238 lines
27 KiB
XML
238 lines
27 KiB
XML
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<mods:title id="9AACEEE49E2F1C221F4681995FF7CA79">Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals</mods:title>
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<taxonomicName id="2ECF4D5E3F05FF92FF76FF11FD96FF50" ID-CoL="34K5X" authority="(Risso, 1820)" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1820" box="[151,630,151,178]" class="Actinopterygii" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="collonianus">
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<emphasis id="DBBBEACF3F05FF92FF76FF11FE2FFF53" bold="true" box="[151,463,151,178]" italics="true" pageId="48" pageNumber="49">Deltentosteus collonianus</emphasis>
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(
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<bibRefCitation id="8D5E4B2C3F05FF92FE3EFF11FD8EFF50" author="Risso, A." box="[479,622,151,177]" pageId="48" pageNumber="49" pagination="241 - 255" refId="ref63831" refString="Risso, A. (1820) Memoire sur quelques poissons observes dans la mer de Nice. Journal de Physique, de Chimie et d'Histoire Naturelle, 91, 241 - 255." type="journal article" year="1820">Risso, 1820</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="71F42A583F05FF92FD67FF11FD3BFF50" box="[646,731,151,177]" captionStart="FIGURE 51" captionStartId="48.[151,250,1863,1887]" captionTargetBox="[151,1436,448,1839]" captionTargetId="figure-100@48.[151,1436,448,1839]" captionTargetPageId="48" captionText="FIGURE 51. Live coloration in its natural habitat of Deltentosteus collonianus. A) photo by S. Guerrieri, Tuscany, Italy. B) Breeding male (the rounded, notably enlarged first dorsal fin, has been injured), photo by S. Le Bris, Saména, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601727" httpUri="https://zenodo.org/record/6601727/files/figure.png" pageId="48" pageNumber="49">Fig. 51</figureCitation>
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)—Toothed Goby
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="A1D565563F05FF92FF76FF5DFD07FEDC" pageId="48" pageNumber="49" type="reference_group">
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<treatmentCitation id="686E10CC3F05FF92FF76FF5DFDF1FF12" author="Risso, A." box="[151,529,219,244]" page="252" pageId="48" pageNumber="49" year="1820">
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<taxonomicName id="2ECF4D5E3F05FF92FF76FF5DFDF1FF12" ID-CoL="3GPQJ" authority="Risso, 1820: 252" authorityName="Risso" authorityPageNumber="252" authorityYear="1820" box="[151,529,219,244]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="collonianus">
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<emphasis id="DBBBEACF3F05FF92FF76FF5DFEBDFF12" box="[151,349,219,243]" italics="true" pageId="48" pageNumber="49">Gobius collonianus</emphasis>
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<bibRefCitation id="8D5E4B2C3F05FF92FE82FF5DFDF1FF12" author="Risso, A." box="[355,529,219,244]" pageId="48" pageNumber="49" pagination="241 - 255" refId="ref63831" refString="Risso, A. (1820) Memoire sur quelques poissons observes dans la mer de Nice. Journal de Physique, de Chimie et d'Histoire Naturelle, 91, 241 - 255." type="journal article" year="1820">Risso, 1820: 252</bibRefCitation>
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;
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<materialsCitation id="59A73C803F05FF92FDFDFF5AFB61FF12" ID-GBIF-Occurrence="3878848342" box="[540,1153,219,244]" country="France" location="northwestern Mediterranean Sea" municipality="Nice" pageId="48" pageNumber="49" specimenCount="1">
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<typeStatus id="3674887F3F05FF92FDFDFF5AFDA8FF15" box="[540,584,220,244]" pageId="48" pageNumber="49">type</typeStatus>
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locality:
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,
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<collectingCountry id="91D8764D3F05FF92FC1CFF5AFBA5FF12" box="[1021,1093,220,243]" name="France" pageId="48" pageNumber="49">France</collectingCountry>
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,
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<collectingMunicipality id="0914ACA73F05FF92FBAFFF5DFB61FF12" box="[1102,1153,219,243]" pageId="48" pageNumber="49">Nice</collectingMunicipality>
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.
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</paragraph>
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<paragraph id="E97036DD3F05FF92FF76FEA4FD07FEDC" blockId="48.[151,1437,290,424]" box="[151,743,290,317]" pageId="48" pageNumber="49">
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Size. Maximum size
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<quantity id="2E379B383F05FF92FE65FEA4FE5BFEDD" box="[388,443,290,316]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="48" pageNumber="49" unit="cm" value="7.0">7 cm</quantity>
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total length (Miller 1986).
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</paragraph>
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<subSubSection id="A1D565563F05FF93FF26FEC0FBFBFD2F" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
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<paragraph id="E97036DD3F05FF93FF26FEC0FD90FEFC" blockId="48.[151,1437,290,424]" lastBlockId="49.[151,1437,150,718]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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Morphology.
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<emphasis id="DBBBEACF3F05FF92FEBEFEC0FD72FE81" bold="true" box="[351,658,325,352]" pageId="48" pageNumber="49">D VI + I,10–11; A I,10–11;</emphasis>
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P 17 (Miller 1986;
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<bibRefCitation id="8D5E4B2C3F05FF92FC8AFEC0FB68FE80" author="Kovacic, M. & Golani, D." box="[875,1160,326,353]" pageId="48" pageNumber="49" pagination="417 - 425" refId="ref58262" refString="Kovacic, M. & Golani, D. (2007 b) First record of Papillogobius melanobranchus in the Mediterranean Sea and new data on geographic distributions, bathymetric ranges and morphology of several benthic fishes in the Levant. Cybium, 31 (4), 417 - 425. https: // doi. org / 10.26028 / cybium / 2007 - 314 - 002" type="journal article" year="2007">Kovačić & Golani 2007b</bibRefCitation>
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). Moderately small goby with subcylindrical body, laterally compressed towards caudal fin, head slightly depressed and moderately large. Snout large and pointed. Eyes lateral, interorbital moderately wide. Mouth superior, oblique, tip of lower jaw at horizontal level of pupil.
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<emphasis id="DBBBEACF3F04FF93FE53FF10FC96FF51" bold="true" box="[434,886,150,177]" pageId="49" pageNumber="50">Angle of jaws below rear half of orbit.</emphasis>
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Predorsal area and nape scaled. Caudal peduncle slender, lower than body. The first dorsal fin higher than the second dorsal fin, no elongated spines, but
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<emphasis id="DBBBEACF3F04FF93FAD2FF3CFCDCFF19" bold="true" pageId="49" pageNumber="50">breeding males with notably enlarged, sail-shaped, first dorsal fin</emphasis>
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(
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||
<figureCitation id="71F42A583F04FF93FCADFF59FC4CFF18" box="[844,940,222,249]" captionStart="FIGURE 51" captionStartId="48.[151,250,1863,1887]" captionTargetBox="[151,1436,448,1839]" captionTargetId="figure-100@48.[151,1436,448,1839]" captionTargetPageId="48" captionText="FIGURE 51. Live coloration in its natural habitat of Deltentosteus collonianus. A) photo by S. Guerrieri, Tuscany, Italy. B) Breeding male (the rounded, notably enlarged first dorsal fin, has been injured), photo by S. Le Bris, Saména, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601727" httpUri="https://zenodo.org/record/6601727/files/figure.png" pageId="49" pageNumber="50">Fig. 51b</figureCitation>
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) (
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<bibRefCitation id="8D5E4B2C3F04FF93FC25FF59FBB6FF18" author="Louisy, P." box="[964,1110,222,249]" pageId="49" pageNumber="50" refId="ref60639" refString="Louisy, P. (2015) Europe and Mediterranean Marine Fish Identification Guide. 3 rd Edition. Eugen Ulmer Ed., Paris, 512 pp." type="book" year="2015">Louisy 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FB82FF59FADBFF18" author="Le Bris, S. & Berenger, L. & Louisy, P." box="[1123,1339,223,249]" pageId="49" pageNumber="50" refId="ref59988" refString="Le Bris, S., Berenger, L. & Louisy, P. (2017) Signalisation de / Record of Deltentosteus collonianus, 12 March 2017 (obs. 31 January 2017, photo # 3514). Fish Watch Forum, Louisy, P. & Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2017">
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||
Le Bris
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||
<emphasis id="DBBBEACF3F04FF93FB5EFF59FB11FF18" box="[1215,1265,223,249]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
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. 2017
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||
</bibRefCitation>
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||
;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FAA6FF59FA71FF18" author="Le Bris, S. & Pillon, R. & Louisy, P." box="[1351,1425,223,249]" pageId="49" pageNumber="50" refId="ref60064" refString="Le Bris, S., Pillon, R. & Louisy, P. (2019 a) Signalisation de / Record of Deltentosteus collonianus, 02 March 2019 (obs. 27 February 2019, photo # 5152). Fish Watch Forum, Louisy, P. & Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2019">2019a</bibRefCitation>
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||
). Caudal fin truncate to slightly emarginate.
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</paragraph>
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<caption id="BDB066553F05FF92FF76F8C1FF03F847" ID-DOI="http://doi.org/10.5281/zenodo.6601727" ID-Zenodo-Dep="6601727" httpUri="https://zenodo.org/record/6601727/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1863,1887]" targetBox="[151,1436,448,1839]" targetPageId="48">
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<paragraph id="E97036DD3F05FF92FF76F8C1FF03F847" blockId="48.[151,1437,1863,1958]" pageId="48" pageNumber="49">
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<emphasis id="DBBBEACF3F05FF92FF76F8C1FEC4F8BE" bold="true" box="[151,292,1863,1887]" pageId="48" pageNumber="49">FIGURE 51.</emphasis>
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Live coloration in its natural habitat of
|
||
<taxonomicName id="2ECF4D5E3F05FF92FD34F8C1FC3FF8BE" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1820" box="[725,991,1863,1887]" class="Actinopterygii" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="collonianus">
|
||
<emphasis id="DBBBEACF3F05FF92FD34F8C1FC3FF8BE" box="[725,991,1863,1887]" italics="true" pageId="48" pageNumber="49">Deltentosteus collonianus</emphasis>
|
||
</taxonomicName>
|
||
. A) photo by S. Guerrieri, Tuscany, Italy. B) Breeding male (the rounded, notably enlarged first dorsal fin, has been injured), photo by S. Le Bris,
|
||
<emphasis id="DBBBEACF3F05FF92FB03F8EDFAD8F863" box="[1250,1336,1899,1922]" italics="true" pageId="48" pageNumber="49">Saména,</emphasis>
|
||
Southern France.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E97036DD3F04FF93FF26FEA1FBB2FE14" blockId="49.[151,1437,150,718]" pageId="49" pageNumber="50">
|
||
Live coloration. Body dorsally mainly brown or beige with
|
||
<emphasis id="DBBBEACF3F04FF93FC90FEA1FEC4FE84" bold="true" pageId="49" pageNumber="50">5 pale dorsal saddles, each above a midlateral dark blotch</emphasis>
|
||
including one at the caudal-fin base (
|
||
<figureCitation id="71F42A583F04FF93FD2FFECDFCC2FE84" box="[718,802,330,357]" captionStart="FIGURE 51" captionStartId="48.[151,250,1863,1887]" captionTargetBox="[151,1436,448,1839]" captionTargetId="figure-100@48.[151,1436,448,1839]" captionTargetPageId="48" captionText="FIGURE 51. Live coloration in its natural habitat of Deltentosteus collonianus. A) photo by S. Guerrieri, Tuscany, Italy. B) Breeding male (the rounded, notably enlarged first dorsal fin, has been injured), photo by S. Le Bris, Saména, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601727" httpUri="https://zenodo.org/record/6601727/files/figure.png" pageId="49" pageNumber="50">Fig. 51</figureCitation>
|
||
).
|
||
<emphasis id="DBBBEACF3F04FF93FCD6FECDFEA7FE4C" bold="true" pageId="49" pageNumber="50">The saddles on the back generally conspicuous and usually with black anterior and posterior outlines. The midlateral dark blotches of irregular rhomboid to rounded shape</emphasis>
|
||
, anteriorly about 1/3 of body height.
|
||
<emphasis id="DBBBEACF3F04FF93FD0AFE15FE57FE30" bold="true" pageId="49" pageNumber="50">Black mark on caudal-fin base triangular and larger than other midlateral marks.</emphasis>
|
||
Ventral half of body whitish.
|
||
<emphasis id="DBBBEACF3F04FF93FCEFFE31FA74FE31" bold="true" box="[782,1428,438,465]" pageId="49" pageNumber="50">The first dorsal fin without dark markings in females</emphasis>
|
||
; one or two black spots on rear margin in males (
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FD53FE5DFC68FE14" author="Le Bris, S. & Berenger, L. & Louisy, P." box="[690,904,475,501]" pageId="49" pageNumber="50" refId="ref59988" refString="Le Bris, S., Berenger, L. & Louisy, P. (2017) Signalisation de / Record of Deltentosteus collonianus, 12 March 2017 (obs. 31 January 2017, photo # 3514). Fish Watch Forum, Louisy, P. & Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2017">
|
||
Le Bris
|
||
<emphasis id="DBBBEACF3F04FF93FCEFFE5DFCDFFE14" box="[782,831,475,501]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FC75FE5DFC3AFE14" author="Le Bris, S. & Pillon, R. & Louisy, P." box="[916,986,475,501]" pageId="49" pageNumber="50" refId="ref60064" refString="Le Bris, S., Pillon, R. & Louisy, P. (2019 a) Signalisation de / Record of Deltentosteus collonianus, 02 March 2019 (obs. 27 February 2019, photo # 5152). Fish Watch Forum, Louisy, P. & Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2019">2019a</bibRefCitation>
|
||
) (
|
||
<figureCitation id="71F42A583F04FF93FC10FE5DFBA4FE14" box="[1009,1092,474,501]" captionStart="FIGURE 51" captionStartId="48.[151,250,1863,1887]" captionTargetBox="[151,1436,448,1839]" captionTargetId="figure-100@48.[151,1436,448,1839]" captionTargetPageId="48" captionText="FIGURE 51. Live coloration in its natural habitat of Deltentosteus collonianus. A) photo by S. Guerrieri, Tuscany, Italy. B) Breeding male (the rounded, notably enlarged first dorsal fin, has been injured), photo by S. Le Bris, Saména, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601727" httpUri="https://zenodo.org/record/6601727/files/figure.png" pageId="49" pageNumber="50">Fig. 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F04FF93FF26FE79FBFAFDF8" blockId="49.[151,1437,150,718]" box="[199,1050,511,537]" pageId="49" pageNumber="50">
|
||
Similar species.
|
||
<taxonomicName id="2ECF4D5E3F04FF93FE9EFE79FD3DFDF8" authorityName="Benthic" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[383,733,511,537]" class="Actinopterygii" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="quadrimaculatus">
|
||
<emphasis id="DBBBEACF3F04FF93FE9EFE79FD3DFDF8" box="[383,733,511,537]" italics="true" pageId="49" pageNumber="50">Deltentosteus quadrimaculatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2ECF4D5E3F04FF93FD0AFE79FBF3FDF8" authorityName="Miller" authorityYear="1973" box="[747,1043,511,537]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="adriaticus">
|
||
<emphasis id="DBBBEACF3F04FF93FD0AFE79FBF3FDF8" box="[747,1043,511,537]" italics="true" pageId="49" pageNumber="50">Pomatoschistus adriaticus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F04FF93FF26FDA5FC48FD80" blockId="49.[151,1437,150,718]" pageId="49" pageNumber="50">
|
||
Habitat. Infralittoral to circalittoral species, known from
|
||
<quantity id="2E379B383F04FF93FCB6FDA4FC32FDDC" box="[855,978,546,573]" metricMagnitude="1" metricUnit="m" metricValue="6.075" metricValueMax="12.0" metricValueMin="0.15" pageId="49" pageNumber="50" unit="m" value="60.75" valueMax="120.0" valueMin="1.5">1.5–120 m</quantity>
|
||
on soft bottoms and seagrass meadows (Miller 1986;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FED5FDC1FDE8FD80" author="Le Bris, S. & Berenger, L. & Louisy, P." box="[308,520,583,609]" pageId="49" pageNumber="50" refId="ref59988" refString="Le Bris, S., Berenger, L. & Louisy, P. (2017) Signalisation de / Record of Deltentosteus collonianus, 12 March 2017 (obs. 31 January 2017, photo # 3514). Fish Watch Forum, Louisy, P. & Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2017">
|
||
Le Bris
|
||
<emphasis id="DBBBEACF3F04FF93FE6FFDC1FE20FD80" box="[398,448,583,609]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FDF5FDC1FD19FD80" author="Iglesias, S. P. & Bariche, M. & Beau, F. & Berenger, L. & Beucher, R. & Chabrolle, A. & Cottalorda, J. M. & Cousin, B. & Curd, A. & Danet, V. & Duhamel, E. & Duval, A. & Farque, P. A. & Goascoz, N. & Jadaud, A. & Larnaud, P. & Le Bouter, M. & Le Bras, Y. & Le Bris, S. & Lombard, L. & Louisy, P. & Mandine, M. & Mas, L. & Menut, T. & Metral, L. & Poussard, P. & Quero, J. C. & Raybaud, V. & Renoult, J. P. & Richard, T. & Spitz, J. & Ternon, Q. & Thiriet, P. & Tournier-Broer, R." box="[532,761,583,609]" pageId="49" pageNumber="50" pagination="169 - 188" refId="ref56818" refString="Iglesias, S. P., Bariche, M., Beau, F., Berenger, L., Beucher, R., Chabrolle, A., Cottalorda, J. M., Cousin, B., Curd, A., Danet, V., Duhamel, E., Duval, A., Farque, P. A., Goascoz, N., Jadaud, A., Larnaud, P., Le Bouter, M., Le Bras, Y., Le Bris, S., Lombard, L., Louisy, P., Mandine, M., Mas, L., Menut, T., Metral, L., Poussard, P., Quero, J. C., Raybaud, V., Renoult, J. P., Richard, T., Spitz, J., Ternon, Q., Thiriet, P. & Tournier-Broer, R. (2021 b) French Ichthyological Records for 2019. Cybium, 45 (3), 169 - 188. https: // doi. org / 10.26028 / cybium / 2020 - 444 - 001" type="journal article" year="2021">
|
||
Iglésias
|
||
<emphasis id="DBBBEACF3F04FF93FD90FDC1FD42FD80" box="[625,674,583,609]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
||
. 2021b
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FCE4FDC1FC7CFD80" author="Patzner, R. A." box="[773,924,583,609]" pageId="49" pageNumber="50" refId="ref62855" refString="Patzner, R. A. (2021) Gobiidae. Available from: http: // www. patzner. sbg. ac. at / Gobiidae / GobiidaeSpecies. htm (accessed 1 October 2021)" type="url" year="2021">Patzner 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F04FF93FF26FDEDFBFBFD2F" blockId="49.[151,1437,150,718]" pageId="49" pageNumber="50">
|
||
Geographic distribution. Mediterranean and adjacent Atlantic Ocean.
|
||
<taxonomicName id="2ECF4D5E3F04FF93FC31FDEDFB95FD64" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1820" box="[976,1141,619,645]" class="Actinopterygii" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="collonianus">
|
||
<emphasis id="DBBBEACF3F04FF93FC31FDEDFC04FD65" box="[976,996,619,644]" italics="true" pageId="49" pageNumber="50">D</emphasis>
|
||
.
|
||
<emphasis id="DBBBEACF3F04FF93FC13FDEDFB95FD64" box="[1010,1141,619,645]" italics="true" pageId="49" pageNumber="50">collonianus</emphasis>
|
||
</taxonomicName>
|
||
is known from the eastern Atlantic coast of
|
||
<collectingCountry id="91D8764D3F04FF93FEBAFD09FE5AFD48" box="[347,442,655,681]" name="Portugal" pageId="49" pageNumber="50">Portugal</collectingCountry>
|
||
(Miller 1986). In the Mediterranean, it is known from the western basin to the Adriatic Sea (Miller 1986). It was later reported from Levant (
|
||
<bibRefCitation id="8D5E4B2C3F04FF93FD0FFD35FBEFFD2F" author="Kovacic, M. & Golani, D." box="[750,1039,691,718]" pageId="49" pageNumber="50" pagination="417 - 425" refId="ref58262" refString="Kovacic, M. & Golani, D. (2007 b) First record of Papillogobius melanobranchus in the Mediterranean Sea and new data on geographic distributions, bathymetric ranges and morphology of several benthic fishes in the Levant. Cybium, 31 (4), 417 - 425. https: // doi. org / 10.26028 / cybium / 2007 - 314 - 002" type="journal article" year="2007">Kovačić & Golani 2007b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |