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<document id="4F41BAD387150E1E559961F91D8265BF" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFB33C54FEA5FAD5FD3A6AF2" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada moraniana Köhler 2011, n.sp." docType="treatment" docUuid="AC1CBD2D-F8F3-4026-879A-A044D50F9B4C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="177" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="175" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="90089316A0C482DE93F7FAED52241BF7">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="4AD820E7E13E8157D097532AA6D91FF6" type="personal">
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<mods:namePart id="18ADDDBBE9B856A29B4CAAC34B0221D9">Köhler, Frank</mods:namePart>
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<mods:title id="06C110BA9644AB8A42F875533D4880E2">Records of the Australian Museum</mods:title>
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<mods:date id="B199D59F512E02BDB80294B141B07321">2011</mods:date>
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<mods:number id="7204EDC9E0879F734FB49D731766C8CB">2011-11-30</mods:number>
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<mods:number id="D3464B9B991DA50EF6C546D57B685CFA">63</mods:number>
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<mods:url id="9F451AE9D1C76BBCEF38AC6A21A103C8">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
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<mods:classification id="9F3D718C114880DDD2A2F5EE0D300D50">journal article</mods:classification>
<mods:identifier id="D3B6C55FF4227928A0FE0AC4EED057E8" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
<mods:identifier id="C7500475E40D8DCC2B5D9E6A52A71D2F" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="EB3BAF3E5470CD634BDCE78D274E2509" type="Zenodo-Dep">5239547</mods:identifier>
<mods:identifier id="DEEBB1676A91EB550E2361F135A179A0" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
</mods:mods>
<treatment id="0384503AFFB33C54FEA5FAD5FD3A6AF2" LSID="urn:lsid:zoobank.org:act:AC1CBD2D-F8F3-4026-879A-A044D50F9B4C" httpUri="http://treatment.plazi.org/id/0384503AFFB33C54FEA5FAD5FD3A6AF2" lastPageId="10" lastPageNumber="177" pageId="8" pageNumber="175">
<subSubSection id="C337B2A7FFB33C56FEA5FAD5FD4E6B6B" box="[285,675,1287,1313]" pageId="8" pageNumber="175" type="nomenclature">
<paragraph id="8B92E12CFFB33C56FEA5FAD5FD4E6B6B" blockId="8.[285,675,1287,1313]" box="[285,675,1287,1313]" pageId="8" pageNumber="175">
<heading id="D0DA5640FFB33C56FEA5FAD5FD4E6B6B" box="[285,675,1287,1313]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="175" reason="2">
<emphasis id="B9593D3EFFB33C56FEA5FAD5FD4E6B6B" bold="true" box="[285,675,1287,1313]" pageId="8" pageNumber="175">
<taxonomicName id="4C2D9AAFFFB33C56FEA5FAD5FDB26B6B" ID-CoL="D7RN" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[285,607,1287,1313]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="175" phylum="Mollusca" rank="species" species="moraniana" status="sp. nov.">
<emphasis id="B9593D3EFFB33C56FEA5FAD5FDB26B6B" bold="true" box="[285,607,1287,1313]" italics="true" pageId="8" pageNumber="175">Amplirhagada moraniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB33C56FDDFFAD5FD4E6B6B" box="[615,675,1287,1313]" pageId="8" pageNumber="175" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB33C56FF1EFAEBFD4A6812" pageId="8" pageNumber="175" type="materials_examined">
<paragraph id="8B92E12CFFB33C56FF1EFAEBFE836BE2" blockId="8.[166,794,1337,1448]" pageId="8" pageNumber="175">
<materialsCitation id="3B45EB71FFB33C56FF1EFAEBFE856BE2" collectingDate="1987-06-18" collectionCode="V" collectorName="NW Kimberley" latitude="-15.341666" location="Moran River" longLatPrecision="21" longitude="125.701385" pageId="8" pageNumber="175" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B9593D3EFFB33C56FF1EFAEBFED46B1B" bold="true" box="[166,313,1337,1361]" pageId="8" pageNumber="175">
<typeStatus id="54965F8EFFB33C56FF1EFAEBFF326B1B" box="[166,223,1337,1361]" pageId="8" pageNumber="175">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="49E92FCEFFB33C56FEFEFAEBFDEA6B1A" box="[326,519,1337,1360]" country="Australia" name="Western Australia" pageId="8" pageNumber="175">Western Australia</collectingRegion>
,
<collectorName id="26D884FAFFB33C56FDA9FAEBFD556B1A" box="[529,696,1337,1360]" pageId="8" pageNumber="175">NW Kimberley</collectorName>
,
<locationDeviation id="9A20F80EFFB33C56FD7AFAEBFDF56B27" location="Mitchell River Homestead" pageId="8" pageNumber="175">
<quantity id="4CD54CC9FFB33C56FD7AFAEBFCF76B1A" box="[706,794,1337,1360]" metricMagnitude="4" metricUnit="m" metricValue="2.56" pageId="8" pageNumber="175" unit="km" value="25.6">25.6 km</quantity>
SSW of Mitchell River Homestead
</locationDeviation>
, N bank of
<location id="8EF2B7F7FFB33C56FD36FA84FCFB6B27" LSID="urn:lsid:plazi:treatment:0384503AFFB33C54FEA5FAD5FD3A6AF2:8EF2B7F7FFB33C56FD36FA84FCFB6B27" box="[654,790,1366,1389]" latitude="-15.341666" longLatPrecision="21" longitude="125.701385" name="Moran River" pageId="8" pageNumber="175" stateProvince="Western Australia">Moran River</location>
,
<geoCoordinate id="EE1987EBFFB33C56FF1EFAA6FECD6BC1" box="[166,288,1396,1419]" degrees="15" direction="south" minutes="20" orientation="latitude" pageId="8" pageNumber="175" precision="15" seconds="30" value="-15.341666">15°20'30&quot;S</geoCoordinate>
<geoCoordinate id="EE1987EBFFB33C56FE92FAA6FE5E6BC1" box="[298,435,1396,1419]" degrees="125" direction="east" minutes="42" orientation="longitude" pageId="8" pageNumber="175" precision="15" seconds="05" value="125.701385">125°42'05&quot;E</geoCoordinate>
(RFS-18-2: coll.
<collectionCode id="ED3C79E9FFB33C56FDC0FAA6FD666BC1" box="[632,651,1396,1419]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="175" type="Museum">V</collectionCode>
. Kessner,
<date id="FF93C7ECFFB33C56FD46FAA6FEE46BE2" pageId="8" pageNumber="175" value="1987-06-18">
<collectingDate id="EFD73E04FFB33C56FD46FAA6FEE46BE2" pageId="8" pageNumber="175" value="1987-06-18">18 Jun 1987</collectingDate>
</date>
) (
<figureCitation id="1316FDA9FFB33C56FEA7FA43FEB36BE2" box="[287,350,1425,1448]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="8" pageNumber="175">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B92E12CFFB33C56FF1EFA19FD4A6812" blockId="8.[166,794,1483,1624]" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FF1EFA19FEB76BA9" bold="true" box="[166,346,1483,1507]" pageId="8" pageNumber="175">Type material</emphasis>
.
<materialsCitation id="3B45EB71FFB33C56FED7FA1EFE1D684A" accessionNumber="S34713" collectionCode="WAM" pageId="8" pageNumber="175" specimenCount="1" typeStatus="holotype">
<typeStatus id="54965F8EFFB33C56FED7FA1EFE306BA9" box="[367,477,1484,1507]" pageId="8" pageNumber="175" type="holotype">Holotype</typeStatus>
<collectionCode id="ED3C79E9FFB33C56FE50FA1EFDC06BA9" box="[488,557,1484,1507]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="175" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB33C56FD80FA1EFD7F6BA9" box="[568,658,1484,1507]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34713" pageId="8" pageNumber="175" type="EnaNcbi">S34713</accessionNumber>
(preserved specimen) (Pl. 1.4;
<tableCitation id="C6AFD497FFB33C56FE36FA3BFE0E684A" box="[398,483,1513,1536]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="8" pageNumber="175">Table 1</tableCitation>
)
</materialsCitation>
.
<materialsCitation id="3B45EB71FFB33C56FDB8FA3BFE2E6857" accessionNumber="S34714" collectionCode="WAM" pageId="8" pageNumber="175" specimenCount="1" typeStatus="paratype">
<typeStatus id="54965F8EFFB33C56FDB8FA3BFD80684A" box="[512,621,1513,1536]" pageId="8" pageNumber="175" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED3C79E9FFB33C56FDCFFA3BFD57684A" box="[631,698,1513,1536]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="175" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB33C56FD7DFA3BFCF7684A" box="[709,794,1513,1536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34714" pageId="8" pageNumber="175" type="EnaNcbi">S34714</accessionNumber>
(2 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB33C56FE6AF9D4FECD6871" collectionCode="FMNH" pageId="8" pageNumber="175" specimenCode="FMNH 220725" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB33C56FE6AF9D4FD6D6857" box="[466,640,1542,1565]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="175" type="Museum">FMNH 220725</specimenCode>
(3 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB33C56FE91F9F6FD976871" box="[297,634,1572,1595]" collectionCode="FMNH" pageId="8" pageNumber="175" specimenCode="FMNH 220724" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB33C56FE91F9F6FE266871" box="[297,459,1572,1595]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="175" type="Museum">FMNH 220724</specimenCode>
(20 dried shells)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB33C56FD3AF9F6FEBB6812" accessionNumber="S34715" collectionCode="WAM" pageId="8" pageNumber="175" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3C79E9FFB33C56FD3AF9F6FD2E6871" box="[642,707,1572,1595]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="175" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB33C56FD70F9F6FCF46871" box="[712,793,1572,1595]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34715" pageId="8" pageNumber="175" type="EnaNcbi">S34715</accessionNumber>
(15 dried shells)
</materialsCitation>
,
<collectionCode id="ED3C79E9FFB33C56FEE7F993FE676812" box="[351,394,1601,1624]" country="Australia" name="Australian Museum" pageId="8" pageNumber="175" type="Museum">AM</collectionCode>
C.472922 (6 dried shells).
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB33C56FF1EF9A9FD8268D9" box="[166,623,1659,1683]" pageId="8" pageNumber="175" type="etymology">
<paragraph id="8B92E12CFFB33C56FF1EF9A9FD8268D9" blockId="8.[166,623,1659,1683]" box="[166,623,1659,1683]" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FF1EF9A9FECC68D9" bold="true" box="[166,289,1659,1683]" pageId="8" pageNumber="175">Etymology</emphasis>
. Named after the Moran River.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB33C54FE2EF96EFDCD6DD8" lastPageId="10" lastPageNumber="177" pageId="8" pageNumber="175" type="description">
<paragraph id="8B92E12CFFB33C56FE2EF96EFDC7689C" blockId="8.[406,554,1724,1750]" box="[406,554,1724,1750]" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FE2EF96EFDC7689C" bold="true" box="[406,554,1724,1750]" pageId="8" pageNumber="175">Description</emphasis>
</paragraph>
<paragraph id="8B92E12CFFB33C56FF1EF923FA6A6BC0" blockId="8.[166,795,1777,2005]" lastBlockId="8.[839,1467,1249,2005]" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FF1EF923FF366942" box="[166,219,1777,1800]" italics="true" pageId="8" pageNumber="175">Shell</emphasis>
(Pl. 1.4;
<figureCitation id="1316FDA9FFB33C56FEFEF923FE3F6942" box="[326,466,1777,1800]" captionStart="Figure 11" captionStartId="10.[166,229,1846,1867]" captionText="Figure 11 (facing page). Amplirhagada moraniana n.sp. SEM of shell (AM C.472922), radula and jaw (WAM S34714). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) First teleoconch whorls viewed obliquely from above. (D) Close-up showing sculpture on last whorl, viewed from above. (E) Close-up showing sculpture on last whorl, lateral view. (F) Jaw. (G) Close-up showing detail of central and inner marginal teeth. (H) Close-up showing detail of central marginal teeth. (I) Central and inner lateral teeth viewed from above. (J) Marginal teeth viewed from above. Scale bars: AB, DE, 200 µm; C, 1 mm; F, 100 µm; GH, J, 10 µm; I, 20 µm." pageId="8" pageNumber="175">Fig. 11 AE</figureCitation>
). Broadly conical with low to moderately elevated spire; thin, translucent. Periphery well rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus 7090% concealed by columellar reflection. Background and ventral colour pale brownish horn; peripheral and sub-sutural bands usually absent or diffuse, visible on last whorl only; outer and inner lip whitish. Protoconch ~
<quantity id="4CD54CC9FFB33C56FE83F86CFE7F699F" box="[315,402,1982,2005]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="175" unit="mm" value="2.5">2.5 mm</quantity>
in diameter, comprising about 1.5 whorls, with radially elongated pustulations. Teleoconch with regular axial lirae, evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, slightly expanded, slightly reflected; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 10.9±1.5 × 15.9±
<quantity id="4CD54CC9FFB33C56FB71FAA1FAF46BC0" box="[1225,1305,1395,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="175" unit="mm" value="1.5">1.5 mm</quantity>
(
<tableCitation id="C6AFD497FFB33C56FA9FFAA1FA9A6BC0" box="[1319,1399,1395,1418]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="8" pageNumber="175">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B92E12CFFB33C56FCDDFA43FBCA6BA8" blockId="8.[839,1467,1249,2005]" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FCDDFA43FB776BE2" box="[869,1178,1425,1448]" italics="true" pageId="8" pageNumber="175">Radular and jaw morphology</emphasis>
(
<figureCitation id="1316FDA9FFB33C56FB1FFA43FAF96BE2" box="[1191,1300,1425,1448]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="8" pageNumber="175">Fig. 1 FJ</figureCitation>
). Tooth formula C + 1213 + 34 + 1618, with 120 rows of teeth (n = 1). Jaw with nine plates.
</paragraph>
<paragraph id="8B92E12CFFB33C54FCDDFA3BFCED6D3F" blockId="8.[839,1467,1249,2005]" lastBlockId="10.[166,795,803,915]" lastPageId="10" lastPageNumber="177" pageId="8" pageNumber="175">
<emphasis id="B9593D3EFFB33C56FCDDFA3BFBD7684A" box="[869,1082,1513,1536]" italics="true" pageId="8" pageNumber="175">Genital morphology</emphasis>
(
<figureCitation id="1316FDA9FFB33C56FBFEFA3BFB2D684A" box="[1094,1216,1513,1536]" captionStart-0="Figure 12" captionStart-1="Figure 13" captionStartId-0="10.[166,230,670,691]" captionStartId-1="10.[839,900,1162,1183]" captionTargetBox-0="[181,783,163,650]" captionTargetBox-1="[865,1400,162,1144]" captionTargetId-0="figure-323@10.[166,793,151,662]" captionTargetId-1="figure-346@10.[839,1422,149,1158]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionText-0="Figure 12. Genitalia of Amplirhagada moraniana n.sp. (WAM S34714). For labelling of structures see Fig. 3. Scale bar = 5 mm." captionText-1="Figure 13.Penial anatomy of Amplirhagada moraniana n.sp.(WAM S34714). For labelling of structures see Fig. 4. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239570" figureDoi-1="http://doi.org/10.5281/zenodo.5239572" httpUri-0="https://zenodo.org/record/5239570/files/figure.png" httpUri-1="https://zenodo.org/record/5239572/files/figure.png" pageId="8" pageNumber="175">Figs 1213</figureCitation>
). Penis straight, tubular, longer than anterior part of oviduct. Penial sheath very delicate. Length of penial retractor muscle very short, equivalent to about 1/10 of length of penial complex. Penial verge very long, extending about ¾ of length of penial chamber, slender with pointed tip. Inner penial wall almost entirely covered by reloped, rhomboid pustules that are arranged in honey-comb pattern. No main stimulatory pilaster differentiated; three to four longitudinal pilasters formed by fused, more elevated pustules are present, extending most of inner penial chamber, giving rise to corrugated longitudinal pilasters comprising distal end of inner penial wall. Vas deferens very thick, not undulating, entering penial sheath within proximal third of penial complex. Vagina long, tubular; inner vaginal wall and inner wall of bursa copulatrix with continuous, well-developed, smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct. Free oviduct comprising less than half of length of anterior part of oviduct. Spermoviduct about as long as anterior part of oviduct.
</paragraph>
<caption id="DF52B1A4FFB13C54FF1EFD4CFCF66C84" ID-DOI="http://doi.org/10.5281/zenodo.5239570" ID-Zenodo-Dep="5239570" httpUri="https://zenodo.org/record/5239570/files/figure.png" pageId="10" pageNumber="177" startId="10.[166,230,670,691]" targetBox="[181,783,163,650]" targetPageId="10">
<paragraph id="8B92E12CFFB13C54FF1EFD4CFCF66C84" blockId="10.[166,795,670,718]" pageId="10" pageNumber="177">
Figure 12. Genitalia of
<taxonomicName id="4C2D9AAFFFB13C54FE2FFD4DFD7B6CFE" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[407,662,671,692]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="177" phylum="Mollusca" rank="species" species="moraniana" status="sp. nov.">
<emphasis id="B9593D3EFFB13C54FE2FFD4DFD7B6CFE" box="[407,662,671,692]" italics="true" pageId="10" pageNumber="177">Amplirhagada moraniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB13C54FD18FD4CFD236CF9" box="[672,718,670,691]" pageId="10" pageNumber="177" rank="species">n.sp.</taxonomicNameLabel>
(WAM
<accessionNumber id="947E7CCFFFB13C54FF1EFD6BFF1E6C84" box="[166,243,697,718]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34714" pageId="10" pageNumber="177" type="EnaNcbi">S34714</accessionNumber>
). For labelling of structures see Fig. 3. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFB13C54FF7DFCAEFDCD6DD8" blockId="10.[166,795,803,915]" box="[197,544,891,915]" pageId="10" pageNumber="177">
<emphasis id="B9593D3EFFB13C54FF7DFCAEFE7A6DD9" box="[197,407,892,915]" italics="true" pageId="10" pageNumber="177">Aestivation strategy</emphasis>
: Free sealer.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB13C54FF1EFC64FD3A6AF2" pageId="10" pageNumber="177" type="discussion">
<paragraph id="8B92E12CFFB13C54FF1EFC64FD3A6AF2" blockId="10.[166,794,950,1208]" pageId="10" pageNumber="177">
<emphasis id="B9593D3EFFB13C54FF1EFC64FEE36D84" bold="true" box="[166,270,950,974]" pageId="10" pageNumber="177">Remarks</emphasis>
. Anatomical description based on dissection of
<specimenCount id="9D2B2AA5FFB13C54FF1EFC01FED06DA0" box="[166,317,979,1002]" pageId="10" pageNumber="177" type="generic">one specimen</specimenCount>
. Listed by Solem (1991) as “
<taxonomicName id="4C2D9AAFFFB13C54FD38FC06FF366A42" authority="NSP" authorityName="NSP" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="177" phylum="Mollusca" rank="genus">
<emphasis id="B9593D3EFFB13C54FD38FC06FCF76DA1" box="[640,794,980,1003]" italics="true" pageId="10" pageNumber="177">Amplirhagada</emphasis>
NSP
</taxonomicName>
20”. One of the smallest species with respect to shell height and diameter; a similar shell with respect to size and shape is only found in
<taxonomicName id="4C2D9AAFFFB13C54FE1CFBFEFDB76A09" authorityName="Köhler" authorityYear="2011" box="[420,602,1068,1091]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="177" phylum="Mollusca" rank="species" species="angustocauda">
<emphasis id="B9593D3EFFB13C54FE1CFBFEFDB76A09" box="[420,602,1068,1091]" italics="true" pageId="10" pageNumber="177">A. angustocauda</emphasis>
</taxonomicName>
described further below. Elongated shape of penis and markedly elongated penial verge (almost as long as penial chamber), corrugated penial wall sculpture, and very thick vas deferens is combination of features typical only for this species.
</paragraph>
</subSubSection>
</treatment>
</document>