treatments-xml/data/3F/78/28/3F782856250C897EFF23FE1F8F25F958.xml

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<mods:title id="84D0462F02BD8E78FCB55CD8325A5A73">Four new species of the diving beetle genus Laccophilus Leach, 1815 from Madagascar (Coleoptera, Dytiscidae, Laccophilini)</mods:title>
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<mods:namePart id="8F46A6CA786AFEFBA57520100E8B6006">Manuel, Michaël</mods:namePart>
<mods:affiliation id="3DCE579C76761BD2F38109CFCF23D9EC">Sorbonne Université, Institut de Systématique, Evolution, Biodiversité (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, 75005 Paris, France. &amp; michael. manuel @ upmc. fr; http: // orcid. org / 0000 - 0001 - 9728 - 5580</mods:affiliation>
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<mods:namePart id="15B29423E9F450A34B23AD4BBE05E7E1">Ramahandrison, Andriamirado T.</mods:namePart>
<mods:affiliation id="3792F2621B5645DEFDF3FB95FCA2D551">Sorbonne Université, Institut de Systématique, Evolution, Biodiversité (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, 75005 Paris, France. &amp; Département de Biologie et Ecologie Végétales, Faculté des Sciences, BP 906, Université d’Antananarivo, Antananarivo, Madagascar. mirad. 95 @ gmail. com; http: // orcid. org / 0000 - 0002 - 0833 - 8730</mods:affiliation>
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<paragraph id="B76E9940250C897CFF23FE1F8DBCFE10" blockId="9.[151,488,474,537]" box="[151,488,474,501]" pageId="9" pageNumber="491">
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<taxonomicName id="70D1E2C3250C897CFF23FE1F8DD7FE10" ID-CoL="8LZ6N" authority="Manuel &amp; Ramahandrison, 2020" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[151,387,474,501]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="491" phylum="Arthropoda" rank="species" species="makay" status="sp. nov.">
<emphasis id="85A54552250C897CFF23FE1F8DD7FE10" bold="true" box="[151,387,474,501]" italics="true" pageId="9" pageNumber="491">Laccophilus makay</emphasis>
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<taxonomicNameLabel id="9E96F829250C897CFE3EFE1E8DBCFE10" box="[394,488,475,501]" pageId="9" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="FFCBCACB250C897CFF23FE3A8D9EFDFC" box="[151,458,511,537]" pageId="9" pageNumber="491" type="description">
<paragraph id="B76E9940250C897CFF23FE3A8D9EFDFC" blockId="9.[151,488,474,537]" box="[151,458,511,537]" pageId="9" pageNumber="491">
<figureCitation id="2FEA85C5250C897CFF23FE3A8D55FDFC" box="[151,257,511,537]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="9" pageNumber="491">Figures 3</figureCitation>
, 
<figureCitation id="2FEA85C5250C897CFEBAFE3A8D4AFDFC" box="[270,286,511,537]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="9" pageNumber="491">7</figureCitation>
, 
<figureCitation id="2FEA85C5250C897CFE9EFE3A8D25FDFC" box="[298,369,511,537]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="9" pageNumber="491">11, 18</figureCitation>
, 
<figureCitation id="2FEA85C5250C897CFEC9FE3A8D92FDFC" box="[381,454,511,537]" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="9" pageNumber="491">27, 28</figureCitation>
.
</paragraph>
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<paragraph id="B76E9940250C897CFF23FD838EC1FD61" blockId="9.[151,1437,581,2027]" pageId="9" pageNumber="491">
<materialsCitation id="07B9931D250C897CFF23FD838EC1FD61" ID-GBIF-Occurrence="3019656343" country="Madagascar" elevation="540" latitude="-21.216667" location="Makay" longLatPrecision="1264" longitude="45.3" pageId="9" pageNumber="491" specimenCount="540" stateProvince="Toliara" typeStatus="holotype">
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<typeStatus id="686A27E2250C897CFF23FD838C86FD85" box="[151,210,582,608]" pageId="9" pageNumber="491">Type</typeStatus>
locality.
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<collectingCountry id="CFC6D9D0250C897CFEF4FD808D9DFDBA" box="[320,457,581,607]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
, former province of 
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, 
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massif, ca. 
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North-West of Tsivoky
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, GPS coordinates 
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, altitude 
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<elevation id="3CFC7E73250C897CFDF3FDAC8EC5FD61" box="[583,657,617,644]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="491" unit="m" value="540.0">540 m</elevation>
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.
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</paragraph>
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<emphasis id="85A54552250C897CFF73FD488D27FD43" bold="true" box="[199,371,652,679]" pageId="9" pageNumber="491">Type material.</emphasis>
<materialsCitation id="07B9931D250C897CFECDFD488833FD0B" ID-GBIF-Occurrence="3019656361" collectingDate="2016-06-07" collectionCode="MNHN" collectorName="Manuel" country="Madagascar" elevation="540" latitude="-21.216667" location="Toliara" longLatPrecision="1264" longitude="45.3" pageId="9" pageNumber="491" specimenCount="541" specimenCount-male="1" stateProvince="Toliara" typeStatus="holotype">
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<typeStatus id="686A27E2250C897CFECDFD488DB2FD42" box="[377,486,653,679]" pageId="9" pageNumber="491" type="holotype">Holotype</typeStatus>
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(
<specimenCount id="A1D752C9250C897CFE41FD488E59FD42" box="[501,525,653,679]" pageId="9" pageNumber="491" type="male">♂</specimenCount>
): ” 
<collectingCountry id="CFC6D9D0250C897CFD9EFD498EE2FD43" box="[554,694,652,678]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. Ex-prov. / 
<collectingRegion id="751557A2250C897CFC86FD498FD1FD43" box="[818,901,652,678]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
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NW / Tsivoky. 
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<collectingDate id="D32B4668250C897CFB05FD488965FD43" box="[1201,1329,652,679]" pageId="9" pageNumber="491" value="2016-06-07">07 VI 2016</collectingDate>
</date>
. / 
<collectorName id="1A24FC96250C897CFAF3FD4989C9FD43" box="[1351,1437,652,678]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
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<geoCoordinate id="D2E5FF87250C897CFE89FD758DCAFD2E" box="[317,414,688,715]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.3">E45°18’</geoCoordinate>
. Alt. 
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<quantity id="702934A5250C897CFE6CFD758E4FFD2E" box="[472,539,688,715]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="491" unit="m" value="540.0">
<elevation id="3CFC7E73250C897CFE6CFD758E4FFD2E" box="[472,539,688,715]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="491" unit="m" value="540.0">540m</elevation>
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.
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/ Shadowed pools beneath / rock wall in a canyon. / Makay massif” [pr], ” 
<typeStatus id="686A27E2250C897CFAECFD758C9EFD0B" pageId="9" pageNumber="491" type="holotype">Holotype</typeStatus>
, 
<taxonomicName id="70D1E2C3250C897CFF62FD168DE5FD08" authority="Manuel &amp; Ramahandrison, 2020" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[214,433,723,749]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="491" phylum="Arthropoda" rank="species" species="makay" status="sp. nov.">Laccophilus makay</taxonomicName>
<taxonomicNameLabel id="9E96F829250C897CFE03FD118E44FD0B" box="[439,528,724,750]" pageId="9" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
, Manuel &amp; Ramahandrison 2020” [red, pr] (
<collectionCode id="D1C00185250C897CFBB7FD11880BFD0B" box="[1027,1119,724,750]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="491">MNHN</collectionCode>
)
</materialsCitation>
. 
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<emphasis id="85A54552250C897CFBC5FD1188BDFD0B" bold="true" box="[1137,1257,724,750]" pageId="9" pageNumber="491">
<typeStatus id="686A27E2250C897CFBC5FD1188BDFD0B" box="[1137,1257,724,750]" pageId="9" pageNumber="491" type="paratype">Paratypes</typeStatus>
</emphasis>
(in total 
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, 
<specimenCount id="A1D752C9250C897CFF23FD328C89FCF4" box="[151,221,759,785]" pageId="9" pageNumber="491" type="female">34♀♀</specimenCount>
) (
<collectionCode id="D1C00185250C897CFF41FD328D19FCF4" box="[245,333,759,785]" pageId="9" pageNumber="491">BMNH</collectionCode>
, 
<collectionCode id="D1C00185250C897CFEECFD328DCBFCF4" box="[344,415,759,785]" country="Brazil" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="9" pageNumber="491">CMM</collectionCode>
, 
<collectionCode id="D1C00185250C897CFE1EFD328DAFFCF4" box="[426,507,759,785]" pageId="9" pageNumber="491">DEUA</collectionCode>
, 
<collectionCode id="D1C00185250C897CFDB2FD328E0AFCF4" box="[518,606,759,785]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="491">MNHN</collectionCode>
, 
<collectionCode id="D1C00185250C897CFDDDFD328EE2FCF4" box="[617,694,759,785]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="9" pageNumber="491">NHRS</collectionCode>
, 
<collectionCode id="D1C00185250C897CFD75FD328F47FCF4" box="[705,787,759,785]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="491">NMPC</collectionCode>
, 
<collectionCode id="D1C00185250C897CFCAAFD328F32FCF4" box="[798,870,759,785]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="9" pageNumber="491">PBZT</collectionCode>
/ 
<collectionCode id="D1C00185250C897CFCDBFD328FFBFCF4" box="[879,943,759,785]" country="USA" name="Montgomery Botanical Center" pageId="9" pageNumber="491">MBC</collectionCode>
)
</materialsCitation>
: 
<materialsCitation id="07B9931D250C897CFC75FD328900FCF4" ID-GBIF-Occurrence="3019656353" box="[961,1364,759,785]" collectingDate="2016-06-07" collectionCode="MNHN" collectorName="Manuel" country="Madagascar" elevation="540" latitude="-21.216667" location="Toliara" longLatPrecision="1264" longitude="45.3" pageId="9" pageNumber="491" specimenCount="18" specimenCount-female="6" specimenCount-male="12" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFC75FD32885DFCF4" box="[961,1033,759,785]" pageId="9" pageNumber="491" type="male">12♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFBA0FD32881AFCF4" box="[1044,1102,759,785]" pageId="9" pageNumber="491" type="female">6♀♀</specimenCount>
, same data as holotype
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFAE9FD328FA4FCBD" ID-GBIF-Occurrence="3019656339" collectingDate="2016-06-04" collectorName="Manuel" country="Madagascar" county="Ex-prov." elevation="500" latitude="-21.216667" location="river Andranomanitsy" longLatPrecision="1264" longitude="45.333332" municipality="Tsivoky." pageId="9" pageNumber="491" specimenCount="515" specimenCount-female="7" specimenCount-male="8" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFAE9FD3289CCFCF4" box="[1373,1432,759,785]" pageId="9" pageNumber="491" type="male">8♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFF23FCDE8C84FCD0" box="[151,208,795,821]" pageId="9" pageNumber="491" type="female">7♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFF5CFCDF8D20FCD1" box="[232,372,794,820]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingCounty id="5E0FE1CC250C897CFECBFCDE8DB5FCD0" box="[383,481,795,821]" pageId="9" pageNumber="491">Ex-prov.</collectingCounty>
/ 
<collectingRegion id="751557A2250C897CFE4CFCDF8E1FFCD1" box="[504,587,794,820]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFDC2FCDF8EEAFCD1" box="[630,702,794,820]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NNW / 
<collectingMunicipality id="570A033A250C897CFC94FCDF8FD4FCD1" box="[800,896,794,820]" pageId="9" pageNumber="491">Tsivoky.</collectingMunicipality>
<date id="C36FBF80250C897CFC33FCDF8858FCD1" box="[903,1036,794,821]" pageId="9" pageNumber="491" value="2016-06-04">
<collectingDate id="D32B4668250C897CFC33FCDF8858FCD1" box="[903,1036,794,821]" pageId="9" pageNumber="491" value="2016-06-04">04 VI 2016</collectingDate>
</date>
. / 
<collectorName id="1A24FC96250C897CFB93FCDF8829FCD1" box="[1063,1149,794,820]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFB79FCDF8977FCD0" box="[1229,1315,794,821]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
/ 
<geoCoordinate id="D2E5FF87250C897CFA98FCDF89D9FCD0" box="[1324,1421,794,821]" degrees="45" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.333332">E45°20’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFF7EFCFB8D45FCBC" box="[202,273,830,857]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFF7EFCFB8D59FCBC" box="[202,269,830,857]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">
<elevation id="3CFC7E73250C897CFF7EFCFB8D59FCBC" box="[202,269,830,857]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">500m</elevation>
</quantity>
.
</specimenCount>
Sandy pool / on bank of 
<location id="B20ECF9B250C897CFD87FCFB8F7EFCBD" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFD87FCFB8F7EFCBD" box="[563,810,830,856]" country="Madagascar" county="Ex-prov." latitude="-21.216667" longLatPrecision="1264" longitude="45.333332" municipality="Tsivoky." name="river Andranomanitsy" pageId="9" pageNumber="491" stateProvince="Toliara">river Andranomanitsy</location>
/ Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFC48FCFB8E01FC7A" ID-GBIF-Occurrence="3019656317" collectingDate="2016-06-04" collectorName="Manuel" country="Madagascar" county="Ex-prov." elevation="500" latitude="-21.216667" location="river Andranomanitsy" longLatPrecision="1264" longitude="45.333332" municipality="Tsivoky." pageId="9" pageNumber="491" specimenCount="503" specimenCount-female="1" specimenCount-male="2" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFC48FCFB8863FCBD" box="[1020,1079,830,856]" pageId="9" pageNumber="491" type="male">2♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFBF7FCFB882FFCBD" box="[1091,1147,830,856]" pageId="9" pageNumber="491" type="female">1♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFB20FCFB8974FCBD" box="[1172,1312,830,856]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingCounty id="5E0FE1CC250C897CFA9FFCFB89D9FCBD" box="[1323,1421,830,856]" pageId="9" pageNumber="491">Ex-prov.</collectingCounty>
/ 
<collectingRegion id="751557A2250C897CFF23FCA48CBEFC9E" box="[151,234,865,891]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFEA3FCA48D34FC9E" box="[279,352,865,891]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NNW / 
<collectingMunicipality id="570A033A250C897CFE77FCA48E77FC9E" box="[451,547,865,891]" pageId="9" pageNumber="491">Tsivoky.</collectingMunicipality>
<date id="C36FBF80250C897CFD9FFCA48EF9FC9E" box="[555,685,865,892]" pageId="9" pageNumber="491" value="2016-06-04">
<collectingDate id="D32B4668250C897CFD9FFCA48EF9FC9E" box="[555,685,865,892]" pageId="9" pageNumber="491" value="2016-06-04">04 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFD72FCA48F48FC9E" box="[710,796,865,891]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFCDAFCA48F9CFC9E" box="[878,968,865,891]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFC64FCA48878FC99" box="[976,1068,865,892]" degrees="45" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.333332">E45°20’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFBC3FCA488EAFC99" box="[1143,1214,865,892]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFBC3FCA488EEFC99" box="[1143,1210,865,892]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">
<elevation id="3CFC7E73250C897CFBC3FCA488EEFC99" box="[1143,1210,865,892]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">500m</elevation>
</quantity>
.
</specimenCount>
Spring, / alongside 
<location id="B20ECF9B250C897CFF23FC408DDAFC7A" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFF23FC408DDAFC7A" box="[151,398,901,927]" country="Madagascar" county="Ex-prov." latitude="-21.216667" longLatPrecision="1264" longitude="45.333332" municipality="Tsivoky." name="river Andranomanitsy" pageId="9" pageNumber="491" stateProvince="Toliara">river Andranomanitsy</location>
/ Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFDD4FC408D01FC03" ID-GBIF-Occurrence="3019656330" collectingDate="2016-06-05" collectorName="Ramahandrison &amp; Manuel" country="Madagascar" elevation="650" latitude="-21.216667" location="Tsivoky" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="655" specimenCount-female="1" specimenCount-male="4" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFDD4FC408ECFFC7A" box="[608,667,901,927]" pageId="9" pageNumber="491" type="male">4♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFD1CFC408EB4FC7A" box="[680,736,901,927]" pageId="9" pageNumber="491" type="female">1♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFD4CFC408FD0FC7A" box="[760,900,901,927]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFC24FC408FA6FC7A" box="[912,1010,901,927]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFBBDFC408808FC7A" box="[1033,1116,901,927]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFB3CFC408884FC7A" box="[1160,1232,901,927]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NW / 
<location id="B20ECF9B250C897CFAAAFC40892CFC7A" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFAAAFC40892CFC7A" box="[1310,1400,901,927]" country="Madagascar" latitude="-21.216667" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky" pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky</location>
<date id="C36FBF80250C897CFA34FC408CADFC27" pageId="9" pageNumber="491" value="2016-06-05">
<collectingDate id="D32B4668250C897CFA34FC408CADFC27" pageId="9" pageNumber="491" value="2016-06-05">05 VI 2016</collectingDate>
</date>
. / 
<collectorName id="1A24FC96250C897CFEA0FC6D8D90FC27" box="[276,452,936,962]" pageId="9" pageNumber="491">Ramahandrison</collectorName>
&amp; 
<collectorName id="1A24FC96250C897CFE5EFC6D8E14FC27" box="[490,576,936,962]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFD25FC6D8EBFFC27" box="[657,747,936,962]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFD40FC6D8F04FC26" box="[756,848,936,963]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFC2DFC6D8FB4FC26" box="[921,992,936,963]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFC2DFC6D8F88FC26" box="[921,988,936,963]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="491" unit="m" value="650.0">
<elevation id="3CFC7E73250C897CFC2DFC6D8F88FC26" box="[921,988,936,963]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="491" unit="m" value="650.0">650m</elevation>
</quantity>
.
</specimenCount>
Lake above / natural dam in a canyon. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFED4FC098DD5FBC8" ID-GBIF-Occurrence="3019656362" collectingDate="2016-06-05" collectorName="Ramahandrison &amp; Manuel" country="Madagascar" elevation="650" latitude="-21.216667" location="Tsivoky." longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="653" specimenCount-male="3" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFED4FC098DCFFC03" box="[352,411,972,998]" pageId="9" pageNumber="491" type="male">3♂♂</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFE04FC098E68FC03" box="[432,572,972,998]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFDFCFC098EFDFC03" box="[584,681,972,998]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFD0BFC098F46FC03" box="[703,786,972,998]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFC89FC098FD1FC03" box="[829,901,972,998]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NW / 
<location id="B20ECF9B250C897CFC65FC098865FC03" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFC65FC098865FC03" box="[977,1073,972,998]" country="Madagascar" latitude="-21.216667" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFB8CFC0E88ECFC03" box="[1080,1208,971,998]" pageId="9" pageNumber="491" value="2016-06-05">
<collectingDate id="D32B4668250C897CFB8CFC0E88ECFC03" box="[1080,1208,971,998]" pageId="9" pageNumber="491" value="2016-06-05">05 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFB7BFC09892BFC03" box="[1231,1407,972,998]" pageId="9" pageNumber="491">Ramahandrison</collectorName>
&amp; 
<collectorName id="1A24FC96250C897CFF23FC2A8CB9FBEC" box="[151,237,1007,1033]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFE8EFC2A8DC0FBEC" box="[314,404,1007,1033]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFE2FFC2A8DA3FBEF" box="[411,503,1007,1034]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFD88FC2A8ED7FBEF" box="[572,643,1007,1034]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFD88FC2A8E2BFBEF" box="[572,639,1007,1034]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="491" unit="m" value="650.0">
<elevation id="3CFC7E73250C897CFD88FC2A8E2BFBEF" box="[572,639,1007,1034]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="491" unit="m" value="650.0">650m</elevation>
</quantity>
.
</specimenCount>
Slowly flowing / stream in a canyon, bottom / with much organic matter. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFE38FBD68D11FB96" ID-GBIF-Occurrence="3019656308" collectingDate="2016-06-05" collectorName="Manuel" country="Madagascar" elevation="700" latitude="-21.2" location="Tsivoky." longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="703" specimenCount-female="2" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFE38FBD68D92FBC8" box="[396,454,1043,1069]" pageId="9" pageNumber="491" type="male">1♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFE67FBD68E59FBC8" box="[467,525,1043,1069]" pageId="9" pageNumber="491" type="female">2♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFD90FBD68EE4FBC8" box="[548,688,1043,1069]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFD0FFBD68F49FBC8" box="[699,797,1043,1069]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFC87FBD68FD2FBC8" box="[819,902,1043,1069]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFC06FBD68FAEFBC8" box="[946,1018,1043,1069]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NW / 
<location id="B20ECF9B250C897CFBF3FBD688F3FBC8" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFBF3FBD688F3FBC8" box="[1095,1191,1043,1069]" country="Madagascar" latitude="-21.2" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFB1AFBD7897BFBC8" box="[1198,1327,1042,1069]" pageId="9" pageNumber="491" value="2016-06-05">
<collectingDate id="D32B4668250C897CFB1AFBD7897BFBC8" box="[1198,1327,1042,1069]" pageId="9" pageNumber="491" value="2016-06-05">05 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFAF3FBD689C9FBC8" box="[1351,1437,1043,1069]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. 
<geoCoordinate id="D2E5FF87250C897CFF7CFBF38D76FBB5" box="[200,290,1078,1104]" degrees="21" direction="south" minutes="12" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.2">S21°12’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFE9DFBF38DD1FBB4" box="[297,389,1078,1105]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
// Alt. 
<specimenCount id="A1D752C9250C897CFE67FBF38E4EFBB5" box="[467,538,1078,1104]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFE67FBF38E42FBB5" box="[467,534,1078,1104]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="491" unit="m" value="700.0">
<elevation id="3CFC7E73250C897CFE67FBF38E42FBB5" box="[467,534,1078,1104]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="491" unit="m" value="700.0">700m</elevation>
</quantity>
.
</specimenCount>
Quiet zone / alongside brook in deep / canyon, with masses of / orange deposit. Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFEE4FB9F8FA1FB72" ID-GBIF-Occurrence="3019656318" collectingDate="2016-06-05" collectorName="Manuel" country="Madagascar" elevation="620" latitude="-21.2" location="Tsivoky." longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="622" specimenCount-female="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFEE4FB9F8DDEFB91" box="[336,394,1114,1140]" pageId="9" pageNumber="491" type="male">1♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFE22FB9F8D9AFB91" box="[406,462,1114,1140]" pageId="9" pageNumber="491" type="female">1♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFE52FB9C8E26FB96" box="[486,626,1113,1139]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFDC9FB9F8E8BFB91" box="[637,735,1114,1140]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFD40FB9C8F13FB96" box="[756,839,1113,1139]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFCC5FB9F8FECFB96" box="[881,952,1113,1140]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NW / 
<location id="B20ECF9B250C897CFBB7FB9C8837FB96" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFBB7FB9C8837FB96" box="[1027,1123,1113,1139]" country="Madagascar" latitude="-21.2" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFBDDFB9C88BDFB96" box="[1129,1257,1113,1140]" pageId="9" pageNumber="491" value="2016-06-05">
<collectingDate id="D32B4668250C897CFBDDFB9C88BDFB96" box="[1129,1257,1113,1140]" pageId="9" pageNumber="491" value="2016-06-05">05 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFB4BFB9C8901FB96" box="[1279,1365,1113,1139]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFF23FBB88CA5FB72" box="[151,241,1149,1175]" degrees="21" direction="south" minutes="12" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.2">S21°12’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFF4DFBB88D01FB7D" box="[249,341,1149,1176]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFE29FBB88DB0FB72" box="[413,484,1149,1175]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFE29FBB88DB4FB72" box="[413,480,1149,1175]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="491" unit="m" value="620.0">
<elevation id="3CFC7E73250C897CFE29FBB88DB4FB72" box="[413,480,1149,1175]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="491" unit="m" value="620.0">620m</elevation>
</quantity>
.
</specimenCount>
Pool full / of orange deposit. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFBB4FBB88E69FB3B" ID-GBIF-Occurrence="3019656355" collectingDate="2016-06-05" collectorName="Manuel" country="Madagascar" elevation="550" latitude="-21.216667" location="Tsivoky." longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="570" specimenCount-female="9" specimenCount-male="11" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFBB4FBB8881CFB72" box="[1024,1096,1149,1175]" pageId="9" pageNumber="491" type="male">11♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFBE0FBB888D9FB72" box="[1108,1165,1149,1175]" pageId="9" pageNumber="491" type="female">9♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFB10FBB88964FB72" box="[1188,1328,1149,1175]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFA8FFBBB89C9FB7D" box="[1339,1437,1150,1176]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFF13FB658CAEFB5F" box="[167,250,1184,1210]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFE93FB648D3BFB5F" box="[295,367,1184,1211]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="491" unit="km" value="11.0">11 km</quantity>
NNW / 
<location id="B20ECF9B250C897CFE66FB658E66FB5F" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFE66FB658E66FB5F" box="[466,562,1184,1210]" country="Madagascar" latitude="-21.216667" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFD8EFB658EE8FB5F" box="[570,700,1184,1211]" pageId="9" pageNumber="491" value="2016-06-05">
<collectingDate id="D32B4668250C897CFD8EFB658EE8FB5F" box="[570,700,1184,1211]" pageId="9" pageNumber="491" value="2016-06-05">05 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFD61FB658F7FFB5F" box="[725,811,1184,1210]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFCCAFB648F8CFB5E" box="[894,984,1185,1211]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFC54FB658868FB5E" box="[992,1084,1184,1211]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFB33FB65889AFB5E" box="[1159,1230,1184,1211]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFB33FB65889EFB5E" box="[1159,1226,1184,1211]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="491" unit="m" value="550.0">
<elevation id="3CFC7E73250C897CFB33FB65889EFB5E" box="[1159,1226,1184,1211]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="491" unit="m" value="550.0">550m</elevation>
</quantity>
.
</specimenCount>
Large pool / with masses of tree roots. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFDF2FB018D07FAC0" ID-GBIF-Occurrence="3019656320" collectingDate="2016-06-06" collectorName="Manuel" country="Madagascar" elevation="600" latitude="-21.216667" location="Tsivoky." longLatPrecision="1264" longitude="45.35" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="606" specimenCount-female="4" specimenCount-male="2" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFDF2FB018ED5FB3B" box="[582,641,1220,1246]" pageId="9" pageNumber="491" type="male">2♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFD38FB018E91FB3B" box="[652,709,1220,1246]" pageId="9" pageNumber="491" type="female">4♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFD6FFB018F33FB3B" box="[731,871,1220,1246]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFCC5FB008F87FB3A" box="[881,979,1221,1247]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFC52FB01886DFB3B" box="[998,1081,1220,1246]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFBD6FB0188CEFB3B" box="[1122,1178,1220,1246]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="491" unit="km" value="9.0">9 km</quantity>
NNW / 
<location id="B20ECF9B250C897CFB43FB018903FB3B" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFB43FB018903FB3B" box="[1271,1367,1220,1246]" country="Madagascar" latitude="-21.216667" longLatPrecision="1264" longitude="45.35" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFAE9FB018C9BFAE4" pageId="9" pageNumber="491" value="2016-06-06">
<collectingDate id="D32B4668250C897CFAE9FB018C9BFAE4" pageId="9" pageNumber="491" value="2016-06-06">06 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFF50FB228D6EFAE4" box="[228,314,1255,1281]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFE31FB2D8D8BFAE7" box="[389,479,1256,1282]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFE51FB228E15FAE7" box="[485,577,1255,1282]" degrees="45" direction="east" minutes="21" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.35">E45°21’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFD30FB228E98FAE4" box="[644,716,1255,1281]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFD30FB228E9CFAE4" box="[644,712,1255,1281]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="491" unit="m" value="600.0">
<elevation id="3CFC7E73250C897CFD30FB228E9CFAE4" box="[644,712,1255,1281]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="491" unit="m" value="600.0">600m</elevation>
</quantity>
.
</specimenCount>
Small pools, / muddy-sandy bottom, / with tree roots, in canyon. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFEE9FACE8853FAAD" ID-GBIF-Occurrence="3019656360" collectingDate="2016-06-07" collectorName="Manuel" country="Madagascar" elevation="570" latitude="-21.216667" location="Tsivoky." longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." pageId="9" pageNumber="491" specimenCount="575" specimenCount-female="2" specimenCount-male="3" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFEE9FACE8DC3FAC0" box="[349,407,1291,1317]" pageId="9" pageNumber="491" type="male">3♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFE17FACE8D88FAC0" box="[419,476,1291,1317]" pageId="9" pageNumber="491" type="female">2♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFE46FACE8E2AFAC0" box="[498,638,1291,1317]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingMunicipality id="570A033A250C897CFD3CFACE8EBEFAC0" box="[648,746,1291,1317]" pageId="9" pageNumber="491">Ex-prov.</collectingMunicipality>
/ 
<collectingRegion id="751557A2250C897CFD49FACE8F04FAC0" box="[765,848,1291,1317]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFCCDFACE8F81FAC0" box="[889,981,1291,1317]" metricMagnitude="4" metricUnit="m" metricValue="1.08" pageId="9" pageNumber="491" unit="km" value="10.8">10.8 km</quantity>
NW / 
<location id="B20ECF9B250C897CFBAAFACE882AFAC0" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFBAAFACE882AFAC0" box="[1054,1150,1291,1317]" country="Madagascar" latitude="-21.216667" longLatPrecision="1264" longitude="45.316666" municipality="Ex-prov." name="Tsivoky." pageId="9" pageNumber="491" stateProvince="Toliara">Tsivoky.</location>
<date id="C36FBF80250C897CFB30FACE8956FAC0" box="[1156,1282,1291,1317]" pageId="9" pageNumber="491" value="2016-06-07">
<collectingDate id="D32B4668250C897CFB30FACE8956FAC0" box="[1156,1282,1291,1317]" pageId="9" pageNumber="491" value="2016-06-07">07 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFAA3FACE8939FAC0" box="[1303,1389,1291,1317]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFF19FAEB8D53FAAD" box="[173,263,1326,1352]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFEBAFAEB8D3EFAAC" box="[270,362,1326,1353]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFE19FAEB8DA1FAAC" box="[429,501,1326,1353]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFE19FAEB8DA5FAAC" box="[429,497,1326,1353]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="491" unit="m" value="570.0">
<elevation id="3CFC7E73250C897CFE19FAEB8DA5FAAC" box="[429,497,1326,1353]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="491" unit="m" value="570.0">570m</elevation>
</quantity>
.
</specimenCount>
Shallow / stream, under trees. / Makay massif”
</materialsCitation>
. 
<materialsCitation id="07B9931D250C897CFBA5FAEB8958FA6A" ID-GBIF-Occurrence="3019656323" collectingDate="2016-06-07" collectorName="Manuel" country="Madagascar" county="Ex-prov." elevation="500" latitude="-21.216667" location="river Andranomanitsy" longLatPrecision="1264" longitude="45.316666" municipality="Tsivoky." pageId="9" pageNumber="491" specimenCount="502" specimenCount-female="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="A1D752C9250C897CFBA5FAEB881FFAAD" box="[1041,1099,1326,1352]" pageId="9" pageNumber="491" type="male">1♂♂</specimenCount>
, 
<specimenCount id="A1D752C9250C897CFBE2FAEB88DAFAAD" box="[1110,1166,1326,1352]" pageId="9" pageNumber="491" type="female">1♀♀</specimenCount>
, “ 
<collectingCountry id="CFC6D9D0250C897CFB11FAEB8965FAAD" box="[1189,1329,1326,1352]" name="Madagascar" pageId="9" pageNumber="491">Madagascar</collectingCountry>
. 
<collectingCounty id="5E0FE1CC250C897CFA8FFAEA89C9FAAC" box="[1339,1437,1327,1353]" pageId="9" pageNumber="491">Ex-prov.</collectingCounty>
/ 
<collectingRegion id="751557A2250C897CFF11FA978CACFA89" box="[165,248,1362,1388]" country="Madagascar" name="Toliara" pageId="9" pageNumber="491">Toliara</collectingRegion>
. ca 
<quantity id="702934A5250C897CFE96FA978D3DFA89" box="[290,361,1362,1388]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="491" unit="km" value="10.0">10 km</quantity>
NW / 
<collectingMunicipality id="570A033A250C897CFE06FA978E46FA89" box="[434,530,1362,1388]" pageId="9" pageNumber="491">Tsivoky.</collectingMunicipality>
<date id="C36FBF80250C897CFDACFA978EC3FA89" box="[536,663,1362,1388]" pageId="9" pageNumber="491" value="2016-06-07">
<collectingDate id="D32B4668250C897CFDACFA978EC3FA89" box="[536,663,1362,1388]" pageId="9" pageNumber="491" value="2016-06-07">07 VI 2016</collectingDate>
</date>
/ 
<collectorName id="1A24FC96250C897CFD19FA978F57FA89" box="[685,771,1362,1388]" pageId="9" pageNumber="491">Manuel</collectorName>
leg. // 
<geoCoordinate id="D2E5FF87250C897CFCFBFA978FFEFA89" box="[847,938,1362,1388]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="491" precision="925" value="-21.216667">S21°13’</geoCoordinate>
<geoCoordinate id="D2E5FF87250C897CFC04FA978858FA88" box="[944,1036,1362,1389]" degrees="45" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="491" precision="925" value="45.316666">E45°19’</geoCoordinate>
/ Alt. 
<specimenCount id="A1D752C9250C897CFBE5FA9788CDFA88" box="[1105,1177,1362,1389]" pageId="9" pageNumber="491" type="generic">
<quantity id="702934A5250C897CFBE5FA9788C1FA88" box="[1105,1173,1362,1389]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">
<elevation id="3CFC7E73250C897CFBE5FA9788C1FA88" box="[1105,1173,1362,1389]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="491" unit="m" value="500.0">500m</elevation>
</quantity>
.
</specimenCount>
Pool with / vegetation, alongside / 
<location id="B20ECF9B250C897CFEACFAB08E5AFA6A" LSID="urn:lsid:plazi:treatment:3F782856250C897EFF23FE1F8F25F958:B20ECF9B250C897CFEACFAB08E5AFA6A" box="[280,526,1397,1423]" country="Madagascar" county="Ex-prov." latitude="-21.216667" longLatPrecision="1264" longitude="45.316666" municipality="Tsivoky." name="river Andranomanitsy" pageId="9" pageNumber="491" stateProvince="Toliara">river Andranomanitsy</location>
/ Makay massif”. All 
<typeStatus id="686A27E2250C897CFCB1FAB38F24FA75" box="[773,880,1398,1424]" pageId="9" pageNumber="491" type="paratype">paratypes</typeStatus>
with the respective printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FFCBCACB250C897EFF73FA5D8E40FDFC" lastPageId="11" lastPageNumber="493" pageId="9" pageNumber="491" type="description">
<paragraph id="B76E9940250C897CFF73FA5D8F08F9F8" blockId="9.[151,1437,581,2027]" pageId="9" pageNumber="491">
<emphasis id="85A54552250C897CFF73FA5D8DB6FA56" bold="true" box="[199,482,1432,1459]" pageId="9" pageNumber="491">
Description of 
<typeStatus id="686A27E2250C897CFECFFA5C8DB6FA56" box="[379,482,1433,1459]" pageId="9" pageNumber="491" type="holotype">holotype</typeStatus>
</emphasis>
. 
<emphasis id="85A54552250C897CFE46FA5D8E1AFA56" bold="true" box="[498,590,1432,1459]" italics="true" pageId="9" pageNumber="491">Habitus</emphasis>
(
<figureCitation id="2FEA85C5250C897CFDEBFA5C8EF7FA56" box="[607,675,1433,1459]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="9" pageNumber="491">Fig. 3</figureCitation>
) oval-oblong, with lateral sides widely rounded, posteriorly rather strongly tapered to rounded apex; lateral outline continuous between pronotum and elytra; maximum body width at about 43% of total body length and about 31% of elytral length; dorsal surface evenly and moderately convex; in lateral view maximum height at about first third of total length.
</paragraph>
<paragraph id="B76E9940250C897CFF73F9E389C9F92B" blockId="9.[151,1437,581,2027]" pageId="9" pageNumber="491">
<emphasis id="85A54552250C897CFF73F9E38D0EF9A4" bold="true" box="[199,346,1574,1601]" italics="true" pageId="9" pageNumber="491">Colouration.</emphasis>
Dorsal surface of head rufo-testaceous. Pronotum rufo-testaceous, very narrowly darkened along posterior margin. Elytron with typical 
<emphasis id="85A54552250C897CFDFCF98F8EFBF981" box="[584,687,1610,1636]" italics="true" pageId="9" pageNumber="491">alluaudi-</emphasis>
group colour pattern as in 
<figureCitation id="2FEA85C5250C897CFC63F98F884FF981" box="[983,1051,1610,1636]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="9" pageNumber="491">Fig. 3</figureCitation>
; brown longitudinal vittae weakly contrasting with testaceous background, anteriorly vaguely delimited, leaving rather wide basal area and humeral region entirely testaceous; pale stripes between dark longitudinal vittae irregularly interrupted. Ventral surface entirely rufo-testaceous (
<figureCitation id="2FEA85C5250C897CFE20F9718D8CF92A" box="[404,472,1716,1743]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="9" pageNumber="491">Fig. 7</figureCitation>
). Antennae, palps, anterior and median legs testaceous; posterior legs rufo-testaceous.
</paragraph>
<paragraph id="B76E9940250C897CFF73F9128E10F8B9" blockId="9.[151,1437,581,2027]" pageId="9" pageNumber="491">
<emphasis id="85A54552250C897CFF73F9128D51F917" bold="true" box="[199,261,1751,1778]" italics="true" pageId="9" pageNumber="491">Head</emphasis>
. Anterior outline of clypeus widely rounded, subtruncated in middle. Surface shiny, with double reticulation; large meshes irregular and of variable sizes, more strongly impressed than small meshes; with sparse micropunctures, on vertex denser and coarser; larger punctures present along inner eye margins and in two irregular transverse rows sub-laterally on frons.
</paragraph>
<paragraph id="B76E9940250C897CFF73F8A38D9CF80E" blockId="9.[151,1437,581,2027]" pageId="9" pageNumber="491">
<emphasis id="85A54552250C897CFF73F8A38D68F865" bold="true" box="[199,316,1894,1920]" italics="true" pageId="9" pageNumber="491">Pronotum</emphasis>
. Lateral margins evenly arcuate. Surface shiny, with double reticulation; large meshes irregular and of variable sizes, more strongly impressed than small meshes; small meshes rounded, more strongly impressed than on head; punctation inconspicuous except in anterior quarter and close to posterior angles with sparse, moderately large and coarse punctures.
</paragraph>
<paragraph id="B76E9940250F897FFF73FF53886CFF1C" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="492">
<emphasis id="85A54552250F897FFF73FF538D71FF54" bold="true" box="[199,293,150,177]" italics="true" pageId="10" pageNumber="492">Elytron.</emphasis>
Surface shiny, with double reticulation; large meshes indistinct except in scutellar region; small meshes irregularly polygonal, of rather homogeneous sizes, larger and much more strongly impressed than on pronotum; punctation inconspicuous beyond rather indistinct discal and lateral puncture rows.
</paragraph>
<paragraph id="B76E9940250F897FFF73FEC7888CFE10" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="492">
<emphasis id="85A54552250F897FFF73FEC78D23FEF9" bold="true" box="[199,375,258,285]" italics="true" pageId="10" pageNumber="492">Ventral surface</emphasis>
. Prosternal process narrowly lanceolate, strongly convex, with apex acutely pointed. Metacoxal lines anteriorly subparallel. Apical abdominal ventrite strongly convex, in posterior half almost tectiform; without protuberance; posterior margin unmodified, very weakly arcuate on each side of slightly produced, evenly rounded apex. Entire ventral surface very shiny; on elytral epipleura, metaventrite, metacoxa and abdominal ventrites with very fine regular microstriae; on abdominal ventrites with oblique striae, more strongly impressed on ventrites I–III; surface impunctate except on antero-medial prosternal region, anterior region of metaventrite and apical abdominal ventrite with shallow punctures of moderate size. Metacoxal plate without stridulatory apparatus.
</paragraph>
<paragraph id="B76E9940250F897FFF73FE3A890FFDFC" blockId="10.[151,1437,150,537]" box="[199,1371,511,537]" pageId="10" pageNumber="492">
<emphasis id="85A54552250F897FFF73FE3A8D51FDFC" bold="true" box="[199,261,511,537]" italics="true" pageId="10" pageNumber="492">Legs.</emphasis>
Pro- and mesotarsi not distinctly broadened, but with four transverse rows of small adhesive setae.
</paragraph>
<caption id="E3AEC9C8250F897FFF23F8B08F4AF830" ID-DOI="http://doi.org/10.5281/zenodo.4401795" ID-Zenodo-Dep="4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="10" pageNumber="492" startId="10.[151,264,1909,1933]" targetBox="[157,1431,561,1885]" targetPageId="10">
<paragraph id="B76E9940250F897FFF23F8B08F4AF830" blockId="10.[151,1436,1909,2006]" pageId="10" pageNumber="492">
<emphasis id="85A54552250F897FFF23F8B08D00F868" bold="true" box="[151,340,1909,1933]" pageId="10" pageNumber="492">FIGURES 23–30.</emphasis>
Left 
<emphasis id="85A54552250F897FFE3EF8B08E71F868" bold="true" box="[394,549,1909,1933]" pageId="10" pageNumber="492">(23, 25, 27, 29)</emphasis>
and right 
<emphasis id="85A54552250F897FFD38F8B08F73F868" bold="true" box="[652,807,1909,1933]" pageId="10" pageNumber="492">(24, 26, 28, 30)</emphasis>
parameres of: 
<emphasis id="85A54552250F897FFC09F8B08859F868" bold="true" box="[957,1037,1909,1933]" pageId="10" pageNumber="492">(23, 24)</emphasis>
<taxonomicName id="70D1E2C3250F897FFBA7F8B08955F868" authority="Manuel &amp; Ramahandrison, 2020" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[1043,1281,1909,1933]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="492" phylum="Arthropoda" rank="species" species="leguyaderi" status="sp. nov.">
<emphasis id="85A54552250F897FFBA7F8B08955F868" box="[1043,1281,1909,1933]" italics="true" pageId="10" pageNumber="492">Laccophilus leguyaderi</emphasis>
</taxonomicName>
<emphasis id="85A54552250F897FFABCF8B3890CF86B" bold="true" box="[1288,1368,1910,1934]" pageId="10" pageNumber="492">
<taxonomicNameLabel id="9E96F829250F897FFABCF8B3890CF86B" box="[1288,1368,1910,1934]" pageId="10" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); 
<emphasis id="85A54552250F897FFF6CF85C8D7CF854" bold="true" box="[216,296,1945,1969]" pageId="10" pageNumber="492">(25, 26)</emphasis>
<taxonomicName id="70D1E2C3250F897FFE9AF85C8E58F854" authority="Manuel &amp; Ramahandrison, 2020" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[302,524,1945,1969]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="492" phylum="Arthropoda" rank="species" species="bergsteni" status="sp. nov.">
<emphasis id="85A54552250F897FFE9AF85C8E58F854" box="[302,524,1945,1969]" italics="true" pageId="10" pageNumber="492">Laccophilus bergsteni</emphasis>
</taxonomicName>
<emphasis id="85A54552250F897FFDA6F85F8E36F857" bold="true" box="[530,610,1946,1970]" pageId="10" pageNumber="492">
<taxonomicNameLabel id="9E96F829250F897FFDA6F85F8E36F857" box="[530,610,1946,1970]" pageId="10" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); 
<emphasis id="85A54552250F897FFD6BF85C8F7BF854" bold="true" box="[735,815,1945,1969]" pageId="10" pageNumber="492">(27, 28)</emphasis>
<taxonomicName id="70D1E2C3250F897FFC81F85C8FACF854" authority="Manuel &amp; Ramahandrison, 2020" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[821,1016,1945,1969]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="492" phylum="Arthropoda" rank="species" species="makay" status="sp. nov.">
<emphasis id="85A54552250F897FFC81F85C8FACF854" box="[821,1016,1945,1969]" italics="true" pageId="10" pageNumber="492">Laccophilus makay</emphasis>
</taxonomicName>
<emphasis id="85A54552250F897FFC4AF85F881AF857" bold="true" box="[1022,1102,1946,1970]" pageId="10" pageNumber="492">
<taxonomicNameLabel id="9E96F829250F897FFC4AF85F881AF857" box="[1022,1102,1946,1970]" pageId="10" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); 
<emphasis id="85A54552250F897FFB7FF85C894EF854" bold="true" box="[1227,1306,1945,1969]" pageId="10" pageNumber="492">(29, 30)</emphasis>
<taxonomicName id="70D1E2C3250F897FFA94F85C8CBFF830" authorityName="Regimbart" authorityYear="1900" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="492" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250F897FFA94F85C8CBFF830" italics="true" pageId="10" pageNumber="492">Laccophilus alluaudi</emphasis>
</taxonomicName>
(male specimen from Isalo massif). Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="B76E9940250E897EFF73FF538CEEFF30" blockId="11.[151,1437,150,645]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FF538D63FF54" bold="true" box="[199,311,150,177]" italics="true" pageId="11" pageNumber="493">Aedeagus</emphasis>
. Median lobe in lateral view as in 
<figureCitation id="2FEA85C5250E897EFD73FF528F4DFF54" box="[711,793,151,177]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Fig. 11</figureCitation>
, in ventral view as in 
<figureCitation id="2FEA85C5250E897EFBACFF528838FF54" box="[1048,1132,151,177]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Fig. 18</figureCitation>
; parameres as in 
<figureCitation id="2FEA85C5250E897EFA87FF528CE2FF30" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="11" pageNumber="493">Figs. 27– 28</figureCitation>
.
</paragraph>
<paragraph id="B76E9940250E897EFF73FF1A8F3EFF1C" blockId="11.[151,1437,150,645]" box="[199,874,223,249]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FF1A8D64FF1C" bold="true" box="[199,304,223,249]" pageId="11" pageNumber="493">Females.</emphasis>
Pro- and mesotarsi without ventral adhesive setae.
</paragraph>
<paragraph id="B76E9940250E897EFF73FEC78E9DFE48" blockId="11.[151,1437,150,645]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FEC78D19FEF9" bold="true" box="[199,333,258,284]" pageId="11" pageNumber="493">Variability.</emphasis>
Specimens vary slightly in dorsal colour pattern, mainly with respect to homogeneity of colouration within the elytral brown longitudinal markings (in some specimens entirely pale brown, in others with gradually paler medial area enclosed in dark markings) and, to a lesser extent, degree of fragmentation of the pale interspaces between dark bands. However, the overall elytral pattern (notably, position of the anterior extremities of the dark bands) is highly stable among studied individuals.
</paragraph>
<paragraph id="B76E9940250E897EFF73FE728E40FDFC" blockId="11.[151,1437,150,645]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FE728D2AFE34" bold="true" box="[199,382,439,465]" pageId="11" pageNumber="493">Measurements.</emphasis>
<typeStatus id="686A27E2250E897EFE37FE728DA4FE34" box="[387,496,439,465]" pageId="11" pageNumber="493" type="holotype">Holotype</typeStatus>
: TL = 
<quantity id="702934A5250E897EFD80FE728EDEFE34" box="[564,650,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="493" unit="mm" value="3.9">3.9 mm</quantity>
, TL without head = 
<quantity id="702934A5250E897EFCD1FE728FE8FE34" box="[869,956,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="493" unit="mm" value="3.7">3.7 mm</quantity>
, MW = 
<quantity id="702934A5250E897EFBA7FE72883EFE34" box="[1043,1130,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="493" unit="mm" value="2.1">2.1 mm</quantity>
, ratio TL/MW = 1.83. 
<typeStatus id="686A27E2250E897EFAD5FE728C82FE10" pageId="11" pageNumber="493" type="paratype">Paratypes</typeStatus>
: TL = 
<quantity id="702934A5250E897EFEABFE1E8DF2FE10" box="[287,422,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="11" pageNumber="493" unit="mm" value="3.9" valueMax="4.2" valueMin="3.6">3.6–4.2 mm</quantity>
(3.91±0.17), TL without head = 3.4–4.0 mm (3.74±0.16), MW = 2.0– 
<quantity id="702934A5250E897EFB01FE1E895FFE10" box="[1205,1291,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="493" unit="mm" value="2.4">2.4 mm</quantity>
(2.18±0.10), TL/MW = 1.71–1.94 (1.79±0.04).
</paragraph>
</subSubSection>
<subSubSection id="FFCBCACB250E897EFF73FDE7882CF9B4" pageId="11" pageNumber="493" type="diagnosis">
<paragraph id="B76E9940250E897EFF73FDE78EE6FD60" blockId="11.[151,1437,150,645]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FDE78D9AFDD9" bold="true" box="[199,462,546,572]" pageId="11" pageNumber="493">Differential diagnosis.</emphasis>
This species is close to 
<taxonomicName id="70D1E2C3250E897EFD68FDE68F03FDD8" authorityName="Regimbart" authorityYear="1900" box="[732,855,547,573]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFD68FDE68F03FDD8" box="[732,855,547,573]" italics="true" pageId="11" pageNumber="493">L. alluaudi</emphasis>
</taxonomicName>
particularly with respect to the shape of the median lobe of aedeagus (compare 
<figureCitation id="2FEA85C5250E897EFE67FD828E0FFD84" box="[467,603,583,609]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Figs. 11, 18</figureCitation>
and 
<figureCitation id="2FEA85C5250E897EFD21FD828F26FD84" box="[661,882,582,609]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Figs. 12–15, 19–22</figureCitation>
) and parameres (compare 
<figureCitation id="2FEA85C5250E897EFB10FD82897EFD84" box="[1188,1322,583,609]" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="11" pageNumber="493">Figs. 27–28</figureCitation>
and 
<figureCitation id="2FEA85C5250E897EFAD1FD828CB6FD60" captionStart="FIGURES 23–30" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[157,1431,561,1885]" captionTargetId="figure-175@10.[157,1431,561,1885]" captionTargetPageId="10" captionText="FIGURES 23–30. Left (23, 25, 27, 29) and right (24, 26, 28, 30) parameres of: (23, 24) Laccophilus leguyaderi sp. nov. (holotype); (25, 26) Laccophilus bergsteni sp. nov. (holotype); (27, 28) Laccophilus makay sp. nov. (holotype); (29, 30) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401795" httpUri="https://zenodo.org/record/4401795/files/figure.png" pageId="11" pageNumber="493">Figs. 29–30</figureCitation>
). However, it differs from 
<taxonomicName id="70D1E2C3250E897EFDB8FDAE8ED2FD60" authorityName="Regimbart" authorityYear="1900" box="[524,646,619,645]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFDB8FDAE8ED2FD60" box="[524,646,619,645]" italics="true" pageId="11" pageNumber="493">L. alluaudi</emphasis>
</taxonomicName>
by:
</paragraph>
<paragraph id="B76E9940250E897EFF23FD768D27FD14" blockId="11.[151,1436,690,1041]" pageId="11" pageNumber="493">
- slightly larger size (
<emphasis id="85A54552250E897EFE3DFD768E5AFD28" box="[393,526,691,717]" italics="true" pageId="11" pageNumber="493">
<taxonomicName id="70D1E2C3250E897EFE3DFD768E5CFD28" authorityName="Regimbart" authorityYear="1900" box="[393,520,691,717]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">L. alluaudi</taxonomicName>
:
</emphasis>
specimens examined in this study, TL = 
<quantity id="702934A5250E897EFC6BFD7788D1FD28" box="[991,1157,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.85" metricValueMin="3.25" pageId="11" pageNumber="493" unit="mm" value="3.55" valueMax="3.85" valueMin="3.25">3.25–3.85 mm</quantity>
; 
<bibRefCitation id="D340E4B1250E897EFB25FD768924FD28" author="Bistrom, O. &amp; Nilsson, A. N. &amp; Bergsten, J." box="[1169,1392,690,717]" pageId="11" pageNumber="493" pagination="1 - 379" refId="ref13976" refString="Bistrom, O., Nilsson, A. N. &amp; Bergsten, J. (2015) Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae). ZooKeys, 542, 1 - 379. https: // doi. org / 10.3897 / zookeys. 542.5975" type="journal article" year="2015">
Biström 
<emphasis id="85A54552250E897EFB47FD768979FD28" box="[1267,1325,691,717]" italics="true" pageId="11" pageNumber="493">et al.</emphasis>
2015
</bibRefCitation>
: TL = 
<quantity id="702934A5250E897EFF69FD128D3CFD14" box="[221,360,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="11" pageNumber="493" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 mm</quantity>
),
</paragraph>
<paragraph id="B76E9940250E897EFF23FD3E8E54FCB8" blockId="11.[151,1436,690,1041]" pageId="11" pageNumber="493">
- elytral colour pattern fainter, with pale area between anterior margin of elytron and anterior extremity of dark longitudinal markings distinctly broader, and with no basal dark marking in shoulder area (present in 
<taxonomicName id="70D1E2C3250E897EFAACFCDA89C7FCDC" authorityName="Regimbart" authorityYear="1900" box="[1304,1427,799,825]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFAACFCDA89C7FCDC" box="[1304,1427,799,825]" italics="true" pageId="11" pageNumber="493">L. alluaudi</emphasis>
</taxonomicName>
) (compare 
<figureCitation id="2FEA85C5250E897EFE83FC868D2CFCB8" box="[311,376,835,861]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="11" pageNumber="493">Fig. 3</figureCitation>
and 
<figureCitation id="2FEA85C5250E897EFE1AFC868DA6FCB8" box="[430,498,835,861]" captionStart="FIGURES 1–4" captionStartId="6.[151,264,1939,1963]" captionTargetBox="[185,1405,193,1915]" captionTargetId="figure-17@6.[157,1431,181,1915]" captionTargetPageId="6" captionText="FIGURES 1–4. Habitus in dorsal view of: (1) Laccophilus leguyaderi sp. nov. (holotype); (2) Laccophilus bergsteni sp. nov. (holotype); (3) Laccophilus makay sp. nov. (holotype); (4) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401789" httpUri="https://zenodo.org/record/4401789/files/figure.png" pageId="11" pageNumber="493">Fig. 4</figureCitation>
),
</paragraph>
<paragraph id="B76E9940250E897EFF23FCA28DDBFC40" blockId="11.[151,1436,690,1041]" pageId="11" pageNumber="493">
- ventral surface entirely testaceous (in 
<taxonomicName id="70D1E2C3250E897EFDE0FCA28E9BFC64" authorityName="Regimbart" authorityYear="1900" box="[596,719,871,897]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFDE0FCA28E9BFC64" box="[596,719,871,897]" italics="true" pageId="11" pageNumber="493">L. alluaudi</emphasis>
</taxonomicName>
, metacoxal plates always largely dark brown to black; compare 
<figureCitation id="2FEA85C5250E897EFF72FC4E8D53FC40" box="[198,263,907,933]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="11" pageNumber="493">Fig. 7</figureCitation>
and 
<figureCitation id="2FEA85C5250E897EFE8AFC4E8DD5FC40" box="[318,385,907,933]" captionStart="FIGURES 5–8" captionStartId="7.[151,264,1896,1920]" captionTargetBox="[159,1418,196,1873]" captionTargetId="figure-26@7.[150,1437,181,1873]" captionTargetPageId="7" captionText="FIGURES 5–8. Ventral surface of: (5) Laccophilus leguyaderi sp. nov. (holotype); (6) Laccophilus bergsteni sp. nov. (holotype); (7) Laccophilus makay sp. nov. (holotype); (8) Laccophilus alluaudi (male specimen from Isalo massif). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401791" httpUri="https://zenodo.org/record/4401791/files/figure.png" pageId="11" pageNumber="493">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="B76E9940250E897EFF23FC6A8E0FFBF4" blockId="11.[151,1436,690,1041]" pageId="11" pageNumber="493">
- median lobe in lateral view with basal piece proportionally larger and with keel (absent in southern populations of 
<taxonomicName id="70D1E2C3250E897EFF72FC168D14FC08" authorityName="Regimbart" authorityYear="1900" box="[198,320,979,1005]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFF72FC168D14FC08" box="[198,320,979,1005]" italics="true" pageId="11" pageNumber="493">L. alluaudi</emphasis>
</taxonomicName>
; compare 
<figureCitation id="2FEA85C5250E897EFE01FC168E57FC08" box="[437,515,979,1005]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Fig. 11</figureCitation>
with 
<figureCitation id="2FEA85C5250E897EFDF6FC168E9DFC08" box="[578,713,978,1005]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Figs. 13–15</figureCitation>
), and in ventral view with apical quarter more strongly narrowed (compare 
<figureCitation id="2FEA85C5250E897EFE83FC328DD2FBF4" box="[311,390,1015,1041]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Fig. 18</figureCitation>
with 
<figureCitation id="2FEA85C5250E897EFE72FC328E1BFBF4" box="[454,591,1015,1041]" captionStart="FIGURES 9–22" captionStartId="8.[151,264,1906,1930]" captionTargetBox="[184,1403,181,1882]" captionTargetId="figure-17@8.[184,1403,181,1882]" captionTargetPageId="8" captionText="FIGURES 9–22. Median lobe of aedeagus in lateral (9–15) and ventral (16–22) view of: (9, 16) Laccophilus leguyaderi sp. nov. (holotype); (10, 17) Laccophilus bergsteni sp. nov. (holotype); (11, 18) Laccophilus makay sp. nov. (holotype); Laccophilus alluaudi, male specimens from (12, 19) Montagne d’Ambre, (13, 14, 20, 21) Isalo massif, and (15, 22) Ibity massif. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401793" httpUri="https://zenodo.org/record/4401793/files/figure.png" pageId="11" pageNumber="493">Figs. 19–22</figureCitation>
).
</paragraph>
<paragraph id="B76E9940250E897EFF73FBFA8DD4FB0C" blockId="11.[151,1437,1087,1725]" pageId="11" pageNumber="493">
In external morphology, 
<taxonomicName id="70D1E2C3250E897EFE6FFBFA8E16FBBC" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[475,578,1087,1113]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="makay">
<emphasis id="85A54552250E897EFE6FFBFA8E16FBBC" box="[475,578,1087,1113]" italics="true" pageId="11" pageNumber="493">L. makay</emphasis>
</taxonomicName>
shares with 
<taxonomicName id="70D1E2C3250E897EFD7AFBFA8F1DFBBD" authorityName="Guignot" authorityYear="1955" box="[718,841,1087,1112]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="pseustes">
<emphasis id="85A54552250E897EFD7AFBFA8F1DFBBD" box="[718,841,1087,1112]" italics="true" pageId="11" pageNumber="493">L. pseustes</emphasis>
</taxonomicName>
the entirely testaceous colouration of the ventral surface (all other known 
<taxonomicName id="70D1E2C3250E897EFE3EFBA68DB3FB98" authorityName="Regimbart" authorityYear="1900" box="[394,487,1123,1149]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="85A54552250E897EFE3EFBA68DB3FB98" box="[394,487,1123,1149]" italics="true" pageId="11" pageNumber="493">alluaudi</emphasis>
</taxonomicName>
-group species have the metacoxal plates darkened to a variable extent) and the rather faint elytral colouration pattern. However, in 
<taxonomicName id="70D1E2C3250E897EFD2DFB428F54FB44" authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" box="[665,768,1159,1185]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="makay">
<emphasis id="85A54552250E897EFD2DFB428F54FB44" box="[665,768,1159,1185]" italics="true" pageId="11" pageNumber="493">L. makay</emphasis>
</taxonomicName>
the dark elytral vittae end farther from the anterior margin of elytron. The median lobes of these two species have very different shapes (for 
<taxonomicName id="70D1E2C3250E897EFB86FB6E88FAFB21" authorityName="Guignot" authorityYear="1955" box="[1074,1198,1195,1220]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="493" phylum="Arthropoda" rank="species" species="pseustes">
<emphasis id="85A54552250E897EFB86FB6E88FAFB21" box="[1074,1198,1195,1220]" italics="true" pageId="11" pageNumber="493">L. pseustes</emphasis>
</taxonomicName>
, see Fig. 
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<bibRefCitation id="D340E4B1250E897EFF23FB0A8D20FB0C" author="Bistrom, O. &amp; Nilsson, A. N. &amp; Bergsten, J." box="[151,372,1230,1257]" pageId="11" pageNumber="493" pagination="1 - 379" refId="ref13976" refString="Bistrom, O., Nilsson, A. N. &amp; Bergsten, J. (2015) Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae). ZooKeys, 542, 1 - 379. https: // doi. org / 10.3897 / zookeys. 542.5975" type="journal article" year="2015">
Biström 
<emphasis id="85A54552250E897EFF4CFB0A8D7EFB0C" box="[248,298,1231,1257]" italics="true" pageId="11" pageNumber="493">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
<paragraph id="B76E9940250E897EFF73FB37882CF9B4" blockId="11.[151,1437,1087,1725]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73FB378D7EFAE9" bold="true" box="[199,298,1266,1292]" pageId="11" pageNumber="493">Habitat.</emphasis>
This species was collected at an altitude range of 
<quantity id="702934A5250E897EFCE1FB368F82FAE8" box="[853,982,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="5.8" metricValueMax="6.7" metricValueMin="4.9" pageId="11" pageNumber="493" unit="m" value="580.0" valueMax="670.0" valueMin="490.0">
490– 
<specimenCount id="A1D752C9250E897EFC39FB368F82FAE8" box="[909,982,1267,1293]" pageId="11" pageNumber="493" type="generic">670 m</specimenCount>
</quantity>
. It inhabits streams and stream-fed pools or ponds (
<figureCitation id="2FEA85C5250E897EFEBCFAD28DC5FAD4" box="[264,401,1303,1329]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1648,1672]" captionTargetBox="[156,1431,181,1625]" captionTargetId="figure-17@16.[156,1431,181,1625]" captionTargetPageId="16" captionText="FIGURES 47–52. Habitats of Laccophilus makay sp. nov. (47–49) and Laccophilus rakouthae sp. nov. (50–52). (47) Spring on sandy bank of river Andranomanitsy (Makay massif, 500 m); (48) Large pool with clear water, sandy bottom and masses of tree roots (Makay massif, 550 m). (49) Shadowed pool with sandy bottom and lots of orange deposit, beneath rock wall in a canyon (Makay massif, 540 m; type locality of L. makay); (50) Small uncultivated rice paddy field, in agricultural landscape (Ankaratra massif, Manjakatompo, 1,650 m); (51) Large eutrophic pond full of helophytes, surrounded by degraded forest (Ankaratra massif, Manjakatompo, Étang des Tilapia, 1,700 m; type locality of L. rakouthae); (52) Close-up view of the margin of the same pond." figureDoi="http://doi.org/10.5281/zenodo.4401803" httpUri="https://zenodo.org/record/4401803/files/figure.png" pageId="11" pageNumber="493">Figs. 47–49</figureCitation>
), generally surrounded by gallery forest. Most collecting sites were at least partly shaded by trees (only one fully sun-exposed site). The bottom consisted of sand (5/11 sites), sand together with siliceous stones or rock mass (3/11 sites), or clay (3/11 sites). Organic matter in the substratum was generally moderately abundant (8/11 sites); at two sites it was inconspicuous and on the contrary at one site (in which the species was not abundant) it formed a thick cover on the bottom. Water was very clear (except at one site) and the pH was moderately acidic. At all sites, helophytes were absent, and in the vast majority of sites (9/11) there was no submerged aquatic vegetation. Three sampling sites were characterised by strong development of orange deposits (e.g. 
<figureCitation id="2FEA85C5250E897EFB76FA2A8941F9EC" box="[1218,1301,1519,1545]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1648,1672]" captionTargetBox="[156,1431,181,1625]" captionTargetId="figure-17@16.[156,1431,181,1625]" captionTargetPageId="16" captionText="FIGURES 47–52. Habitats of Laccophilus makay sp. nov. (47–49) and Laccophilus rakouthae sp. nov. (50–52). (47) Spring on sandy bank of river Andranomanitsy (Makay massif, 500 m); (48) Large pool with clear water, sandy bottom and masses of tree roots (Makay massif, 550 m). (49) Shadowed pool with sandy bottom and lots of orange deposit, beneath rock wall in a canyon (Makay massif, 540 m; type locality of L. makay); (50) Small uncultivated rice paddy field, in agricultural landscape (Ankaratra massif, Manjakatompo, 1,650 m); (51) Large eutrophic pond full of helophytes, surrounded by degraded forest (Ankaratra massif, Manjakatompo, Étang des Tilapia, 1,700 m; type locality of L. rakouthae); (52) Close-up view of the margin of the same pond." figureDoi="http://doi.org/10.5281/zenodo.4401803" httpUri="https://zenodo.org/record/4401803/files/figure.png" pageId="11" pageNumber="493">Fig. 49</figureCitation>
) and at one site moderately abundant green filamentous algae were observed. All sites in which this species was sampled are situated in a well-preserved area with no significant signs of anthropogenic perturbation.
</paragraph>
</subSubSection>
<subSubSection id="FFCBCACB250E897EFF73F99F894FF990" box="[199,1307,1626,1653]" pageId="11" pageNumber="493" type="distribution">
<paragraph id="B76E9940250E897EFF73F99F894FF990" blockId="11.[151,1437,1087,1725]" box="[199,1307,1626,1653]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73F99F8D35F991" bold="true" box="[199,353,1626,1652]" pageId="11" pageNumber="493">Distribution.</emphasis>
So far known only from the Makay massif in central-southern 
<collectingCountry id="CFC6D9D0250E897EFB97F99E88FEF990" box="[1059,1194,1627,1653]" name="Madagascar" pageId="11" pageNumber="493">Madagascar</collectingCountry>
(
<figureCitation id="2FEA85C5250E897EFB0EF99E8959F990" box="[1210,1293,1626,1653]" captionStart="FIGURE 53" captionStartId="18.[151,250,1530,1554]" captionTargetBox="[391,1197,597,1507]" captionTargetId="figure-243@18.[391,1197,597,1507]" captionTargetPageId="18" captionText="FIGURE 53. Map showing the distribution of known localities of the newly described species. Red dot: L. leguyaderi; Blue dots: L. bergsteni; Magenta dot: L. bergsteni and L. rakouthae; yellow dot: L. makay. Names of corresponding protected areas or massifs indicated in italics above dots." figureDoi="http://doi.org/10.5281/zenodo.4401805" httpUri="https://zenodo.org/record/4401805/files/figure.png" pageId="11" pageNumber="493">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FFCBCACB250E897EFF73F9BB8F25F958" pageId="11" pageNumber="493" type="etymology">
<paragraph id="B76E9940250E897EFF73F9BB8F25F958" blockId="11.[151,1437,1087,1725]" pageId="11" pageNumber="493">
<emphasis id="85A54552250E897EFF73F9BB8DF1F97D" bold="true" box="[199,421,1662,1688]" pageId="11" pageNumber="493">Derivatio nominis.</emphasis>
This species is named after the 
<typeStatus id="686A27E2250E897EFCA4F9BA8F14F97C" box="[784,832,1663,1689]" pageId="11" pageNumber="493">type</typeStatus>
locality, the Makay massif in central-southern 
<collectingCountry id="CFC6D9D0250E897EFAE7F9BA8CB5F958" name="Madagascar" pageId="11" pageNumber="493">Madagascar</collectingCountry>
. It is a noun in the nominative case standing in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>