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<mods:title id="DFC18EEA7664743DC6D688FF00D268BE">Taxonomic revision of Artines Godman, 1901 (Hesperiidae: Hesperiinae: Moncini) with the description of nine new species</mods:title>
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<mods:namePart id="45AEBD434D5579BDB9A29F193E1AB239">Medeiros, Adalberto Dantas De</mods:namePart>
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<mods:namePart id="D95B02C5C4F0F9EA9AC5B9331F1E4A56">Dolibaina, Diego Rodrigo</mods:namePart>
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<mods:namePart id="02DC63166C1957D020B18E05EFD6DBE7">Carneiro, Eduardo</mods:namePart>
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<mods:namePart id="8175000BDC7D9F9EB1AEF52AEC1DF61C">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<treatment id="CB52563B9D4ACA5DFF7CFECEFB97F9C5" ID-GBIF-Taxon="157436013" LSID="urn:lsid:plazi:treatment:CB52563B9D4ACA5DFF7CFECEFB97F9C5" httpUri="http://treatment.plazi.org/id/CB52563B9D4ACA5DFF7CFECEFB97F9C5" lastPageNumber="17" pageId="16" pageNumber="17">
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<paragraph id="4344E72D9D4ACA5DFF7CFECEFD60FEF0" blockId="16.[151,643,333,395]" box="[151,643,333,359]" pageId="16" pageNumber="17">
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<heading id="180C50419D4ACA5DFF7CFECEFD60FEF0" bold="true" box="[151,643,333,359]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="84FB9CAE9D4ACA5DFF7CFECEFD60FEF0" authority="Steinhauser & Austin, 1993" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[151,643,333,359]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
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<emphasis id="718F3B3F9D4ACA5DFF7CFECEFD60FEF0" bold="true" box="[151,643,333,359]" pageId="16" pageNumber="17">
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<emphasis id="718F3B3F9D4ACA5DFF7CFECEFEC5FEF0" bold="true" box="[151,294,333,359]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
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<bibRefCitation id="276A9ADC9D4ACA5DFEC6FECEFD60FEF0" author="Steinhauser, S. R. & Austin, G. T." box="[301,643,333,359]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin, 1993</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
|
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</subSubSection>
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<subSubSection id="0BE1B4A69D4ACA5DFF7CFEF2FE58FE1C" box="[151,443,368,395]" pageId="16" pageNumber="17" type="description">
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<paragraph id="4344E72D9D4ACA5DFF7CFEF2FE58FE1C" blockId="16.[151,643,333,395]" box="[151,443,368,395]" pageId="16" pageNumber="17">
|
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<figureCitation id="DBC0FBA89D4ACA5DFF7CFEF2FF08FE1C" box="[151,235,369,395]" captionStart="FIGURES 9–13" captionStartId="7.[151,264,1180,1205]" captionTargetBox="[152,1436,180,1162]" captionTargetId="figure@7.[151,1436,174,1169]" captionTargetPageId="7" captionText="FIGURES 9–13. Species of the “aepitus group” in dorsal and ventral views. 9. Artines maya, A–B: holotype male. 10. Artines litoralis sp. nov., A–B: holotype male, C–D: allotype female. 11. Artines rica, A–B: male, Turrialba, Cartago, Costa Rica (DZ 41.890), C–D: female from same locality of male (DZ 41.840). 12. Artines liege sp. nov., A–B: holotype male, C–D: allotype female. 13. Artines bamba sp. nov., A–B: holotype male, C–D: tentative female, Moyobamba, San Martin, Peru (OM 79.413). Scale bar = 1cm." httpUri="https://zenodo.org/record/3241043/files/figure.png" pageId="16" pageNumber="17">Figs 11</figureCitation>
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,
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<figureCitation id="DBC0FBA89D4ACA5DFF1CFEF2FEF6FE1C" box="[247,277,369,395]" captionStart="FIGURES 33–35" captionStartId="25.[151,264,1776,1801]" captionTargetBox="[151,1436,187,1743]" captionTargetId="figure@25.[151,1436,181,1750]" captionTargetPageId="25" captionText="FIGURES 33–35. Male genitalia of the “aepitus group” species. 33. Artines rica, Turrialba, Cartago, Costa Rica (DZ 4.061). 34. Artines liege sp. nov., holotype. 35. Artines bamba sp. nov., paratype, Pakitza, Parque Nacional del Manu, Madre de Díos, Peru (DZ 9.728). For reference of the letters see legend of the figure 27. Scale bar = 1 mm." httpUri="https://zenodo.org/record/3241051/files/figure.png" pageId="16" pageNumber="17">33</figureCitation>
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,
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<figureCitation id="DBC0FBA89D4ACA5DFECAFEF2FEA3FE1C" box="[289,320,369,395]" captionStart="FIGURES 42–48" captionStartId="33.[151,264,1892,1917]" captionTargetBox="[151,1431,625,1874]" captionTargetId="figure@33.[151,1436,625,1880]" captionText="FIGURES 42–48. Dorsal view of vesica and cornutus of the "aepitus group" species. 42. Artines aepitus. 43. Artines panama sp. nov.. 44. Artines delfos sp. nov.. 45. Artines litoralis sp. nov.. 46. Artines rica. 47. Artines liege sp. nov. 48. Artines bamba sp. nov. Scale bar = 0,2 mm." httpUri="https://zenodo.org/record/3406274/files/figure.png" pageId="16" pageNumber="17">46</figureCitation>
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, 58, 70, 78
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF7CFE35FF18FC81" pageId="16" pageNumber="17" type="reference_group">
|
||
<paragraph id="4344E72D9D4ACA5DFF7CFE35FC4DFD12" blockId="16.[151,1437,437,646]" pageId="16" pageNumber="17">
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||
<taxonomicName id="84FB9CAE9D4ACA5DFF7CFE35FDD3FE5A" authority="Steinhauser & Austin, 1993" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[151,560,437,462]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
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||
<emphasis id="718F3B3F9D4ACA5DFF7CFE35FEECFE5A" box="[151,271,438,461]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
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||
<bibRefCitation id="276A9ADC9D4ACA5DFEFEFE36FDD3FE5A" author="Steinhauser, S. R. & Austin, G. T." box="[277,560,437,462]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin, 1993</bibRefCitation>
|
||
</taxonomicName>
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||
.
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<emphasis id="718F3B3F9D4ACA5DFDD2FE36FD5DFE59" bold="true" box="[569,702,437,462]" pageId="16" pageNumber="17">Trop. Lep. 4</emphasis>
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, Suppl. 2: 14, figs 9, 10 (male d, v), 11, 12 (female d, v), 19 (male gen.), 20 (female gen.);
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<typeStatus id="9C40598F9D4ACA5DFE97FE50FE36FE7B" box="[380,469,467,492]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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male,
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<date id="3745C1ED9D4ACA5DFDF1FE57FD64FE7B" box="[538,647,468,492]" pageId="16" pageNumber="17" value="1972-06-01">1-VI-1972</date>
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, Turrialba, Província
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<collectingRegion id="813F29CF9D4ACA5DFC8FFE57FC54FE7B" box="[868,951,468,492]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
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,
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<collectingCountry id="3BECA7BD9D4ACA5DFC2AFE57FBD1FE7B" box="[961,1074,467,492]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
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, H. L. King
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<emphasis id="718F3B3F9D4ACA5DFB59FE57FB32FE7B" box="[1202,1233,468,492]" italics="true" pageId="16" pageNumber="17">leg</emphasis>
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.; AME [MGCL].—
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<bibRefCitation id="276A9ADC9D4ACA5DFF2CFE71FEB8FD9C" author="Bridges, C. A." box="[199,347,498,523]" pageId="16" pageNumber="17" refId="ref38783" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Vols. 1 - 13 & Apps. 1 - 3. Author, Urbana, 3. [11] + 598 pp. [11 pp., Vol. 1: 1 pp., Vol. 2: 1 pp., Vol. 3: 1 pp., Vol. 4: 36 pp., Vol. 5: 9 pp., Vol. 6: 20 pp., Vol. 7: 6 pp., Vol. 8: 246 pp., Vol. 9: 76 pp., Vol. 10: 90 pp., Vol. 11: 71 pp., Vol. 12: 28 pp., Vol. 13: 9 pp., App. 1: 1 pp., App. 2: 2 pp., App. 3: 1 pp.]" type="book" year="1994">Bridges, 1994</bibRefCitation>
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||
.
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<emphasis id="718F3B3F9D4ACA5DFE8DFE71FB88FD9C" bold="true" box="[358,1131,498,523]" pageId="16" pageNumber="17">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
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, p. 194;
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<emphasis id="718F3B3F9D4ACA5DFB21FE71FB34FD9C" bold="true" box="[1226,1239,498,523]" pageId="16" pageNumber="17">9</emphasis>
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, p. 6.—O.
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<bibRefCitation id="276A9ADC9D4ACA5DFAA5FE71FF1EFDBE" author="Mielke, O. H. H." pageId="16" pageNumber="17" pagination="3 - 11" refId="ref43459" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A, Hesperioidea-Papilionoidea. In: Heppner, J. (Ed.), Atlas of Neotropical Lepidoptera 5 A. Scientific Publishers, Association for Tropical Lepidoptera, Gainesville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
|
||
. Hesperioidea, p. 62,
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<emphasis id="718F3B3F9D4ACA5DFE0FFD92FE1AFDBE" bold="true" box="[484,505,529,553]" italics="true" pageId="16" pageNumber="17">in</emphasis>
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Lamas (ed.).
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<emphasis id="718F3B3F9D4ACA5DFD65FD92FB75FDBD" bold="true" box="[654,1174,529,554]" pageId="16" pageNumber="17">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
|
||
,
|
||
<emphasis id="718F3B3F9D4ACA5DFB4DFD92FB58FDBE" bold="true" box="[1190,1211,529,553]" italics="true" pageId="16" pageNumber="17">in</emphasis>
|
||
Heppner (ed.).
|
||
<emphasis id="718F3B3F9D4ACA5DFA8FFD92FE60FDDF" bold="true" pageId="16" pageNumber="17">
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFA8FFD92FEBEFDDF" authority="Neotrop. Lep." authorityName="Neotrop. Lep." class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
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5A
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</emphasis>
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||
.—O.
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||
<bibRefCitation id="276A9ADC9D4ACA5DFE29FDACFDAEFDDF" author="Mielke, O. H. H." box="[450,589,559,584]" pageId="16" pageNumber="17" pagination="775 - 1055" refId="ref43529" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 4. Hesperiinae 1. Adlerodea- Lychnuchus. Sociedade Brasileira de Zoologia, Curitiba. pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
|
||
.
|
||
<emphasis id="718F3B3F9D4ACA5DFDBDFDACFC97FDDF" bold="true" box="[598,884,559,584]" pageId="16" pageNumber="17">Cat. Amer. Hesperioidea 4</emphasis>
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, p. 833.—
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFC36FDACFB15FDDF" author="Garwood, K. & Lehman, R." box="[989,1270,559,584]" pageId="16" pageNumber="17" refId="ref40900" refString="Garwood, K. & Lehman, R. (2013) Butterflies of Central America. A photographic checklist of common species 3: Hesperiidae, The skippers. RiCale Publishing, Mc Allen, xii + 288 pp." type="book" year="2013">Garwood & Lehman, 2013</bibRefCitation>
|
||
.
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||
<emphasis id="718F3B3F9D4ACA5DFB14FDB3FEF4FDF0" bold="true" pageId="16" pageNumber="17">Butt. C. Amer. 3, Hesp</emphasis>
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., p. 228, fig. (v).—Carneiro; O. Mielke & Casagrande, 2015.
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||
<emphasis id="718F3B3F9D4ACA5DFC62FDCCFBF8FDF0" bold="true" box="[905,1051,590,615]" pageId="16" pageNumber="17">Zootaxa 3931</emphasis>
|
||
(2): 198.—Dolibaina; A. Warren; Carneiro & O. Mielke, 2015.
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||
<emphasis id="718F3B3F9D4ACA5DFE25FDEEFD77FD11" bold="true" box="[462,660,621,646]" pageId="16" pageNumber="17">Trop. Lep. Res. 25</emphasis>
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(2): 53, figs 3-4 (male d, v).
|
||
</paragraph>
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||
<paragraph id="4344E72D9D4ACA5DFF7CFD30FF18FC81" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF7CFD30FE98FD59" bold="true" box="[151,379,691,718]" pageId="16" pageNumber="17">Taxonomic history.</emphasis>
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||
<taxonomicName id="84FB9CAE9D4ACA5DFE69FD37FDE4FD5A" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[386,519,692,718]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
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||
<emphasis id="718F3B3F9D4ACA5DFE69FD37FDE4FD5A" box="[386,519,692,718]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
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||
</taxonomicName>
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||
was described by
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFD3CFD37FBFBFD59" author="Steinhauser, S. R. & Austin, G. T." box="[727,1048,692,718]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin (1993)</bibRefCitation>
|
||
based in
|
||
<specimenCount id="55FD2CA49D4ACA5DFB68FD37FAF8FD59" box="[1155,1307,692,718]" pageId="16" pageNumber="17" type="generic">12 specimens</specimenCount>
|
||
(males and females) from Turrialba,
|
||
<collectingRegion id="813F29CF9D4ACA5DFE5EFD5BFDF3FD65" box="[437,528,728,754]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BECA7BD9D4ACA5DFDF6FD5BFD7FFD65" box="[541,668,728,754]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
|
||
. The other authors mentioned this species in taxonomic studies and catalogs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFC9CFDD1FC5D" pageId="16" pageNumber="17" type="diagnosis">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFC9CFDD1FC5D" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFC9CFEA1FCAD" bold="true" box="[199,322,799,826]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFEA1FCA3FE33FCAE" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[330,464,800,826]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
|
||
<emphasis id="718F3B3F9D4ACA5DFEA1FCA3FE33FCAE" box="[330,464,800,826]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
|
||
</taxonomicName>
|
||
differs from the other species of the “
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFC6FFCA3FC36FCAE" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[900,981,800,825]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aepitus">
|
||
<emphasis id="718F3B3F9D4ACA5DFC6FFCA3FC36FCAE" box="[900,981,800,825]" italics="true" pageId="16" pageNumber="17">aepitus</emphasis>
|
||
</taxonomicName>
|
||
group” in the following combination of characters: harpe with ventral process narrow and cylindrical, shorter than tegumen+uncus, with setae on apical portion; cornutus with thin and straight spines, directed posteriorly; proximo-ventral folds of lamella antevaginalis similar to
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFEE3FC0FFE94FC32" baseAuthorityName="Geyer" baseAuthorityYear="1832" box="[264,375,908,934]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aepitus">
|
||
<emphasis id="718F3B3F9D4ACA5DFEE3FC0FFE94FC32" box="[264,375,908,934]" italics="true" pageId="16" pageNumber="17">A. aepitus</emphasis>
|
||
</taxonomicName>
|
||
, but less developed, distal projections of lamella postvaginalis separated by deeper cleft; distribution restricted to Central America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFC50FE5FFB85" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFC50FE5FFB85" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFC50FEDBFC79" bold="true" box="[199,312,979,1006]" pageId="16" pageNumber="17">Variation</emphasis>
|
||
. According to
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFE35FC57FCFEFC79" author="Steinhauser, S. R. & Austin, G. T." box="[478,797,980,1006]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin (1993)</bibRefCitation>
|
||
there is variation in the presence or absence of discal and apical marks on forewing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFB98FB4FFB7D" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFB98FAFCFB35" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFB98FEA5FBA1" bold="true" box="[199,326,1051,1078]" pageId="16" pageNumber="17">Comments</emphasis>
|
||
.
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFEBFFB9FFD6CFBA1" author="Steinhauser, S. R. & Austin, G. T." box="[340,655,1052,1078]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin (1993)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFCECFB9FFC6AFBA2" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[775,905,1052,1078]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
|
||
<emphasis id="718F3B3F9D4ACA5DFCECFB9FFC6AFBA2" box="[775,905,1052,1078]" italics="true" pageId="16" pageNumber="17">Artines rica</emphasis>
|
||
</taxonomicName>
|
||
and mentioned some specimens from
|
||
<collectingRegion id="813F29CF9D4ACA5DFADAFB9FFA7FFBA1" box="[1329,1436,1052,1078]" country="Brazil" name="Rondonia" pageId="16" pageNumber="17">Rondônia</collectingRegion>
|
||
and Distrito Federal,
|
||
<collectingCountry id="3BECA7BD9D4ACA5DFE94FBC3FE26FBCD" box="[383,453,1088,1114]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
|
||
. They did not designate any of them as
|
||
<typeStatus id="9C40598F9D4ACA5DFC9EFBC3FC02FBCD" box="[885,993,1088,1114]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
|
||
. The identification of the specimen from Distrito Federal is dubious because its genitalia, according to the authors, is damaged. The specimens from these localities studied by us belong to
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFE23FB0BFDF8FB35" authority="Medeiros & Dolibaina & Carneiro & Mielke & Casagrande, 2019" authorityName="Medeiros & Dolibaina & Carneiro & Mielke & Casagrande" authorityYear="2019" box="[456,539,1160,1186]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="liege" status="sp. nov.">
|
||
<emphasis id="718F3B3F9D4ACA5DFE23FB0BFDF8FB35" box="[456,539,1160,1186]" italics="true" pageId="16" pageNumber="17">A. liege</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="718F3B3F9D4ACA5DFDC9FB0BFD9AFB35" bold="true" box="[546,633,1160,1186]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="6ABC86449D4ACA5DFDC9FB0BFD9AFB35" box="[546,633,1160,1186]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Therefore,
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFD13FB0BFCA0FB36" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[760,835,1160,1186]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
|
||
<emphasis id="718F3B3F9D4ACA5DFD13FB0BFCA0FB36" box="[760,835,1160,1186]" italics="true" pageId="16" pageNumber="17">A. rica</emphasis>
|
||
</taxonomicName>
|
||
probably does not occur in South America.
|
||
</paragraph>
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFB28FB4FFB7D" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFB28FE0BFB51" bold="true" box="[199,488,1195,1222]" pageId="16" pageNumber="17">Geographic distribution</emphasis>
|
||
(
|
||
<figureCitation id="DBC0FBA89D4ACA5DFE1CFB2FFDAFFB51" box="[503,588,1196,1222]" captionStart="FIGURE 78" captionStartId="42.[151,250,910,935]" captionTargetBox="[151,1436,174,898]" captionTargetId="figure@42.[151,1436,174,898]" captionTargetPageId="42" captionText="FIGURE 78. Geographic distribution of the “aepitus group” species. The symbols with white inferior half represent records from literature." httpUri="https://zenodo.org/record/3241067/files/figure.png" pageId="16" pageNumber="17">Fig. 78</figureCitation>
|
||
). The species occurs in forested areas of Central America. COSTA RICA—
|
||
<emphasis id="718F3B3F9D4ACA5DFF7CFB53FF17FB7D" box="[151,244,1232,1258]" italics="true" pageId="16" pageNumber="17">
|
||
<collectingRegion id="813F29CF9D4ACA5DFF7CFB53FF17FB7D" box="[151,244,1232,1258]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
|
||
</emphasis>
|
||
: Turrialba.
|
||
<emphasis id="718F3B3F9D4ACA5DFE93FB53FE1DFB7D" box="[376,510,1232,1258]" italics="true" pageId="16" pageNumber="17">
|
||
<collectingRegion id="813F29CF9D4ACA5DFE93FB53FE1DFB7D" box="[376,510,1232,1258]" country="Costa Rica" name="Guanacaste" pageId="16" pageNumber="17">Guanacaste</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="813F29CF9D4ACA5DFDE6FB53FD6DFB7D" box="[525,654,1232,1258]" country="Costa Rica" name="Guanacaste" pageId="16" pageNumber="17">Guanacaste</collectingRegion>
|
||
Conservation Area (
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFC9FFB53FB43FB7D" author="Janzen, D. H. & Hallwachs, W." box="[884,1184,1232,1258]" pageId="16" pageNumber="17" refId="ref41792" refString="Janzen, D. H. & Hallwachs, W. (2009) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available from: http: // janzen. sas. upenn. edu / (accessed 15 November 2018)" type="url" year="2009">Janzen & Hallwachs 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFB77FC22FA99" box="[199,961,1267,1294]" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFB77FC22FA99" blockId="16.[151,1437,691,1618]" box="[199,961,1267,1294]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFB77FE33FA99" bold="true" box="[199,464,1267,1294]" pageId="16" pageNumber="17">Temporal distribution</emphasis>
|
||
. This species occurs in April, June and July.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFA9BFB6BFAA5" box="[199,1160,1304,1330]" pageId="16" pageNumber="17" type="biology_ecology">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFA9BFB6BFAA5" blockId="16.[151,1437,691,1618]" box="[199,1160,1304,1330]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFA9BFEA7FAA5" bold="true" box="[199,324,1304,1330]" pageId="16" pageNumber="17">Host plant</emphasis>
|
||
.
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFEB9FA9BFE16FAA5" authorityName="Trin" authorityYear="1836" box="[338,501,1304,1330]" class="Actinopterygii" family="Olyridae" genus="Olyra" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="caudata">
|
||
<emphasis id="718F3B3F9D4ACA5DFEB9FA9BFE16FAA5" box="[338,501,1304,1330]" italics="true" pageId="16" pageNumber="17">Olyra caudata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFDC0FA9BFCA1FAA5" box="[555,834,1304,1330]" class="Actinopterygii" family="Olyridae" genus="Olyra" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="latifolia">
|
||
<emphasis id="718F3B3F9D4ACA5DFDC0FA9BFD2EFAA5" box="[555,717,1304,1330]" italics="true" pageId="16" pageNumber="17">Olyra latifolia</emphasis>
|
||
(Poaceae)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFCBBFA9BFB9FFAA5" author="Janzen, D. H. & Hallwachs, W." box="[848,1148,1304,1330]" pageId="16" pageNumber="17" refId="ref41792" refString="Janzen, D. H. & Hallwachs, W. (2009) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available from: http: // janzen. sas. upenn. edu / (accessed 15 November 2018)" type="url" year="2009">Janzen & Hallwachs 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFABFFA76FAC1" box="[199,1429,1340,1366]" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFABFFA76FAC1" blockId="16.[151,1437,691,1618]" box="[199,1429,1340,1366]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFABFFEA4FAC1" bold="true" box="[199,327,1340,1366]" pageId="16" pageNumber="17">Etymology</emphasis>
|
||
. According to
|
||
<bibRefCitation id="276A9ADC9D4ACA5DFE00FABFFCCAFAC1" author="Steinhauser, S. R. & Austin, G. T." box="[491,809,1340,1366]" pageId="16" pageNumber="17" pagination="12 - 20" refId="ref45097" refString="Steinhauser, S. R. & Austin, G. T. (1993) New species of Hesperiidae from Costa Rica. Tropical Lepidopter a, 4 (Supplement 2), 12 - 20." type="journal article" year="1993">Steinhauser & Austin (1993)</bibRefCitation>
|
||
the term "
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFC76FABFFC29FAC2" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[925,970,1340,1365]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
|
||
<emphasis id="718F3B3F9D4ACA5DFC76FABFFC29FAC2" box="[925,970,1340,1365]" italics="true" pageId="16" pageNumber="17">rica</emphasis>
|
||
</taxonomicName>
|
||
" refers to the
|
||
<typeStatus id="9C40598F9D4ACA5DFB8CFABFFB74FAC1" box="[1127,1175,1340,1366]" pageId="16" pageNumber="17">type</typeStatus>
|
||
locality of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BE1B4A69D4ACA5DFF2CFAE3FB97F9C5" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFAE3FDD9FA71" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFAE3FE93FAED" bold="true" box="[199,368,1375,1402]" pageId="16" pageNumber="17">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="F393ED709D4ACA5DFE6AFAE3FB08FAED" ID-GBIF-Occurrence="2248478880" box="[385,1259,1376,1402]" collectionCode="MGCL, AME" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="9C40598F9D4ACA5DFE6AFAE3FE0AFAED" box="[385,489,1376,1402]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||
male deposited at the
|
||
<collectionCode id="25EA7FE89D4ACA5DFD1AFAE3FCA2FAED" box="[753,833,1376,1402]" pageId="16" pageNumber="17">MGCL</collectionCode>
|
||
(ex
|
||
<collectionCode id="25EA7FE89D4ACA5DFC9DFAE3FC55FAED" box="[886,950,1376,1402]" pageId="16" pageNumber="17">AME</collectionCode>
|
||
) with the following labels
|
||
</materialsCitation>
|
||
: /
|
||
<materialsCitation id="F393ED709D4ACA5DFAE3FAE3FDCEFA71" ID-GBIF-Occurrence="2248478804" collectingDate="1972-06" collectorName="H. L. King & Database & Allyn Museum Photo & No. & A. C. Allyn Ace. & Specimen" country="Costa Rica" location="Turriable" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
|
||
<typeStatus id="9C40598F9D4ACA5DFAE3FAE3FA7FFAED" box="[1288,1436,1376,1402]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="84FB9CAE9D4ACA5DFF7CFA07FD43FA09" authority=", S. R. Steinhauser & G. T. Austin" authorityName=", S. R. Steinhauser & G. T. Austin" authorityYear="1993" box="[151,672,1412,1438]" class="Insecta" family="Hesperiidae" genus="Artines" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rica">
|
||
<emphasis id="718F3B3F9D4ACA5DFF7CFA07FEC6FA0A" box="[151,293,1412,1438]" italics="true" pageId="16" pageNumber="17">Artines rica,</emphasis>
|
||
S. R. Steinhauser & G. T. Austin
|
||
</taxonomicName>
|
||
/
|
||
<collectingCountry id="3BECA7BD9D4ACA5DFD5CFA07FCBDFA09" box="[695,862,1412,1438]" name="Costa Rica" pageId="16" pageNumber="17">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="813F29CF9D4ACA5DFC81FA07FC13FA09" box="[874,1008,1412,1438]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">CARTAGO</collectingRegion>
|
||
,
|
||
<location id="4624B1F69D4ACA5DFC17FA07FB80FA09" LSID="urn:lsid:plazi:treatment:CB52563B9D4ACA5DFF7CFECEFB97F9C5:4624B1F69D4ACA5DFC17FA07FB80FA09" box="[1020,1123,1412,1438]" country="Costa Rica" name="Turriable" pageId="16" pageNumber="17" stateProvince="Cartago">Turriable</location>
|
||
[sic!] l.
|
||
<date id="3745C1ED9D4ACA5DFB51FA07FAF7FA09" box="[1210,1300,1412,1438]" pageId="16" pageNumber="17" value="1972-06">
|
||
<collectingDate id="270138059D4ACA5DFB51FA07FAF7FA09" box="[1210,1300,1412,1438]" pageId="16" pageNumber="17" value="1972-06">vi.1972</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EE0E82FB9D4ACA5DFACAFA07FA7FFA09" box="[1313,1436,1412,1438]" pageId="16" pageNumber="17">H. L. King</collectorName>
|
||
/ SRS
|
||
<collectorName id="EE0E82FB9D4ACA5DFF09FA2BFEAFFA55" box="[226,332,1448,1474]" pageId="16" pageNumber="17">Database</collectorName>
|
||
, Nº 568 /
|
||
<collectorName id="EE0E82FB9D4ACA5DFE2BFA2BFD56FA55" box="[448,693,1448,1474]" pageId="16" pageNumber="17">Allyn Museum Photo</collectorName>
|
||
/
|
||
<collectorName id="EE0E82FB9D4ACA5DFD24FA2BFD1BFA55" box="[719,760,1448,1474]" pageId="16" pageNumber="17">No.</collectorName>
|
||
890626-19.20 / 890719-21, 22 /
|
||
<collectorName id="EE0E82FB9D4ACA5DFB98FA2BFAD7FA55" box="[1139,1332,1448,1474]" pageId="16" pageNumber="17">A. C. Allyn Ace.</collectorName>
|
||
1972-5 / MGCLFLMNH
|
||
<collectorName id="EE0E82FB9D4ACA5DFEBAFA4FFE5CFA71" box="[337,447,1484,1510]" pageId="16" pageNumber="17">Specimen</collectorName>
|
||
nº 47907
|
||
</materialsCitation>
|
||
/.
|
||
</paragraph>
|
||
<paragraph id="4344E72D9D4ACA5DFF2CFA6CFB97F9C5" blockId="16.[151,1437,691,1618]" pageId="16" pageNumber="17">
|
||
<emphasis id="718F3B3F9D4ACA5DFF2CFA6CFE4EF99D" bold="true" box="[199,429,1519,1546]" pageId="16" pageNumber="17">Examined material</emphasis>
|
||
. Illustration of the
|
||
<typeStatus id="9C40598F9D4ACA5DFD6FFA73FD05F99D" box="[644,742,1520,1546]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
and the following specimens: COSTA RICA—
|
||
<emphasis id="718F3B3F9D4ACA5DFB17FA73FABAF99D" box="[1276,1369,1520,1546]" italics="true" pageId="16" pageNumber="17">
|
||
<collectingRegion id="813F29CF9D4ACA5DFB17FA73FABAF99D" box="[1276,1369,1520,1546]" country="Costa Rica" name="Cartago" pageId="16" pageNumber="17">Cartago</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="718F3B3F9D4ACA5DFA83FA73FF3CF9B9" bold="true" pageId="16" pageNumber="17">Turrialba</emphasis>
|
||
,
|
||
<specimenCount id="55FD2CA49D4ACA5DFF07F997FEACF9B9" box="[236,335,1556,1582]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="3745C1ED9D4ACA5DFEB1F990FE3BF9B9" box="[346,472,1555,1582]" pageId="16" pageNumber="17" value="1971-07-05">5.VII.1971</date>
|
||
,
|
||
<materialsCitation id="F393ED709D4ACA5DFE09F997FDFCF9C6" ID-GBIF-Occurrence="2248478873" collectingDate="1973-04-15" collectingDateMax="1973-04-30" collectingDateMin="1973-04-15" collectionCode="DZUP" collectorName="Becker" elevation="600" pageId="16" pageNumber="17" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
|
||
<specimenCount id="55FD2CA49D4ACA5DFE09F997FDD8F9B9" box="[482,571,1556,1582]" pageId="16" pageNumber="17" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="55FD2CA49D4ACA5DFDAEF997FD4AF9B9" box="[581,681,1556,1582]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="3745C1ED9D4ACA5DFD58F990FCBFF9B9" box="[691,860,1555,1582]" pageId="16" pageNumber="17" value="1973-04-15" valueMax="1973-04-30" valueMin="1973-04-15">
|
||
<collectingDate id="270138059D4ACA5DFD58F990FCBFF9B9" box="[691,860,1555,1582]" pageId="16" pageNumber="17" value="1973-04-15" valueMax="1973-04-30" valueMin="1973-04-15">15–30.IV.1973</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EE0E82FB9D4ACA5DFC8CF997FC55F9B9" box="[871,950,1556,1582]" pageId="16" pageNumber="17">Becker</collectorName>
|
||
<emphasis id="718F3B3F9D4ACA5DFC57F997FC06F9B9" box="[956,997,1556,1582]" italics="true" pageId="16" pageNumber="17">leg.</emphasis>
|
||
, DZ 31.561, DZ 4.061, DZ 41.890, DZ 30.956, DZ 41.840 (
|
||
<collectionCode id="25EA7FE89D4ACA5DFE91F9BBFE2AF9C5" box="[378,457,1592,1618]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="16" pageNumber="17" type="University or college">DZUP</collectionCode>
|
||
),
|
||
<quantity id="84034AC89D4ACA5DFE37F9B4FDFCF9C6" box="[476,543,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="m" value="600.0">
|
||
<elevation id="C8D6001E9D4ACA5DFE37F9B4FDFCF9C6" box="[476,543,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="m" value="600.0">600m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="F393ED709D4ACA5DFDC1F9BBFD25F9C5" ID-GBIF-Occurrence="2248478863" box="[554,710,1592,1618]" collectionCode="VI" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="55FD2CA49D4ACA5DFDC1F9BBFD9BF9C5" box="[554,632,1592,1618]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
|
||
,
|
||
<date id="3745C1ED9D4ACA5DFD68F9BBFD25F9C5" box="[643,710,1592,1618]" pageId="16" pageNumber="17" value="1973-06-29">
|
||
29.
|
||
<collectionCode id="25EA7FE89D4ACA5DFD4DF9BBFD25F9C5" box="[678,710,1592,1618]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="17">VI</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
.1973,
|
||
<collectorName id="EE0E82FB9D4ACA5DFCFAF9BBFC83F9C5" box="[785,864,1592,1618]" pageId="16" pageNumber="17">Becker</collectorName>
|
||
<emphasis id="718F3B3F9D4ACA5DFC8CF9BBFC6AF9C5" box="[871,905,1592,1618]" italics="true" pageId="16" pageNumber="17">leg</emphasis>
|
||
., DZ 8.870 (
|
||
<collectionCode id="25EA7FE89D4ACA5DFBF2F9BBFB8AF9C5" box="[1049,1129,1592,1618]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="16" pageNumber="17" type="University or college">DZUP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |