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<document id="16794C14E04D874D8958A1106201E7A4" ID-ISSN="1536-9307" ID-ZooBank="73EA0C25-1792-4147-917E-0C9CC1A0C857" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638239876550" checkinUser="felipe" docAuthor="de Armas, Luis F. &amp; Trujillo, Rony E." docDate="2013" docId="443E87F3FFD3FFFBFCFDF92346FD0F4E" docLanguage="en" docName="Euscorpius.172.1-5.pdf" docOrigin="Euscorpius 172" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Centruroides caral Armas et Trujillo 2013, sp. n." docType="treatment" docUuid="F18CC52C-1783-4608-B1D5-5B15CB909DB9" docUuidSource="ZooBank" docVersion="3" masterDocId="B807FF8BFFD1FFFEFF91FFB14366082B" masterDocTitle="A new species of the genus Centruroides Marx, 1890 (Scorpiones: Buthidae) from Guatemala" updateTime="1699434925182" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9ECFBAAB49472461C83BBD02018B9C32">A new species of the genus Centruroides Marx, 1890 (Scorpiones: Buthidae) from Guatemala</mods:title>
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<mods:namePart id="3B09A82DFFE26B67F8BCD6C221B7FCB9">de Armas, Luis F.</mods:namePart>
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<mods:namePart id="36ED55E3B644792598F0B3E61A1EE3DB">Trujillo, Rony E.</mods:namePart>
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<mods:title id="5CBB57102933FB10D71A009AF63BD07F">Euscorpius</mods:title>
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<mods:date id="C1DE9689A5C46F233B18427EFE054694">2013</mods:date>
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<subSubSection id="848D656EFFD3FFFCFCFDF92346150E87" box="[876,1395,1682,1709]" pageId="2" type="nomenclature">
<paragraph id="CC2836E5FFD3FFFCFCFDF92346150E87" blockId="2.[876,1395,1682,1739]" box="[876,1395,1682,1709]" pageId="2">
<heading id="97608189FFD3FFFCFCFDF92346150E87" box="[876,1395,1682,1709]" centered="true" fontSize="10" level="2" pageId="2" reason="4">
<taxonomicName id="0B974D66FFD3FFFCFCFDF92346430E86" authority="Armas et Trujillo" authorityName="Armas et Trujillo" authorityYear="2013" box="[876,1317,1682,1709]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="caral" status="sp. nov.">
<emphasis id="FEE3EAF7FFD3FFFCFCFDF92347290E87" bold="true" box="[876,1103,1682,1708]" italics="true" pageId="2">Centruroides caral</emphasis>
Armas et Trujillo
</taxonomicName>
,
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<taxonomicNameLabel id="E5D0578CFFD3FFFCFAA0F92346150E87" box="[1329,1395,1682,1708]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="848D656EFFD3FFFBFB9BF90246FD0F4E" lastPageId="5" pageId="2" type="description">
<paragraph id="CC2836E5FFD3FFFCFB9BF90247B30EE0" blockId="2.[876,1395,1682,1739]" box="[1034,1237,1715,1739]" pageId="2">
(
<figureCitation id="54AC2A60FFD3FFFCFB83F90247090EE0" box="[1042,1135,1715,1739]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[192,267,999,1023]" captionStartId-1="4.[192,267,1503,1527]" captionStartId-2="5.[192,267,1161,1185]" captionTargetBox-0="[191,1439,191,969]" captionTargetBox-1="[191,1441,191,1473]" captionTargetBox-2="[191,1441,191,1131]" captionTargetId-0="figure-433@3.[192,1438,192,968]" captionTargetId-1="figure-209@4.[192,1440,192,1472]" captionTargetId-2="figure-362@5.[192,1440,192,1131]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1: Centruroides caral sp. n. Male holotype. Habitus: dorsal (A) and ventral (B)." captionText-1="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." captionText-2="Figure 3: Centruroides caral sp. n. Male holotype. A, coxasternal area and pectines; B, sternites VIVII; C-D, metasomal segments IIV, lateral aspect; E, telson, lateral aspect." pageId="2">Fig. 13</figureCitation>
,
<tableCitation id="8115035EFFD3FFFCFBEAF90247AD0EE0" box="[1147,1227,1715,1739]" captionStart="Table 1" captionStartId="6.[192,257,861,885]" captionText="Table 1: Measurements (mm) of the male holotype of Centruroides caral sp. n. D, depth; L, length; W, width." pageId="2">Table 1</tableCitation>
)
</paragraph>
<paragraph id="CC2836E5FFD3FFFCFCD1F95446F90ED6" blockId="2.[832,1439,1765,1819]" box="[832,1439,1765,1789]" pageId="2">
<emphasis id="FEE3EAF7FFD3FFFCFCD1F95446F90ED6" bold="true" box="[832,1439,1765,1789]" pageId="2">http://zoobank.org/urn:lsid:zoobank.org:act:F18CC5</emphasis>
</paragraph>
<paragraph id="CC2836E5FFD3FFFCFC0AF8B246250F30" blockId="2.[832,1439,1765,1819]" box="[923,1347,1795,1819]" pageId="2">
<emphasis id="FEE3EAF7FFD3FFFCFC0AF8B246250F30" bold="true" box="[923,1347,1795,1819]" pageId="2">2C-1783-4608-B1D5-5B15CB909DB9</emphasis>
</paragraph>
<paragraph id="CC2836E5FFD3FFFCFCFEF88B46C60F7A" blockId="2.[831,1440,1849,1904]" box="[879,1440,1849,1873]" pageId="2">
<emphasis id="FEE3EAF7FFD3FFFCFCFEF88B47650F7A" box="[879,1027,1849,1873]" italics="true" pageId="2">Type material</emphasis>
.
<materialsCitation id="7CFF3CB8FFD3FFFCFB80F88846C60F7A" box="[1041,1440,1849,1873]" collectionCode="MHN" pageId="2" specimenCount="1" typeStatus="holotype">
Male
<typeStatus id="132C8847FFD3FFFCFBDFF88847CD0F7A" box="[1102,1195,1849,1873]" pageId="2" type="holotype">holotype</typeStatus>
(
<collectionCode id="AA86AE20FFD3FFFCFB2BF888479B0F7A" box="[1210,1277,1849,1873]" pageId="2">MHN</collectionCode>
), Firmeza farm
</materialsCitation>
</paragraph>
<paragraph id="CC2836E5FFD3FFFDFCAEF8E942630CAC" blockId="2.[831,1440,1849,1904]" lastBlockId="3.[192,801,1074,1221]" lastPageId="3" pageId="2">
(
<quantity id="0B6F9B00FFD3FFFCFCD7F8E940EC0F5B" box="[838,906,1880,1904]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="2" unit="m" value="510.0">510 m</quantity>
a.s.l., 15.40684° N, 88.69590° W),
<materialsCitation id="7CFF3CB8FFD3FFFDFA80F8E942670CAC" collectorName="C. Vazquez Almazan" country="Guatemala" lastPageId="3" location="Izabal Department" pageId="2" specimenCount="1" stateProvince="Izabal">
Sierra
<taxonomicName id="0B974D66FFD3FFFDFAC9F8E942710C61" authority=", Morales" authorityName="Morales" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" lastPageId="3" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="caral">Caral, Morales</taxonomicName>
municipality,
<collectingRegion id="0E53F807FFD2FFFDFE23FB83411D0C61" box="[434,635,1074,1098]" country="Guatemala" name="Izabal" pageId="3">Izabal Department</collectingRegion>
,
<collectingCountry id="B4807675FFD2FFFDFD14FB83419D0C61" box="[645,763,1074,1098]" name="Guatemala" pageId="3">Guatemala</collectingCountry>
, 15 August, 2011, leg.
<collectorName id="61625333FFD2FFFDFE0FFBE041F60C42" box="[414,656,1105,1129]" pageId="3">C. Vazquez Almazán</collectorName>
, at night, in house
</materialsCitation>
.
</paragraph>
<caption id="98E8666DFFD2FFFDFF51FC5647700BD4" box="[192,1046,999,1023]" pageId="3" startId="3.[192,267,999,1023]" targetBox="[191,1439,191,969]" targetPageId="3">
<paragraph id="CC2836E5FFD2FFFDFF51FC5647700BD4" blockId="3.[192,1046,999,1023]" box="[192,1046,999,1023]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFF51FC56424F0BD4" bold="true" box="[192,297,999,1023]" pageId="3">Figure 1:</emphasis>
<taxonomicName id="0B974D66FFD2FFFDFEBEFC5842800BD4" authority="de Armas &amp; Trujillo, 2013" authorityName="de Armas &amp; Trujillo" authorityYear="2013" box="[303,486,1001,1023]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="caral" status="sp. nov.">
<emphasis id="FEE3EAF7FFD2FFFDFEBEFC5842800BD4" box="[303,486,1001,1023]" italics="true" pageId="3">Centruroides caral</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D0578CFFD2FFFDFE7DFC5841440BD4" box="[492,546,1001,1023]" pageId="3" rank="species">
<emphasis id="FEE3EAF7FFD2FFFDFE7DFC58417A0BD5" bold="true" box="[492,540,1001,1022]" pageId="3">sp. n</emphasis>
.
</taxonomicNameLabel>
Male holotype. Habitus: dorsal (A) and ventral (B).
</paragraph>
</caption>
<paragraph id="CC2836E5FFD2FFFDFF61FB3F424C0CEE" blockId="3.[192,801,1074,1221]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFF61FB3F42140C8D" box="[240,370,1166,1190]" italics="true" pageId="3">Distribution</emphasis>
. Only known from the
<typeStatus id="132C8847FFD2FFFDFD05FB3E41A40C8C" box="[660,706,1167,1191]" pageId="3">type</typeStatus>
locality (
<figureCitation id="54AC2A60FFD2FFFDFF59FB1C42780CEE" box="[200,286,1197,1221]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="3">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="CC2836E5FFD2FFFDFF61FB5B414A0D0A" blockId="3.[192,801,1258,1314]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFF61FB5B42040D29" box="[240,354,1258,1282]" italics="true" pageId="3">Etymology</emphasis>
. The specific name is a noun in apposition, taken from the
<typeStatus id="132C8847FFD2FFFDFE31FABB42A80D09" box="[416,462,1290,1314]" pageId="3">type</typeStatus>
locality.
</paragraph>
<paragraph id="CC2836E5FFD2FFFDFF61FAF647D30CEE" blockId="3.[192,802,1350,1895]" lastBlockId="3.[831,1441,1074,1222]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFF61FAF6423A0D75" box="[240,348,1351,1374]" italics="true" pageId="3">Diagnosis</emphasis>
. Male
<quantity id="0B6F9B00FFD2FFFDFE3EFAF741790D75" box="[431,543,1350,1374]" metricMagnitude="-2" metricUnit="m" metricValue="4.435" pageId="3" unit="mm" value="44.35">44.35 mm</quantity>
in total length; female unknown. Base color light yellow; carapace, tergites and pedipalps (except chelae) with distinct dusky marbling throughout; each tergite bearing a narrow yellow median longitudinal line; cheliceral manus with dark brown reticulations all over; pedipalp chelae with faint spots on the external surface only, with dark brown fingers. Carapace finely granular, with moderate granules on the interocular triangle; anteromedian notch V-shaped. Pedipalp manus slightly wider than patella; movable finger with poorly developed basal lobe. Pectines with 1819 teeth; basal plate rectangular, wider than long. Metasoma: segments I IV with dorsolateral carinae, lateral supramedian carinae, ventrolateral carinae and ventral submedian carinae moderately developed, granular to slightly serrate; segments I IV with two pairs of ventrolateral macrochaetae; intercarinal spaces I IV coriaceous; vesicle deeper than wide; subaculear tubercle strong, spine-like, with tip pointing towards the apex of the aculeus. Metasomal segment III length/width ratio 1.19; metasomal segment V: length/width ratio 2.83, length/height ratio 2.70; carapace length/metasomal segment V length ratio 0.78.
</paragraph>
<paragraph id="CC2836E5FFD2FFFDFCFEFB5A47B50D29" blockId="3.[831,1441,1258,1895]" box="[879,1235,1258,1282]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFCFEFB5A47AA0D29" box="[879,1228,1258,1282]" italics="true" pageId="3">
Description of the male
<typeStatus id="132C8847FFD2FFFDFBE1FB5B47AA0D29" box="[1136,1228,1258,1282]" pageId="3" type="holotype">holotype</typeStatus>
</emphasis>
.
</paragraph>
<paragraph id="CC2836E5FFD2FFFDFCFEFAB840980EC6" blockId="3.[831,1441,1258,1895]" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFCFEFAB8408D0D0A" box="[879,1003,1289,1313]" italics="true" pageId="3">Coloration.</emphasis>
Base color light yellow. Carapace with lateral and posterior margins infuscate and distinct dusky marbling throughout (
<figureCitation id="54AC2A60FFD2FFFDFBB8FAF7471B0D75" box="[1065,1149,1350,1374]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="3">Fig. 2B</figureCitation>
). Tergites I VI with dusky band along posterior margins; anterior portions of tergites with diffuse dusky markings; each tergite bearing a narrow yellow median longitudinal line, and lateromarginal whitish spots (
<figureCitation id="54AC2A60FFD2FFFDFBDBFA7047C20DF2" box="[1098,1188,1473,1497]" captionStart="Figure 1" captionStartId="3.[192,267,999,1023]" captionTargetBox="[191,1439,191,969]" captionTargetId="figure-433@3.[192,1438,192,968]" captionTargetPageId="3" captionText="Figure 1: Centruroides caral sp. n. Male holotype. Habitus: dorsal (A) and ventral (B)." pageId="3">Fig. 1A</figureCitation>
). Metasomal segments I IV light yellow; V and telson slightly darker. Cheliceral manus with dark brown reticulations all over (
<figureCitation id="54AC2A60FFD2FFFDFCD7F9AC40FD0E1E" box="[838,923,1565,1589]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="3">Fig. 2B</figureCitation>
). Pedipalps (
<figureCitation id="54AC2A60FFD2FFFDFBB9F9AC47C30E1E" box="[1064,1189,1565,1589]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="3">Figs. 2D F</figureCitation>
) yellow with moderate dusky markings on femur and patella; chelae with faint spots on the external surface only, with dark brown fingers. Legs pale yellow with faint to moderate dusky markings. Venter yellowish with pale brown diffuse markings on apex of coxapophyses I II and sternites IV VII (
<figureCitation id="54AC2A60FFD2FFFDFC0CF96440960EC6" box="[925,1008,1749,1773]" captionStart="Figure 1" captionStartId="3.[192,267,999,1023]" captionTargetBox="[191,1439,191,969]" captionTargetId="figure-433@3.[192,1438,192,968]" captionTargetPageId="3" captionText="Figure 1: Centruroides caral sp. n. Male holotype. Habitus: dorsal (A) and ventral (B)." pageId="3">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="CC2836E5FFD2FFFAFCFEF94541330E83" blockId="3.[831,1441,1258,1895]" lastBlockId="4.[192,802,1588,1918]" lastPageId="4" pageId="3">
<emphasis id="FEE3EAF7FFD2FFFDFCFEF94540BE0F20" box="[879,984,1780,1803]" italics="true" pageId="3">Carapace</emphasis>
(
<figureCitation id="54AC2A60FFD2FFFDFC60F945471F0F27" box="[1009,1145,1780,1804]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="3">Figs. 2B C</figureCitation>
) 1.06 times longer than wide; anteromedian notch V-shaped, reaching the posterior margin level of the second pair of lateral eyes; interocular triangle with moderate granules, the rest finely granular. Anterior median carinae vestigial, with disperse moderate granules; superciliary carinae strong, subgranulose; posterior median carinae moderate, subgranulose, other carinae indistinct.
</paragraph>
<caption id="98E8666DFFD5FFFAFF51FA6E475D0E24" pageId="4" startId="4.[192,267,1503,1527]" targetBox="[191,1441,191,1473]" targetPageId="4">
<paragraph id="CC2836E5FFD5FFFAFF51FA6E475D0E24" blockId="4.[192,1440,1503,1551]" pageId="4">
<emphasis id="FEE3EAF7FFD5FFFAFF51FA6E424F0DDC" bold="true" box="[192,297,1503,1527]" pageId="4">Figure 2:</emphasis>
<taxonomicName id="0B974D66FFD5FFFAFEBEFA5042810DDC" authority="de Armas &amp; Trujillo, 2013" authorityName="de Armas &amp; Trujillo" authorityYear="2013" box="[303,487,1505,1527]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="caral" status="sp. nov.">
<emphasis id="FEE3EAF7FFD5FFFAFEBEFA5042810DDC" box="[303,487,1505,1527]" italics="true" pageId="4">Centruroides caral</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D0578CFFD5FFFAFE7CFA5041450DDC" box="[493,547,1505,1527]" pageId="4" rank="species">
<emphasis id="FEE3EAF7FFD5FFFAFE7CFA50417B0DDD" bold="true" box="[493,541,1505,1526]" pageId="4">sp. n</emphasis>
.
</taxonomicNameLabel>
A, geographical distribution. B F, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; D F, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela.
</paragraph>
</caption>
<paragraph id="CC2836E5FFD5FFFAFF61F91F475D0F55" blockId="4.[192,802,1588,1918]" lastBlockId="4.[831,1440,1588,1918]" pageId="4">
<emphasis id="FEE3EAF7FFD5FFFAFF61F91F423D0EED" box="[240,347,1710,1734]" italics="true" pageId="4">Pedipalps</emphasis>
orthobothriotaxic
<typeStatus id="132C8847FFD5FFFAFDBEF91E41000EEC" box="[559,614,1711,1735]" pageId="4">Type</typeStatus>
A. Femur (
<figureCitation id="54AC2A60FFD5FFFAFD67F91E43830ECE" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="4">Fig. 2D</figureCitation>
): internal surface with large granules, all other surfaces finely granular; dorsointernal, dorsoexternal and ventrointernal carinae moderate, granular; ventroexternal carina strong, serrate. Patella (
<figureCitation id="54AC2A60FFD5FFFAFD9FF89841020F6A" box="[526,612,1833,1857]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="4">Fig. 2E</figureCitation>
): internal surface with few large conical granules; dorsal, external and ventral surfaces finely granular; dorsointernal and dorsoexternal carinae moderate, finely granular; dorsomedian carina weak, granular; ventrointernal carina weak, finely granular. Manus (
<figureCitation id="54AC2A60FFD5FFFAFC6EF9C047350EA2" box="[1023,1107,1649,1673]" captionStart="Figure 2" captionStartId="4.[192,267,1503,1527]" captionTargetBox="[191,1441,191,1473]" captionTargetId="figure-209@4.[192,1440,192,1472]" captionTargetPageId="4" captionText="Figure 2: Centruroides caral sp. n. A, geographical distribution. BF, male holotype: B, partial view of the carapace, dorsal (in 75% ethanol); C, carapace; DF, right pedipalp, dorsal aspect: D, femur; E, patella; F, chela." pageId="4">Fig. 2F</figureCitation>
): oval shaped, 1.1 times wider than patella; intercarinal surfaces almost smooth, dorsomarginal, dorsal secondary and digital carinae weak to moderate, minutely granular; ventroexternal carina strong, almost smooth; other carinae vestigial. Fixed fingers with eight oblique rows of denticles; trichobothrium
<emphasis id="FEE3EAF7FFD5FFFAFC3FF89840A70F6A" bold="true" box="[942,961,1833,1857]" italics="true" pageId="4">et</emphasis>
basal to
<emphasis id="FEE3EAF7FFD5FFFAFBA9F89847350F6A" bold="true" box="[1080,1107,1833,1857]" italics="true" pageId="4">db</emphasis>
. Movable fingers with eight principal oblique rows of denticles, plus a distal short one with four denticles.
</paragraph>
<caption id="98E8666DFFD4FFFBFF51FB3841AC0C92" pageId="5" startId="5.[192,267,1161,1185]" targetBox="[191,1441,191,1131]" targetPageId="5">
<paragraph id="CC2836E5FFD4FFFBFF51FB3841AC0C92" blockId="5.[192,1441,1161,1209]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFF51FB3842540C8A" bold="true" box="[192,306,1161,1185]" pageId="5">Figure 3:</emphasis>
<taxonomicName id="0B974D66FFD4FFFBFEAAFB3A42900C8A" authority="de Armas &amp; Trujillo, 2013" authorityName="de Armas &amp; Trujillo" authorityYear="2013" box="[315,502,1163,1185]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="caral" status="sp. nov.">
<emphasis id="FEE3EAF7FFD4FFFBFEAAFB3A42900C8A" box="[315,502,1163,1185]" italics="true" pageId="5">Centruroides caral</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D0578CFFD4FFFBFE6EFB3A415E0C8A" box="[511,568,1163,1185]" pageId="5" rank="species">
<emphasis id="FEE3EAF7FFD4FFFBFE6EFB3A41540C8B" bold="true" box="[511,562,1163,1184]" pageId="5">sp. n</emphasis>
.
</taxonomicNameLabel>
Male holotype. A, coxasternal area and pectines; B, sternites VI VII; C-D, metasomal segments II V, lateral aspect; E, telson, lateral aspect.
</paragraph>
</caption>
<paragraph id="CC2836E5FFD4FFFBFF61FB5C415A0D08" blockId="5.[192,802,1261,1893]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFF61FB5C422C0D2F" box="[240,330,1261,1284]" italics="true" pageId="5">Pectines</emphasis>
(
<figureCitation id="54AC2A60FFD4FFFBFECBFB5C42C80D2E" box="[346,430,1261,1285]" captionStart="Figure 3" captionStartId="5.[192,267,1161,1185]" captionTargetBox="[191,1441,191,1131]" captionTargetId="figure-362@5.[192,1440,192,1131]" captionTargetPageId="5" captionText="Figure 3: Centruroides caral sp. n. Male holotype. A, coxasternal area and pectines; B, sternites VIVII; C-D, metasomal segments IIV, lateral aspect; E, telson, lateral aspect." pageId="5">Fig. 3A</figureCitation>
) with 19/18 teeth; basal plate rectangular, 1.7 times wider than long.
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFF61FA9841310DDE" blockId="5.[192,802,1261,1893]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFF61FA98420F0D6B" box="[240,361,1321,1344]" italics="true" pageId="5">
<taxonomicName id="0B974D66FFD4FFFBFF61FA9842030D6B" box="[240,357,1321,1344]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
.
</emphasis>
Tergites I VII finely granular, with some larger granules on the posterior one-half of I VI; longitudinal median carina I VII weak to moderate, granular. Tergite VII lateral and submedian carinae moderate, subserrate. Sternites III V smooth; VI VII finely granular; VII lateral and submedian carinae moderate, granular to subserrate (
<figureCitation id="54AC2A60FFD4FFFBFE67FA6C412C0DDE" box="[502,586,1501,1525]" captionStart="Figure 3" captionStartId="5.[192,267,1161,1185]" captionTargetBox="[191,1441,191,1131]" captionTargetId="figure-362@5.[192,1440,192,1131]" captionTargetPageId="5" captionText="Figure 3: Centruroides caral sp. n. Male holotype. A, coxasternal area and pectines; B, sternites VIVII; C-D, metasomal segments IIV, lateral aspect; E, telson, lateral aspect." pageId="5">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFF61FA4D47F30D0F" blockId="5.[192,802,1261,1893]" lastBlockId="5.[831,1440,1261,1408]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFF61FA4D42390E38" box="[240,351,1532,1555]" italics="true" pageId="5">Metasoma</emphasis>
(
<figureCitation id="54AC2A60FFD4FFFBFEE5FA4A42900E38" box="[372,502,1531,1555]" captionStart="Figure 3" captionStartId="5.[192,267,1161,1185]" captionTargetBox="[191,1441,191,1131]" captionTargetId="figure-362@5.[192,1440,192,1131]" captionTargetPageId="5" captionText="Figure 3: Centruroides caral sp. n. Male holotype. A, coxasternal area and pectines; B, sternites VIVII; C-D, metasomal segments IIV, lateral aspect; E, telson, lateral aspect." pageId="5">Figs. 3C E</figureCitation>
). Intercarinal spaces I IV coriaceous, with sparse minute granules. Segments II IV with two pairs of ventrolateral macrochaetae. Segment I, ten carinae; II IV, eight carinae; V with five carinae; on II, the lateral inframedian carina is only represented by three distal granules; I IV with dorsolateral, lateral supramedian, ventrolateral and ventrosubmedian carinae moderately developed, granular to slightly serrate. Segment V: intercarinal spaces finely granulate; dorsolateral, ventrolateral and ventromedian carinae moderate, granular. Vesicle with ventral and lateral surfaces finely granular, deeper than wide; subaculear tubercle strong, spine-like, with tip pointing towards the apex of the aculeus.
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFCFEFA9A476C0DAB" blockId="5.[831,1440,1261,1408]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFCFEFA9A40C70D69" box="[879,929,1323,1346]" italics="true" pageId="5">Legs</emphasis>
with femora finely granular on the prolateral surface. Telotarsi moderately covered by fine setae on the ventral surface.
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFCAEFA1C477E0DE1" blockId="5.[831,1048,1453,1482]" box="[831,1048,1453,1482]" pageId="5">
<emphasis id="FEE3EAF7FFD4FFFBFCAEFA1C477E0DE1" bold="true" box="[831,1048,1453,1482]" pageId="5">Natural History</emphasis>
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFCFEFA4440D60EE2" blockId="5.[831,1441,1525,1894]" pageId="5">
The male
<typeStatus id="132C8847FFD4FFFBFC4AFA44475E0E26" box="[987,1080,1525,1549]" pageId="5" type="holotype">holotype</typeStatus>
was collected at night, inside the Ecological Station of Sierra Caral (
<quantity id="0B6F9B00FFD4FFFBFB41F9A5467C0E07" box="[1232,1306,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="5" unit="m" value="510.0">510 m</quantity>
a.s.l.), in a rainforest area. In the same place was collected a nymph I of
<taxonomicName id="0B974D66FFD4FFFBFCE0F9E240820E40" baseAuthorityName="Latreille" baseAuthorityYear="1804" box="[881,996,1619,1643]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="FEE3EAF7FFD4FFFBFCE0F9E240820E40" box="[881,996,1619,1643]" italics="true" pageId="5">C. gracilis</emphasis>
</taxonomicName>
. In the same mountains, but at a higher altitude (
<quantity id="0B6F9B00FFD4FFFBFC32F9C3408B0EA1" box="[931,1005,1650,1674]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" unit="m" value="850.0">850 m</quantity>
a.s.l., cloud forest) lives
<taxonomicName id="0B974D66FFD4FFFBFA84F9C347D60E82" authority="Trujillo et Armas, 2012" authorityName="Trujillo et Armas" authorityYear="2012" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="5" phylum="Arthropoda" rank="species" species="landelinoi">
<emphasis id="FEE3EAF7FFD4FFFBFA84F9C340CD0E82" italics="true" pageId="5">Diplocentrus landelinoi</emphasis>
Trujillo et Armas, 2012
</taxonomicName>
(
<taxonomicName id="0B974D66FFD4FFFBFB2FF92046360E82" authorityName="Latreille" authorityYear="1802" box="[1214,1360,1681,1705]" class="Arachnida" family="Scorpionidae" kingdom="Animalia" order="Scorpiones" pageId="5" phylum="Arthropoda" rank="family">Scorpionidae</taxonomicName>
: Diplocentrinae).
</paragraph>
<paragraph id="CC2836E5FFD4FFFBFCFEF96146FD0F4E" blockId="5.[831,1441,1525,1894]" pageId="5">
Sierra Caral is the most biodiverse forest remnant in the Caribbean
<collectingCountry id="B4807675FFD4FFFBFC7DF95E47050F2C" box="[1004,1123,1775,1799]" name="Guatemala" pageId="5">Guatemala</collectingCountry>
. As part of the Merendon Range that stretches from
<collectingCountry id="B4807675FFD4FFFBFBF5F8BE47B10F0C" box="[1124,1239,1807,1831]" name="Guatemala" pageId="5">Guatemala</collectingCountry>
to
<collectingCountry id="B4807675FFD4FFFBFA90F8BE460D0F0C" box="[1281,1387,1807,1831]" name="Honduras" pageId="5">Honduras</collectingCountry>
, this mountainous system of
<quantity id="0B6F9B00FFD4FFFBFBD5F89F47EC0F6D" box="[1092,1162,1838,1862]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="5" unit="km" value="26.0">26 km</quantity>
long and
<quantity id="0B6F9B00FFD4FFFBFB6AF89F462E0F6D" box="[1275,1352,1838,1862]" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="5" unit="km" value="9.5">9.5 km</quantity>
wide is an important remnant of broadleaf evergreen forest,
</paragraph>
</subSubSection>
</treatment>
</document>