226 lines
28 KiB
XML
226 lines
28 KiB
XML
<document id="4BA3B5E06890F3E6B99AA4CDDD063EBB" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFBE3C58FF57F88DFB956AC3" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada carsoniana Köhler 2011, n.sp." docType="treatment" docUuid="79C0B2D0-CEAD-461E-9DB6-717A14B3C5F6" docUuidSource="ZooBank" docVersion="3" lastPageNumber="173" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="172" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3B640BCD9AB9E88D38649ED3350EEB2C">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
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<mods:roleTerm id="BB8C0B502176FD6018402859DAA308B5">Author</mods:roleTerm>
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<mods:namePart id="28D9D1A6F9547BDFEFA90DFB8FA17A8C">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="AC60F0A7AF5DE8DDCD6B4BF11959DC57">text</mods:typeOfResource>
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<mods:title id="9DEDFAA7AADDAB5D680CA9ECB3D74F99">Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part id="6725C00964B334503BD91B059E40DD1A">
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<mods:date id="0BC57BCCD6B9B901832B29F9C8E57404">2011</mods:date>
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<mods:number id="134263759D8A9A75161B44AA1B671D9C">2011-11-30</mods:number>
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<mods:number id="2F0E532376207286A2B6CD16F3177299">63</mods:number>
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<mods:number id="D18037ABD2BAF9F993E0ECC4DFFD4CB6">2</mods:number>
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<mods:start id="EE6CDECBCA978D13721106517A4A8CB4">167</mods:start>
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<mods:url id="2B726C46B50249B8B353AA62C3C49DA1">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
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<mods:classification id="0F2C7F39A13E09EFBF02E7E755DCBB08">journal article</mods:classification>
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<mods:identifier id="1343E05BD3E205B2BEA108C659C8D997" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
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<mods:identifier id="BA3999A39D307B4C309060BECF7E24BB" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="0FBB997C6FA16A723E730B922569FBA3" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
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<treatment id="0384503AFFBE3C58FF57F88DFB956AC3" LSID="urn:lsid:zoobank.org:act:79C0B2D0-CEAD-461E-9DB6-717A14B3C5F6" httpUri="http://treatment.plazi.org/id/0384503AFFBE3C58FF57F88DFB956AC3" lastPageId="6" lastPageNumber="173" pageId="5" pageNumber="172">
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<subSubSection id="C337B2A7FFBE3C5BFF57F88DFD9B6933" box="[239,630,1887,1913]" pageId="5" pageNumber="172" type="nomenclature">
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<paragraph id="8B92E12CFFBE3C5BFF57F88DFD9B6933" blockId="5.[239,630,1887,1913]" box="[239,630,1887,1913]" pageId="5" pageNumber="172">
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<heading id="D0DA5640FFBE3C5BFF57F88DFD9B6933" box="[239,630,1887,1913]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="172" reason="2">
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<emphasis id="B9593D3EFFBE3C5BFF57F88DFD9B6933" bold="true" box="[239,630,1887,1913]" pageId="5" pageNumber="172">
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<taxonomicName id="4C2D9AAFFFBE3C5BFF57F88DFDDE6933" ID-CoL="66H53" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[239,563,1887,1913]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="172" phylum="Mollusca" rank="species" species="carsoniana" status="sp. nov.">
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<emphasis id="B9593D3EFFBE3C5BFF57F88DFDDE6933" bold="true" box="[239,563,1887,1913]" italics="true" pageId="5" pageNumber="172">Amplirhagada carsoniana</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A26A8045FFBE3C5BFD82F88DFD9B6933" box="[570,630,1887,1913]" pageId="5" pageNumber="172" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C337B2A7FFBE3C5BFFC1F849FB2169C9" pageId="5" pageNumber="172" type="materials_examined">
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<paragraph id="8B92E12CFFBE3C5BFFC1F849FD00699A" blockId="5.[121,749,1947,2000]" pageId="5" pageNumber="172">
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<materialsCitation id="3B45EB71FFBE3C5BFFC1F849FD04699A" collectorName="NW Kimberley" location="Face Point" pageId="5" pageNumber="172" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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||
<emphasis id="B9593D3EFFBE3C5BFFC1F849FEE369F9" bold="true" box="[121,270,1947,1971]" pageId="5" pageNumber="172">
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<typeStatus id="54965F8EFFBE3C5BFFC1F849FF5F69F9" box="[121,178,1947,1971]" pageId="5" pageNumber="172">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingRegion id="49E92FCEFFBE3C5BFEA4F84EFE0D69F9" box="[284,480,1948,1971]" country="Australia" name="Western Australia" pageId="5" pageNumber="172">Western Australia</collectingRegion>
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,
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<collectorName id="26D884FAFFBE3C5BFE54F84EFD7969F9" box="[492,660,1948,1971]" pageId="5" pageNumber="172">NW Kimberley</collectorName>
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||
, Carson Escarpment,
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<quantity id="4CD54CC9FFBE3C5BFEBEF86BFEBC699A" box="[262,337,1977,2000]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="172" unit="km" value="2.5">2.5 km</quantity>
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N of
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||
<location id="8EF2B7F7FFBE3C5BFE2CF86BFDE4699A" LSID="urn:lsid:plazi:treatment:0384503AFFBE3C58FF57F88DFB956AC3:8EF2B7F7FFBE3C5BFE2CF86BFDE4699A" box="[404,521,1977,2000]" name="Face Point" pageId="5" pageNumber="172" stateProvince="Western Australia">Face Point</location>
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, base of escarpment
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</materialsCitation>
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||
;
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</paragraph>
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<paragraph id="8B92E12CFFBE3C5BFCA2F974FC0C6891" blockId="5.[794,1421,1702,1755]" pageId="5" pageNumber="172">
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<materialsCitation id="3B45EB71FFBE3C5BFCA2F974FC316891" collectingDate="1987-06-10" collectionCode="V" latitude="-14.838888" longLatPrecision="21" longitude="126.81944" pageId="5" pageNumber="172" specimenCode="RFS 10-4" specimenCount="1">
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<geoCoordinate id="EE1987EBFFBE3C5BFCA2F974FC7E68F7" box="[794,915,1702,1725]" degrees="14" direction="south" minutes="50" orientation="latitude" pageId="5" pageNumber="172" precision="15" seconds="20" value="-14.838888">14°50'20"S</geoCoordinate>
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||
<geoCoordinate id="EE1987EBFFBE3C5BFC25F974FBCB68F7" box="[925,1062,1702,1725]" degrees="126" direction="east" minutes="49" orientation="longitude" pageId="5" pageNumber="172" precision="15" seconds="10" value="126.81944">126°49'10"E</geoCoordinate>
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||
(
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||
<specimenCode id="DB8B4957FFBE3C5BFB8FF974FB4B68F7" box="[1079,1190,1702,1725]" collectionCode="RFS" pageId="5" pageNumber="172">RFS 10-4</specimenCode>
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||
, coll.
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||
<collectionCode id="ED3C79E9FFBE3C5BFB54F974FB1268F7" box="[1260,1279,1702,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="172" type="Museum">V</collectionCode>
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||
. Kessner,
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||
<date id="FF93C7ECFFBE3C5BFACAF974FC916891" pageId="5" pageNumber="172" value="1987-06-10">
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<collectingDate id="EFD73E04FFBE3C5BFACAF974FC916891" pageId="5" pageNumber="172" value="1987-06-10">10 Jun 1987</collectingDate>
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||
</date>
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||
) (
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<figureCitation id="1316FDA9FFBE3C5BFC2AF916FC3C6891" box="[914,977,1732,1755]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="5" pageNumber="172">Fig. 1</figureCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B92E12CFFBE3C5BFCA2F924FB2169C9" blockId="5.[794,1422,1782,1923]" pageId="5" pageNumber="172">
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||
<emphasis id="B9593D3EFFBE3C5BFCA2F924FC236944" bold="true" box="[794,974,1782,1806]" pageId="5" pageNumber="172">Type material</emphasis>
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||
.
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||
<materialsCitation id="3B45EB71FFBE3C5BFC5AF924FB8E6961" accessionNumber="S34707" collectionCode="WAM" pageId="5" pageNumber="172" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54965F8EFFBE3C5BFC5AF924FBBD6947" box="[994,1104,1782,1805]" pageId="5" pageNumber="172" type="holotype">Holotype</typeStatus>
|
||
<collectionCode id="ED3C79E9FFBE3C5BFBE3F924FB4D6947" box="[1115,1184,1782,1805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="172" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFBE3C5BFB14F924FAEB6947" box="[1196,1286,1782,1805]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34707" pageId="5" pageNumber="172" type="EnaNcbi">S34707</accessionNumber>
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||
(preserved specimen) (Pl. 1.2;
|
||
<tableCitation id="C6AFD497FFBE3C5BFBB9F8C6FBBA6961" box="[1025,1111,1812,1835]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximum–minimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="5" pageNumber="172">Table 1</tableCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B45EB71FFBE3C5BFBCBF8C6FC166902" accessionNumber="S34708" collectionCode="WAM" pageId="5" pageNumber="172" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54965F8EFFBE3C5BFBCBF8C6FB0D6961" box="[1139,1248,1812,1835]" pageId="5" pageNumber="172" type="paratype">Paratypes</typeStatus>
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||
<collectionCode id="ED3C79E9FFBE3C5BFB53F8C6FAC36961" box="[1259,1326,1812,1835]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="172" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFBE3C5BFA80F8C6FA636961" box="[1336,1422,1812,1835]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34708" pageId="5" pageNumber="172" type="EnaNcbi">S34708</accessionNumber>
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(preserved specimen)
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||
</materialsCitation>
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||
,
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<materialsCitation id="3B45EB71FFBE3C5BFBBBF8E3FA646902" box="[1027,1417,1841,1864]" collectionCode="FMNH" pageId="5" pageNumber="172" specimenCode="FMNH 220344" specimenCount="1" typeStatus="paratype">
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||
<specimenCode id="DB8B4957FFBE3C5BFBBBF8E3FB4E6902" box="[1027,1187,1841,1864]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="172" type="Museum">FMNH 220344</specimenCode>
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||
(preserved specimen)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B45EB71FFBE3C5BFCA2F89CFC8F69C9" accessionNumber="S34709" collectionCode="AM, WAM" pageId="5" pageNumber="172" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED3C79E9FFBE3C5BFCA2F89CFCA8692F" box="[794,837,1870,1893]" country="Australia" name="Australian Museum" pageId="5" pageNumber="172" type="Museum">AM</collectionCode>
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||
C.472920 (5 dried shells),
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<collectionCode id="ED3C79E9FFBE3C5BFBC6F89CFB2D692F" box="[1150,1216,1870,1893]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="172" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFBE3C5BFB72F89CFAF0692F" box="[1226,1309,1870,1893]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34709" pageId="5" pageNumber="172" type="EnaNcbi">S34709</accessionNumber>
|
||
(12 dried shells)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B45EB71FFBE3C5BFCD5F8BEFB2569C9" accessionNumber="S34709" box="[877,1224,1900,1923]" collectionCode="FMNH" isTailEnumeration="true" pageId="5" pageNumber="172" specimenCode="FMNH 220343" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB8B4957FFBE3C5BFCD5F8BEFBFD69C9" box="[877,1040,1900,1923]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="172" type="Museum">FMNH 220343</specimenCode>
|
||
(17 dried shells)
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337B2A7FFBE3C5BFCA2F84CFBFD6999" pageId="5" pageNumber="172" type="etymology">
|
||
<paragraph id="8B92E12CFFBE3C5BFCA2F84CFBFD6999" blockId="5.[794,1421,1950,2003]" pageId="5" pageNumber="172">
|
||
<emphasis id="B9593D3EFFBE3C5BFCA2F84CFC7869FC" bold="true" box="[794,917,1950,1974]" pageId="5" pageNumber="172">Etymology</emphasis>
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||
. In reference to the Carson Escarpment, where this species was found.
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||
</paragraph>
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||
</subSubSection>
|
||
<caption id="DF52B1A4FFBD3C58FF1EFD4AFCF66C82" ID-DOI="http://doi.org/10.5281/zenodo.5239559" ID-Zenodo-Dep="5239559" httpUri="https://zenodo.org/record/5239559/files/figure.png" pageId="6" pageNumber="173" startId="6.[166,231,664,685]" targetBox="[187,769,158,638]" targetPageId="6">
|
||
<paragraph id="8B92E12CFFBD3C58FF1EFD4AFCF66C82" blockId="6.[166,795,664,712]" pageId="6" pageNumber="173">
|
||
Figure 6. Genitalia of
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FE36FD4AFD786CE7" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[398,661,664,685]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="carsoniana" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFBD3C58FE36FD4AFD786CE7" box="[398,661,664,685]" italics="true" pageId="6" pageNumber="173">Amplirhagada carsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFBD3C58FD26FD4AFD206CE7" box="[670,717,664,685]" pageId="6" pageNumber="173" rank="species">n.sp.</taxonomicNameLabel>
|
||
(WAM
|
||
<accessionNumber id="947E7CCFFFBD3C58FF1EFD61FF1E6C82" box="[166,243,691,712]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34708" pageId="6" pageNumber="173" type="EnaNcbi">S34708</accessionNumber>
|
||
). For labelling of structures see Fig. 3. Scale bar = 5 mm.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C337B2A7FFBD3C58FE2EFCD4FDF2694F" pageId="6" pageNumber="173" type="description">
|
||
<paragraph id="8B92E12CFFBD3C58FE2EFCD4FDC76D6A" blockId="6.[406,554,774,800]" box="[406,554,774,800]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FE2EFCD4FDC76D6A" bold="true" box="[406,554,774,800]" pageId="6" pageNumber="173">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFBD3C58FF1EFC91FD346ABF" blockId="6.[166,795,835,1797]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FF1EFC91FF366D10" box="[166,219,835,858]" italics="true" pageId="6" pageNumber="173">Shell</emphasis>
|
||
(Pl. 1.2;
|
||
<figureCitation id="1316FDA9FFBD3C58FE86FC91FE556D10" box="[318,440,835,858]" captionStart="Figure 5" captionStartId="5.[121,184,1558,1579]" captionTargetBox="[121,1421,151,1538]" captionTargetId="figure-61@5.[121,1421,151,1538]" captionTargetPageId="5" captionText="Figure 5. Amplirhagada carsoniana n.sp. SEM of shell (AM C.472920), radula and jaw (WAM S34708). (A) Protoconch viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) First teleoconch whorls viewed obliquely from above. (D) Jaw. (E) Central and lateral teeth viewed from above. (F) Close-up of marginal teeth. (G) Inner and central marginal teeth viewed from above. Scale bars: A–C, 200 µm; D, 100 µm; E–G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239557" httpUri="https://zenodo.org/record/5239557/files/figure.png" pageId="6" pageNumber="173">Fig. 5 A–C</figureCitation>
|
||
). Broadly conical with low spire, almost discoid. Thin to delicate, translucent. Periphery rounded to slightly angulate; upper and basal sectors of whorls well rounded. Umbilicus 10–30% concealed by columellar reflection. Background colour brownish horn; peripheral band absent or well marked, thin, brown, visible on most whorls; sub-sutural diffuse, thin, visible on most whorls; outer and inner lip colour whitish. Protoconch
|
||
<quantity id="4CD54CC9FFBD3C58FD40FBC2FF226A0F" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="173" unit="mm" value="2.6">2.6 mm</quantity>
|
||
in diameter, comprising about one whorl, with weak radially elongated pustulations. Teleoconch with faint, regular axial growth lines, evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, expanded, weakly reflected; basal node absent. Parietal wall of inner lip absent or very inconspicuous. Average shell size 9.8±0.6 × 16.3±
|
||
<quantity id="4CD54CC9FFBD3C58FDA1FB0CFD866ABF" box="[537,619,1246,1269]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="173" unit="mm" value="0.8">0.8 mm</quantity>
|
||
(
|
||
<tableCitation id="C6AFD497FFBD3C58FDC1FB0CFD246ABF" box="[633,713,1246,1269]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximum–minimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="6" pageNumber="173">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF52B1A4FFBD3C58FCFFFC77FA516D9F" ID-DOI="http://doi.org/10.5281/zenodo.5239561" ID-Zenodo-Dep="5239561" httpUri="https://zenodo.org/record/5239561/files/figure.png" pageId="6" pageNumber="173" startId="6.[839,901,933,954]" targetBox="[878,1456,161,900]" targetPageId="6">
|
||
<paragraph id="8B92E12CFFBD3C58FCFFFC77FA516D9F" blockId="6.[839,1468,933,981]" pageId="6" pageNumber="173">
|
||
Figure 7. Penial anatomy of
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FBF4FC77FAAE6DF0" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[1100,1347,933,954]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="carsoniana" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFBD3C58FBF4FC77FAAE6DF0" box="[1100,1347,933,954]" italics="true" pageId="6" pageNumber="173">Amplirhagada carsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFBD3C58FAF0FC77FA986DF0" box="[1352,1397,933,954]" pageId="6" pageNumber="173" rank="species">n.sp.</taxonomicNameLabel>
|
||
(WAM
|
||
<accessionNumber id="947E7CCFFFBD3C58FCFFFC12FC796D9F" box="[839,916,960,981]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34708" pageId="6" pageNumber="173" type="EnaNcbi">S34708</accessionNumber>
|
||
). For labelling of structures see Fig. 4. Scale bar = 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B92E12CFFBD3C58FF7DFB29FE136B07" blockId="6.[166,795,835,1797]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FF7DFB29FE1E6B58" box="[197,499,1275,1298]" italics="true" pageId="6" pageNumber="173">Radular and jaw morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFBD3C58FE47FB29FD9A6B58" box="[511,631,1275,1298]" captionStart="Figure 5" captionStartId="5.[121,184,1558,1579]" captionTargetBox="[121,1421,151,1538]" captionTargetId="figure-61@5.[121,1421,151,1538]" captionTargetPageId="5" captionText="Figure 5. Amplirhagada carsoniana n.sp. SEM of shell (AM C.472920), radula and jaw (WAM S34708). (A) Protoconch viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) First teleoconch whorls viewed obliquely from above. (D) Jaw. (E) Central and lateral teeth viewed from above. (F) Close-up of marginal teeth. (G) Inner and central marginal teeth viewed from above. Scale bars: A–C, 200 µm; D, 100 µm; E–G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239557" httpUri="https://zenodo.org/record/5239557/files/figure.png" pageId="6" pageNumber="173">Fig. 5 D–G</figureCitation>
|
||
). Tooth formula C + 12–14 + 4 + 18–20; average number of rows of teeth 130 (n = 2). Jaw with ten plates.
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFBD3C58FF7DFA81FE3468AD" blockId="6.[166,795,835,1797]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FF7DFA81FE746B20" box="[197,409,1363,1386]" italics="true" pageId="6" pageNumber="173">Genital morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFBD3C58FE1EFA81FDEB6B20" box="[422,518,1363,1386]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="6.[166,231,664,685]" captionStartId-1="6.[839,901,933,954]" captionTargetBox-0="[187,769,158,638]" captionTargetBox-1="[878,1456,161,900]" captionTargetId-0="figure-360@6.[176,780,136,644]" captionTargetId-1="figure-384@6.[865,1467,151,907]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionText-0="Figure 6. Genitalia of Amplirhagada carsoniana n.sp. (WAM S34708). For labelling of structures see Fig. 3. Scale bar = 5 mm." captionText-1="Figure 7. Penial anatomy of Amplirhagada carsoniana n.sp. (WAM S34708). For labelling of structures see Fig. 4. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239559" figureDoi-1="http://doi.org/10.5281/zenodo.5239561" httpUri-0="https://zenodo.org/record/5239559/files/figure.png" httpUri-1="https://zenodo.org/record/5239561/files/figure.png" pageId="6" pageNumber="173">Figs 6–7</figureCitation>
|
||
). Penis coiled within thick penial sheath; penial complex about as long as anterior part of oviduct. Length of penial retractor muscle equivalent to about half of length of penial complex. Penial verge extending about 1/5 of length of penial chamber, slender with pointed tip. Inner penial wall entirely covered by dense pustulation; pustules forming indistinct, corrugated pilasters. Main stimulatory pilaster indistinct, formed by prolonged pustules, comprising proximal half of inner penial chamber. Vas deferens entering penial sheath within proximal third of penial complex. Vagina moderately long, distally inflated. Bursa copulatrix short, reaching base of spermoviduct. Free oviduct coiled, shorter than vagina. Spermoviduct longer than anterior part of oviduct.
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFBD3C58FF7DF93CFDF2694F" blockId="6.[166,795,835,1797]" box="[197,543,1774,1797]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FF7DF93CFE7A694F" box="[197,407,1774,1797]" italics="true" pageId="6" pageNumber="173">Aestivation strategy</emphasis>
|
||
. Free sealer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337B2A7FFBD3C58FF1EF8FAFB956AC3" pageId="6" pageNumber="173" type="discussion">
|
||
<paragraph id="8B92E12CFFBD3C58FF1EF8FAFB956AC3" blockId="6.[166,795,1832,2002]" lastBlockId="6.[839,1467,1050,1161]" pageId="6" pageNumber="173">
|
||
<emphasis id="B9593D3EFFBD3C58FF1EF8FAFEE0690A" bold="true" box="[166,269,1832,1856]" pageId="6" pageNumber="173">Remarks</emphasis>
|
||
. Description based on dissection of
|
||
<specimenCount id="9D2B2AA5FFBD3C58FD3CF8FAFCFA6975" box="[644,791,1832,1855]" pageId="6" pageNumber="173" type="generic">one specimen</specimenCount>
|
||
. Listed by Solem (1991) as “
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FE67F894FD5A6917" authority="NSP" authorityName="NSP" box="[479,695,1862,1885]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9593D3EFFBD3C58FE67F894FD946917" box="[479,633,1862,1885]" italics="true" pageId="6" pageNumber="173">Amplirhagada</emphasis>
|
||
NSP
|
||
</taxonomicName>
|
||
18” to be distinct from
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FE84F8B1FD9A6930" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[316,631,1891,1914]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="drysdaleana">
|
||
<emphasis id="B9593D3EFFBD3C58FE84F8B1FE0C6930" box="[316,481,1891,1914]" italics="true" pageId="6" pageNumber="173">A. drysdaleana</emphasis>
|
||
<bibRefCitation id="EFBC9CDDFFBD3C58FE53F8B1FD9A6930" author="Solem, A" box="[491,631,1891,1914]" pageId="6" pageNumber="173" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
. Both taxa are indeed very similar but
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FE0DF853FDBC69D2" authorityName="Köhler" authorityYear="2011" box="[437,593,1921,1944]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="carsoniana">
|
||
<emphasis id="B9593D3EFFBD3C58FE0DF853FDBC69D2" box="[437,593,1921,1944]" italics="true" pageId="6" pageNumber="173">A. carsoniana</emphasis>
|
||
</taxonomicName>
|
||
differs by coiled, shorter penis with shorter main pilaster, no extended basal pilasters of inner penial wall and shorter vagina. Vas deferens entering sheath half way up in
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FBC5FBC8FAE06A7B" authorityName="Köhler" authorityYear="2011" box="[1149,1293,1050,1073]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="carsoniana">
|
||
<emphasis id="B9593D3EFFBD3C58FBC5FBC8FAE06A7B" box="[1149,1293,1050,1073]" italics="true" pageId="6" pageNumber="173">A. carsoniana</emphasis>
|
||
</taxonomicName>
|
||
but within upper sector of penial sheath in
|
||
<taxonomicName id="4C2D9AAFFFBD3C58FBD5FBEAFAFE6A05" authorityName="Solem" authorityYear="1981" box="[1133,1299,1080,1103]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="173" phylum="Mollusca" rank="species" species="drysdaleana">
|
||
<emphasis id="B9593D3EFFBD3C58FBD5FBEAFAFE6A05" box="[1133,1299,1080,1103]" italics="true" pageId="6" pageNumber="173">A. drysdaleana</emphasis>
|
||
</taxonomicName>
|
||
. Being similar overall and occurring in close proximity to each other, both species are likely sister taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |