" box="[1130,1220,401,427]" captionStart="Table 2" captionStartId="13.[137,207,545,573]" captionTargetPageId="13" captionText="Table 2 (continued on next two pages). Comparison of morphological characters (measurements in fixed specimens). The data are derived from this study and from Erséus et al. (2019), Arslan et al. (2018) and according to the author of original species description (marked with*). The data given by us marked with bold, dimensions in μm." httpUri="http://table.plazi.org/id/64816604FFD2FFBFFF0B5F65FD9FFD5F" pageId="12" pageNumber="13" tableUuid="64816604FFD2FFBFFF0B5F65FD9FFD5F">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78E46507FFD3FFBEFF3F5C96FB44FDC7" pageId="12" pageNumber="13" type="distribution">
<paragraph id="3041368CFFD3FFBEFF3F5C96FE0CFE30" blockId="12.[189,498,466,492]" box="[189,498,466,492]" pageId="12" pageNumber="13">
<heading id="6B0981E0FFD3FFBEFF3F5C96FE0CFE30" bold="true" box="[189,498,466,492]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="028AEA9EFFD3FFBEFF3F5C96FE0CFE30" bold="true" box="[189,498,466,492]" pageId="12" pageNumber="13">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="3041368CFFD3FFBEFF3F5F45FB44FDC7" blockId="12.[189,1210,513,539]" box="[189,1210,513,539]" pageId="12" pageNumber="13">
Kale Cove, Adriatic Sea, Kamenjak Peninsula, 
<collectingRegion id="F23AF86EFFD3FFBEFD6A5F45FCD9FDC7" box="[744,807,513,539]" country="Croatia" name="Istarska" pageId="12" pageNumber="13">Istria</collectingRegion>
, 
<collectingCountry id="48E9761CFFD3FFBEFCB15F45FC72FDC7" box="[819,908,513,539]" name="Croatia" pageId="12" pageNumber="13">Croatia</collectingCountry>
, decaying 
<taxonomicName id="F7FE4D0FFFD3FFBEFB8B5F46FB9AFDC7" box="[1033,1124,514,539]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="028AEA9EFFD3FFBEFB8B5F46FB9AFDC7" box="[1033,1124,514,539]" italics="true" pageId="12" pageNumber="13">Zostera</emphasis>
</taxonomicName>
debris.
</paragraph>
</subSubSection>
</treatment>
</document>