126 lines
18 KiB
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126 lines
18 KiB
XML
<document id="B89E6FF7C9380A00E70160909B581F0F" ID-CLB-Dataset="46266" ID-DOI="10.5281/zenodo.278832" ID-GBIF-Dataset="9d274a58-46e3-471c-8945-bbf84a9b1ba1" ID-ISSN="1175-5326" ID-Zenodo-Dep="278832" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393709508" checkinUser="plazi" docAuthor="Mahony, Stephen" docDate="2011" docId="485787BFFFB9C326FF0BFD44FA02F8AB" docLanguage="en" docName="zt03042p067.pdf" docOrigin="Zootaxa 3042" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hemidactylus Brookii Gray 1845" docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="B46EFFC7FFBDC322FF9CFFCBFF87FFA9" masterDocTitle="Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)" masterLastPageNumber="67" masterPageNumber="37" pageNumber="41" updateTime="1698298507063" updateUser="ExternalLinkService">
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<mods:title id="3DE5B49ACD0796A34D0E050CD2BA7AAB">Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)</mods:title>
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<mods:namePart id="AF5CD42AEC79604B0F5CBBF2B2CA81F6">Mahony, Stephen</mods:namePart>
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<mods:typeOfResource id="DC888271360E4F7BAF691550C101E7D5">text</mods:typeOfResource>
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<mods:title id="EE0EDA299F05CF290BB76D27DDCACF12">Zootaxa</mods:title>
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<mods:part id="F0D3D52D0174B7BF261CD0217F80B083">
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<mods:date id="804153AD49A58E578E8E9710474D9F38">2011</mods:date>
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<mods:number id="CE468244AF937F29A6D9B7D112D1B755">3042</mods:number>
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<mods:classification id="504E86175BD613D2BB3DD9817A264528">journal article</mods:classification>
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<mods:identifier id="0740C44E05AA129D1B1259D31A7FC5D6" type="CLB-Dataset">46266</mods:identifier>
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<mods:identifier id="BB1E5294B82732B1429A668B98079DE2" type="DOI">10.5281/zenodo.278832</mods:identifier>
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<treatment id="485787BFFFB9C326FF0BFD44FA02F8AB" ID-DOI="http://doi.org/10.5281/zenodo.5621712" ID-GBIF-Taxon="119417558" ID-Zenodo-Dep="5621712" LSID="urn:lsid:plazi:treatment:485787BFFFB9C326FF0BFD44FA02F8AB" httpUri="http://treatment.plazi.org/id/485787BFFFB9C326FF0BFD44FA02F8AB" lastPageNumber="41" pageId="4" pageNumber="41">
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<subSubSection id="88E46522FFB9C326FF0BFD44FD41FD00" box="[151,710,655,681]" pageId="4" pageNumber="41" type="nomenclature">
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<paragraph id="C04136A9FFB9C326FF0BFD44FD41FD00" blockId="4.[151,710,655,681]" box="[151,710,655,681]" pageId="4" pageNumber="41">
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<heading id="9B0981C5FFB9C326FF0BFD44FD41FD00" bold="true" box="[151,710,655,681]" fontSize="11" level="1" pageId="4" pageNumber="41" reason="1">
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<emphasis id="F28AEABBFFB9C326FF0BFD44FD41FD00" bold="true" box="[151,710,655,681]" pageId="4" pageNumber="41">
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<typeStatus id="1F45880BFFB9C326FF0BFD44FE80FD00" box="[151,263,655,681]" pageId="4" pageNumber="41" type="syntype">Syntypes</typeStatus>
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of
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<taxonomicName id="07FE4D2AFFB9C326FEB2FD44FD41FD00" authority="Brookii Gray, 1845" authorityName="Brookii Gray" authorityYear="1845" box="[302,710,655,681]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="F28AEABBFFB9C326FEB2FD44FDB1FD00" bold="true" box="[302,566,655,681]" italics="true" pageId="4" pageNumber="41">Hemidactylus Brookii</emphasis>
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<bibRefCitation id="A46F4B58FFB9C326FDA2FD44FD41FD00" author="Gray" box="[574,710,655,681]" pageId="4" pageNumber="41" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees of the British Museum, London. xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="88E46522FFB9C326FF0BFD1DFA02F8AB" pageId="4" pageNumber="41" type="description">
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<paragraph id="C04136A9FFB9C326FF0BFD1DFE7BFA17" blockId="4.[151,1437,726,1795]" pageId="4" pageNumber="41">
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The species
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<taxonomicName id="07FE4D2AFFB9C326FEBCFD1DFD97FD47" box="[288,528,726,750]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFB9C326FEBCFD1DFD97FD47" box="[288,528,726,750]" italics="true" pageId="4" pageNumber="41">Hemidactylus brookii</emphasis>
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</taxonomicName>
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was originally described based on three specimens in the collection of the British Museum of Natural History (BMNH). The specimens were reported to originate from Borneo (two) and
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<collectingCountry id="B8E97639FFB9C326FAA9FD31FA1BFCBB" box="[1333,1436,762,786]" name="Australia" pageId="4" pageNumber="41">Australia</collectingCountry>
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(one), the latter locality doubted by several subsequent authors (e.g.,
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<bibRefCitation id="A46F4B58FFB9C326FC23FCD5FBBBFC9F" author="Gray" box="[959,1084,798,822]" pageId="4" pageNumber="41" refString="Gray, J. E. (1867) The Lizards of Australia and New Zealand in the Collection of the British Museum. (With 18 Plates of the New Species, by Mr. Ford). Bernard Quaritch, London, 8 pp. + Pl. I - XX." type="book" year="1867">Gray 1867</bibRefCitation>
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;
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<bibRefCitation id="A46F4B58FFB9C326FBD0FCD5FA83FC9F" author="Loveridge" box="[1100,1284,798,822]" pageId="4" pageNumber="41" refString="Loveridge, A. (1941) Certain Afro-American geckos of the genus Hemidactylus. Copeia, 4, 245 - 248." type="journal article" year="1941">Loveridge 1941</bibRefCitation>
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;
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<bibRefCitation id="A46F4B58FFB9C326FA8FFCD4FF66FCF3" author="Bauer" pageId="4" pageNumber="41" refString="Bauer, A. M., Jackman, T. R., Greenbaum, E., de Silva, A., Giri, V. B. & Das, I. (2010 b) Molecular evidence for the taxonomic status of Hemidactylus brookii group taxa (Squamata: Gekkonidae). Herpetological Journal, 20, 129 - 138." type="journal article" year="2010" yearSuffix="b">
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Bauer
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<emphasis id="F28AEABBFFB9C326FAFCFCEBFA1BFC9F" box="[1376,1436,798,823]" italics="true" pageId="4" pageNumber="41">et al.</emphasis>
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2010b
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</bibRefCitation>
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). Upon examination of these three specimens it becomes apparent that two morphologically distinct forms are represented in the
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<typeStatus id="1F45880BFFB9C326FE12FCACFE39FCD6" box="[398,446,871,895]" pageId="4" pageNumber="41">type</typeStatus>
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series, reinforcing the original statement of two geographic populations. The two specimens, BMNH 1947.3.6.47 (formerly BM RR1934.9.1.49[.21.a]) from “Borneo”, and BMNH 1947.3.6.49 (formerly BM RR1934.9.1.51[.21.b]) from “
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<collectingCountry id="B8E97639FFB9C326FDFCFC65FD4BFC6F" box="[608,716,942,966]" name="Australia" pageId="4" pageNumber="41">Australia</collectingCountry>
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” form a morphologically unique pair, which considerably differ from the third specimen BMNH 1947.3.6.48 (formerly BM RR1934.9.1.50[.21.b]) from “Borneo”. There are numerous possible hypothesis for the above circumstances, but the most obvious explanation is a simple case of mistagging of specimens. All three specimens are currently maintained in their original separate jars and are individually tagged with the current 1947 labels only. There is no evidence to suggest that these specimens were individually tagged with the old (1934) labels, as the specimens do not have constriction “damage” indicative of multiple specimen tags that may have been removed. It is safe to assume that prior to 1947, the specimens were not individually tagged. Each specimens identity at that time therefore relied on which jar the untagged specimen was placed after examination by scientists. Under this circumstance, the likelihood that two specimens were misplaced in each other’s jars, is very high. In 1947, the specimens were directly tagged with details corresponding to the label on their individual jars, and at this time one Borneo specimen was in the
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<collectingCountry id="B8E97639FFB9C326FBB7FADDFB15FA87" box="[1067,1170,1302,1326]" name="Australia" pageId="4" pageNumber="41">Australia</collectingCountry>
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specimen jar, and vice versa, leading to the current confusion. In this case the morphologically distinct form would represent the single specimen originally labelled from “
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<collectingCountry id="B8E97639FFB9C326FDB4FA95FD14FADF" box="[552,659,1374,1398]" name="Australia" pageId="4" pageNumber="41">Australia</collectingCountry>
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” and the remaining two specimens would represent the Borneo species. A remark regarding this problem has been added to the BMNH specimen register upon recent notification (Patrick Campbell
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<emphasis id="F28AEABBFFB9C326FEF6FA63FE2AFA17" box="[362,429,1446,1471]" italics="true" pageId="4" pageNumber="41">in litt.</emphasis>
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2011).
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</paragraph>
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<paragraph id="C04136A9FFB9C326FF59FA00FA02F8AB" blockId="4.[151,1437,726,1795]" pageId="4" pageNumber="41">
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In the original description,
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<bibRefCitation id="A46F4B58FFB9C326FE69FA01FDF9FA4B" author="Gray" box="[501,638,1482,1506]" pageId="4" pageNumber="41" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees of the British Museum, London. xxviii + 289 pp." type="book" year="1845">Gray (1845)</bibRefCitation>
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neither designates a
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<typeStatus id="1F45880BFFB9C326FCF5FA01FC4CFA4B" box="[873,971,1482,1506]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
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nor does he describe any one of the three specimens in more detail than the others.
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<bibRefCitation id="A46F4B58FFB9C326FDEEFA25FCBEF9AF" author="Boulenger" box="[626,825,1518,1542]" pageId="4" pageNumber="41" refString="Boulenger, G. A. (1898) A list of reptiles and batrachians from Ombaai, East Indian archipelago, Annals and Magazine of Natural History, 7 (1), 122 - 124." type="journal article" year="1898">Boulenger (1898)</bibRefCitation>
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was the first, without explanation, to restrict the
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<typeStatus id="1F45880BFFB9C326FAF7FA24FA1CF9AE" box="[1387,1435,1519,1543]" pageId="4" pageNumber="41">type</typeStatus>
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locality to Borneo (followed by, but often incorrectly attributed to
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<bibRefCitation id="A46F4B58FFB9C326FC18F9D9FB8EF983" author="Smith" box="[900,1033,1554,1578]" pageId="4" pageNumber="41" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II. - Sauria. Taylor and Francis, London. xiii + 440 pp., 2 folding maps, 1 pl." type="book" year="1935">Smith 1935</bibRefCitation>
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), either as an error, or intentionally. Regardless, the species represented by two specimens (BMNH 1947.3.6.47 and BMNH 1947.3.6.49) from Borneo is herein redescribed and retained as the material representing
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<emphasis id="F28AEABBFFB9C326FCC0F990FBFEF9DA" box="[860,1145,1627,1651]" italics="true" pageId="4" pageNumber="41">
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<taxonomicName id="07FE4D2AFFB9C326FCC0F990FBC9F9DA" box="[860,1102,1627,1651]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="species" sensu="stricto" species="brookii">Hemidactylus brookii</taxonomicName>
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<taxonomicNameLabel id="E9B957C0FFB9C326FBC9F997FBFEF9DA" box="[1109,1145,1628,1651]" pageId="4" pageNumber="41" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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According to the BMNH specimen register and comparison with the
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<typeStatus id="1F45880BFFB9C326FDE2F9B4FD64F93E" box="[638,739,1663,1687]" pageId="4" pageNumber="41" type="syntype">syntypes</typeStatus>
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, BMNH 1947.3.6.47 represents the specimen figured by
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<bibRefCitation id="A46F4B58FFB9C326FAF8F9B5FF5BF913" author="Gray" pageId="4" pageNumber="41" refString="Gray, J. E. (1867) The Lizards of Australia and New Zealand in the Collection of the British Museum. (With 18 Plates of the New Species, by Mr. Ford). Bernard Quaritch, London, 8 pp. + Pl. I - XX." type="book" year="1867">Gray (1867</bibRefCitation>
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:Pl. 15,
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<figureCitation id="58C52A2CFFB9C326FEAAF969FEF9F913" box="[310,382,1698,1722]" captionStart="FIGURE 2. A" captionStartId="12.[151,250,1676,1699]" captionTargetBox="[151,1436,337,1655]" captionTargetId="figure@12.[151,1436,337,1655]" captionTargetPageId="12" captionText="FIGURE 2. A. Hemidactylus brookii adult male lectotype (BMNH 1947.3.6.47) by present designation; B. H. gleadowi adult male neotype (BMNH [18] 84.7.25.8) by present designation; C. H. subtriedroides adult male lectotype (BMNH 1946.8.25.54) by present designation; D. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8) by present designation. E & F. Ventral view of the original tails showing the difference of subcaudal proportions and tail shape between the “ litter-dwelling ” and “ rock-dwelling ” morphotypes: E. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8); F. H. tenkatei adult male (BMNH 1926.10.30.47) from Timor." httpUri="https://zenodo.org/record/278834/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
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; 1875:Pl. 15,
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<figureCitation id="58C52A2CFFB9C326FDB8F969FDEFF913" box="[548,616,1698,1722]" captionStart="FIGURE 2. A" captionStartId="12.[151,250,1676,1699]" captionTargetBox="[151,1436,337,1655]" captionTargetId="figure@12.[151,1436,337,1655]" captionTargetPageId="12" captionText="FIGURE 2. A. Hemidactylus brookii adult male lectotype (BMNH 1947.3.6.47) by present designation; B. H. gleadowi adult male neotype (BMNH [18] 84.7.25.8) by present designation; C. H. subtriedroides adult male lectotype (BMNH 1946.8.25.54) by present designation; D. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8) by present designation. E & F. Ventral view of the original tails showing the difference of subcaudal proportions and tail shape between the “ litter-dwelling ” and “ rock-dwelling ” morphotypes: E. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8); F. H. tenkatei adult male (BMNH 1926.10.30.47) from Timor." httpUri="https://zenodo.org/record/278834/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
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–– see here
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<figureCitation id="58C52A2CFFB9C326FD65F969FCBEF913" box="[761,825,1698,1722]" captionStart="FIGURE 1" captionStartId="5.[151,250,1169,1192]" captionTargetBox="[151,1436,447,1147]" captionTargetId="figure@5.[151,1436,447,1147]" captionTargetPageId="5" captionText="FIGURE 1. Historical figures of the Hemidactylus brookii group taxa: A. H. brookii adult male lectotype (BMNH 1947.3.6.47) by present designation, from Gray (1867: Pl. 15, Fig. 2), later reprinted in Gray (1875: Pl. 15, Fig. 2); B. H. subtriedroides adult male paralectotype (ZSI 4135) by present designation, from Annandale (1905 b: Pl. II, Fig. 1); C. H. gleadowi not based on the original type series, from Boulenger (1890: 86, Fig. 27), later reprinted as H. brookii (Boulenger 1912: 42, Fig. 14; Smith 1935: 90, Fig. 28); D. H. gleadowi a syntype from Murray (1884 a: Fig. 3)." httpUri="https://zenodo.org/record/278833/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
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A), which is here designated as the
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<typeStatus id="1F45880BFFB9C326FB78F969FACCF913" box="[1252,1355,1698,1722]" pageId="4" pageNumber="41" type="lectotype">lectotype</typeStatus>
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for
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<taxonomicName id="07FE4D2AFFB9C326FA1DF96FFF60F977" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFB9C326FA1DF96FFF60F977" italics="true" pageId="4" pageNumber="41">H. brookii</emphasis>
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</taxonomicName>
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in order to define the morphology of this species. The third specimen (BMNH 1947.3.6.48) from “
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<collectingCountry id="B8E97639FFB9C326FADAF90DFF30F8AB" name="Australia" pageId="4" pageNumber="41">Australia</collectingCountry>
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”, representing a morphologically distinct species of
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<taxonomicName id="07FE4D2AFFB9C326FC9EF920FC1BF8AA" box="[770,924,1771,1795]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="F28AEABBFFB9C326FC9EF920FC1BF8AA" box="[770,924,1771,1795]" italics="true" pageId="4" pageNumber="41">Hemidactylus</emphasis>
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</taxonomicName>
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, is considered conspecific with
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<taxonomicName id="07FE4D2AFFB9C326FA9FF927FAF9F8AA" box="[1283,1406,1771,1795]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFB9C326FA9FF927FAF9F8AA" box="[1283,1406,1771,1795]" italics="true" pageId="4" pageNumber="41">H. tenkatei</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |