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<document id="092770EA44880BB610A5D19195D775B9" ID-CLB-Dataset="23869" ID-DOI="10.11646/zootaxa.4742.3.1" ID-GBIF-Dataset="84ef451f-d1a1-48f6-871e-f688204fd959" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677869" ID-ZooBank="44ECBE3C-DEA4-4A6B-87D8-D7065591577C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272866112" checkinUser="plazi" docAuthor="Nunes, Benedito Mendes, Campos, Lourival Dias, Mendonca, Maria Thayane Da Silva, Cunha, Eduardo Victor De Paiva &amp; Fernandes, Jose Antonio Marin" docDate="2020" docId="720587AC3174D91344B26B2DFC0B1B91" docLanguage="en" docName="zootaxa.4742.3.1.pdf" docOrigin="Zootaxa 4742 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypoxys dolosus Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes 2020, comb. nov." docType="treatment" docVersion="12" lastPageNumber="413" masterDocId="8E3CFFD4317DD91F44256D6EFF9C1A3E" masterDocTitle="Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)" masterLastPageNumber="441" masterPageNumber="401" pageNumber="410" updateTime="1732689660721" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="07CB11B5271163E9639E5F504E3FA702">Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)</mods:title>
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<mods:namePart id="F16C93229553DEA95BA6362938AE47D0">Nunes, Benedito Mendes</mods:namePart>
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<mods:namePart id="7F1DD207400281842E3747D5BCE48F16">Campos, Lourival Dias</mods:namePart>
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<mods:namePart id="6D5E9F9B110447F2397171BAC67C157F">Mendonca, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="EB2A22B047B8F9FA386AABFD7ECC5D05">Cunha, Eduardo Victor De Paiva</mods:namePart>
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<mods:namePart id="DCF3F829B584EEFB7BC77F148807BDCD">Fernandes, Jose Antonio Marin</mods:namePart>
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<mods:title id="249C746C075B59667757AA56156B6C10">Zootaxa</mods:title>
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<mods:date id="59B0FC44BBB79EAB1E97DAB00F3511A3">2020</mods:date>
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<treatment id="720587AC3174D91344B26B2DFC0B1B91" ID-DOI="http://doi.org/10.5281/zenodo.3684976" ID-GBIF-Taxon="162244893" ID-Zenodo-Dep="3684976" LSID="urn:lsid:plazi:treatment:720587AC3174D91344B26B2DFC0B1B91" httpUri="http://treatment.plazi.org/id/720587AC3174D91344B26B2DFC0B1B91" lastPageId="12" lastPageNumber="413" pageId="9" pageNumber="410" scope_family="Pentatomidae">
<subSubSection id="B2B665313174D91644B26B2DFD261C63" box="[151,698,1602,1629]" pageId="9" pageNumber="410" type="nomenclature">
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<heading id="A15B81D63174D91644B26B2DFD261C63" bold="true" box="[151,698,1602,1629]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="C8D8EAA83174D91644B26B2DFD261C63" bold="true" box="[151,698,1602,1629]" pageId="9" pageNumber="410">
<taxonomicName id="3DAC4D393174D91644B26B2DFDB01C62" authority="(Breddin, 1907)" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[151,556,1602,1629]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="dolosus" status="comb. nov.">
<emphasis id="C8D8EAA83174D91644B26B2DFEFE1C63" bold="true" box="[151,354,1602,1629]" italics="true" pageId="9" pageNumber="410">Hypoxys dolosus</emphasis>
(
<bibRefCitation id="9E3D4B4B3174D91645546B2CFDB81C62" author="Breddin, G." box="[369,548,1602,1628]" pageId="9" pageNumber="410" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="D3EB57D33174D91646116B2DFD261C63" box="[564,698,1603,1629]" pageId="9" pageNumber="410" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2B665313174D91644B26B08FE5E1CBE" box="[151,450,1638,1664]" pageId="9" pageNumber="410" type="description">
<paragraph id="FA1336BA3174D91644B26B08FE5E1CBE" blockId="9.[151,698,1602,1664]" box="[151,450,1638,1664]" pageId="9" pageNumber="410">
(
<figureCitation id="62972A3F3174D91644BA6B08FF711CBE" box="[159,237,1638,1664]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="9" pageNumber="410">Figs. 4</figureCitation>
A–E; 18 G–H; 21)
</paragraph>
</subSubSection>
<subSubSection id="B2B665313174D91644B26BC5FB3D1CDC" pageId="9" pageNumber="410" type="reference_group">
<paragraph id="FA1336BA3174D91644B26BC5FB8A1CFD" blockId="9.[151,1185,1706,1792]" box="[151,1046,1706,1731]" pageId="9" pageNumber="410">
<treatmentCitationGroup id="DABC11943174D91644B26BC5FB8A1CFD" box="[151,1046,1706,1731]" pageId="9" pageNumber="410">
<treatmentCitation id="7B0D10AB3174D91644B26BC5FE6B1CFD" author="Breddin, G." box="[151,503,1706,1731]" page="336" pageId="9" pageNumber="410" year="1907">
<taxonomicName id="3DAC4D393174D91644B26BC5FE6B1CFD" authority="Breddin, 1907: 336" authorityName="Breddin" authorityPageNumber="336" authorityYear="1907" box="[151,503,1706,1731]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="C8D8EAA83174D91644B26BC5FEB51CFD" box="[151,297,1707,1731]" italics="true" pageId="9" pageNumber="410">Edessa dolosa</emphasis>
<bibRefCitation id="9E3D4B4B3174D916450A6BC4FE6B1CFD" author="Breddin, G." box="[303,503,1706,1731]" pageId="9" pageNumber="410" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907: 336</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="7B0D10AB3174D91646246BC4FD501CFD" author="Kirkaldy, G. W." box="[513,716,1706,1731]" page="368" pageId="9" pageNumber="410" year="1909">
<bibRefCitation id="9E3D4B4B3174D91646246BC4FD501CFD" author="Kirkaldy, G. W." box="[513,716,1706,1731]" pageId="9" pageNumber="410" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 368</bibRefCitation>
</treatmentCitation>
(catalog); Froeschner, 1981: 66.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA1336BA3174D91644B26BA4FB3D1CDC" blockId="9.[151,1185,1706,1792]" box="[151,1185,1737,1762]" pageId="9" pageNumber="410">
<materialsCitation id="4AC43CE73174D91644B26BA4FB3D1CDC" ID-GBIF-Occurrence="2570056089" box="[151,1185,1737,1762]" collectionCode="SDEI" country="Ecuador" location="Balzapamba" municipality="Palmas" pageId="9" pageNumber="410" specimenCount="1" typeStatus="lectotype">
<typeStatus id="251788183174D91644B26BA4FF601CDC" box="[151,252,1738,1762]" pageId="9" pageNumber="410" type="lectotype">Lectotype</typeStatus>
male.
<collectingCountry id="82BB762A3174D91645646BA7FE0B1CDC" box="[321,407,1737,1762]" name="Ecuador" pageId="9" pageNumber="410">Ecuador</collectingCountry>
,
<location id="FF7360613174D91645846BA7FDBD1CDC" LSID="urn:lsid:plazi:treatment:720587AC3174D91344B26B2DFC0B1B91:FF7360613174D91645846BA7FDBD1CDC" box="[417,545,1737,1762]" country="Ecuador" municipality="Palmas" name="Balzapamba" pageId="9" pageNumber="410">Balzapamba</location>
,
<collectingMunicipality id="1A77ACC03174D916460E6BA7FDE81CDC" box="[555,628,1737,1762]" pageId="9" pageNumber="410">Palmas</collectingMunicipality>
(
<collectionCode id="9CBDAE7F3174D91646A76BA7FD5C1CDF" box="[642,704,1737,1761]" pageId="9" pageNumber="410">SDEI</collectionCode>
). Designated by
<bibRefCitation id="9E3D4B4B3174D916474D6BA7FBB51CDC" author="Gaedike, H." box="[872,1065,1737,1762]" pageId="9" pageNumber="410" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971:85)</bibRefCitation>
. Examined.
</materialsCitation>
</paragraph>
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<subSubSection id="B2B665313174D91644B26B89FD471D3E" box="[151,731,1767,1792]" pageId="9" pageNumber="410" type="description">
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<typeStatus id="251788183174D91644B26B89FEBD1D3E" box="[151,289,1767,1792]" pageId="9" pageNumber="410" type="paralectotype">Paralectotype</typeStatus>
female. Same data as
<typeStatus id="251788183174D91646216B89FDF81D3E" box="[516,612,1767,1792]" pageId="9" pageNumber="410" type="lectotype">lectotype</typeStatus>
. Examined.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313174D91644B26A46FC3E1DC9" pageId="9" pageNumber="410" type="materials_examined">
<paragraph id="FA1336BA3174D91644B26A46FC3E1DC9" blockId="9.[151,1437,1832,2039]" pageId="9" pageNumber="410">
<emphasis id="C8D8EAA83174D91644B26A46FE1C1D7D" bold="true" box="[151,384,1832,1859]" pageId="9" pageNumber="410">Material examined.</emphasis>
<materialsCitation id="4AC43CE73174D91645A26A46FBCC1DB5" ID-GBIF-Occurrence="2570056050" collectingDate="1983-07-01" collectingDateMax="1983-07-10" collectingDateMin="1983-07-01" collectionCode="CMNH" collectorName="Carnegie &amp; Rawlins &amp; R. Davidson &amp; N. Jacobson &amp; M. Smyers &amp; S. Thompson &amp; S. Weller. Rawlins Collection Donated &amp; Fernandes" country="Ecuador" elevation="1250" latitude="0.93333334" location="Chical" longLatPrecision="1308" longitude="-78.183334" pageId="9" pageNumber="410" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
<emphasis id="C8D8EAA83174D91645A26A46FDE11D7D" bold="true" box="[391,637,1832,1859]" pageId="9" pageNumber="410">
<collectingCountry id="82BB762A3174D91645A26A46FD8B1D7D" box="[391,535,1832,1859]" name="Ecuador" pageId="9" pageNumber="410">ECUADOR</collectingCountry>
.
<collectingRegion id="3868F8583174D91646076A46FDE41D7D" box="[546,632,1832,1859]" country="Ecuador" name="Carchi" pageId="9" pageNumber="410">Carchi</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333174D91646A66A47FD041D7D" box="[643,664,1833,1859]" pageId="9" pageNumber="410" type="female">♀</specimenCount>
<location id="FF7360613174D91646BA6A47FD771D7D" LSID="urn:lsid:plazi:treatment:720587AC3174D91344B26B2DFC0B1B91:FF7360613174D91646BA6A47FD771D7D" box="[671,747,1833,1859]" country="Ecuador" latitude="0.93333334" longLatPrecision="1308" longitude="-78.183334" name="Chical" pageId="9" pageNumber="410" stateProvince="Carchi">Chical</location>
,
<quantity id="3D549B5F3174D91646D36A46FCD31D7D" box="[758,847,1832,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="410" unit="m" value="1250.0">
<elevation id="7181D1893174D91646D36A46FCD31D7D" box="[758,847,1832,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="410" unit="m" value="1250.0">1250 m</elevation>
</quantity>
.,
<geoCoordinate id="9F98507D3174D916477B6A46FC2E1D7D" box="[862,946,1832,1859]" degrees="0" direction="north" minutes="56" orientation="latitude" pageId="9" pageNumber="410" precision="925" value="0.93333334">0º56’N</geoCoordinate>
.
<geoCoordinate id="9F98507D3174D91647996A47FBBC1D7D" box="[956,1056,1833,1859]" degrees="78" direction="west" minutes="11" orientation="longitude" pageId="9" pageNumber="410" precision="925" value="-78.183334">78º11’W</geoCoordinate>
,
<date id="8E12107A3174D916400E6A47FB451D7D" box="[1067,1241,1833,1859]" pageId="9" pageNumber="410" value="1983-07-01" valueMax="1983-07-10" valueMin="1983-07-01">
<collectingDate id="9E56E9923174D916400E6A47FB451D7D" box="[1067,1241,1833,1859]" pageId="9" pageNumber="410" value="1983-07-01" valueMax="1983-07-10" valueMin="1983-07-01">1–10 July 1983</collectingDate>
</date>
, U. T.—
<collectorName id="5759536C3174D91641136A47FA001D7D" box="[1334,1436,1833,1859]" pageId="9" pageNumber="410">Carnegie</collectorName>
expdt. J.
<collectorName id="5759536C3174D91645276A23FEFD1D59" box="[258,353,1869,1895]" pageId="9" pageNumber="410">Rawlins</collectorName>
,
<collectorName id="5759536C3174D916454B6A23FD9C1D59" box="[366,512,1869,1895]" pageId="9" pageNumber="410">R. Davidson</collectorName>
,
<collectorName id="5759536C3174D916462B6A23FD011D59" box="[526,669,1869,1895]" pageId="9" pageNumber="410">N. Jacobson</collectorName>
,
<collectorName id="5759536C3174D916468F6A23FCB71D59" box="[682,811,1869,1895]" pageId="9" pageNumber="410">M. Smyers</collectorName>
,
<collectorName id="5759536C3174D916471D6A23FC4F1D59" box="[824,979,1869,1895]" pageId="9" pageNumber="410">S. Thompson</collectorName>
,
<collectorName id="5759536C3174D91647C46A23FA001D59" box="[993,1436,1869,1895]" pageId="9" pageNumber="410">S. Weller. Rawlins Collection Donated</collectorName>
1988,
<taxonomicName id="3DAC4D393174D91644F86A1FFDEA1DB5" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" box="[221,630,1905,1931]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="C8D8EAA83174D91644F86A1FFEE21DB5" box="[221,382,1905,1931]" italics="true" pageId="9" pageNumber="410">Edessa dolosa</emphasis>
<bibRefCitation id="9E3D4B4B3174D91645A16A1FFDB81DB5" author="Breddin, G." box="[388,548,1905,1931]" pageId="9" pageNumber="410" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w.
<typeStatus id="251788183174D91646B96A1FFD7E1DB5" box="[668,738,1905,1931]" pageId="9" pageNumber="410">TYPE</typeStatus>
<collectorName id="5759536C3174D91646CD6A1FFCC21DB5" box="[744,862,1905,1931]" pageId="9" pageNumber="410">Fernandes</collectorName>
, JAM 1999 (
<collectionCode id="9CBDAE7F3174D91647CB6A1FFBD41DB5" box="[1006,1096,1905,1931]" country="USA" name="The Cleveland Museum of Natural History" pageId="9" pageNumber="410" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AC43CE73174D916407E6A1EFD3A1DED" ID-GBIF-Occurrence="2570056041" collectingDate="1986-05-11" collectingDateMax="1986-05-17" collectingDateMin="1986-05-11" collectionCode="JEE" collectorName="E. Sto. Domingo de Los Colorados &amp; J. E. Eger Coll. &amp; Fernandes" country="Ecuador" elevation="762" location="Ca." municipality="Tinalandia" pageId="9" pageNumber="410" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha">
<emphasis id="C8D8EAA83174D916407E6A1EFB461DB5" bold="true" box="[1115,1242,1904,1931]" pageId="9" pageNumber="410">
<collectingRegion id="3868F8583174D916407E6A1EFB4A1DB5" box="[1115,1238,1904,1931]" country="Ecuador" name="Pichincha" pageId="9" pageNumber="410">Pichincha</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333174D91640C46A1FFB6A1DB5" box="[1249,1270,1905,1931]" pageId="9" pageNumber="410" type="male">♂</specimenCount>
<collectingMunicipality id="1A77ACC03174D91640D96A1FFAEA1DB5" box="[1276,1398,1905,1931]" pageId="9" pageNumber="410">Tinalandia</collectingMunicipality>
,
<quantity id="3D549B5F3174D91641A56A1FFF221D91" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="410" unit="km" value="12.0">12 km</quantity>
.
<collectorName id="5759536C3174D91644ED6AFBFDD31D91" box="[200,591,1941,1967]" pageId="9" pageNumber="410">E. Sto. Domingo de Los Colorados</collectorName>
,
<location id="FF7360613174D916467C6AFBFDE31D91" LSID="urn:lsid:plazi:treatment:720587AC3174D91344B26B2DFC0B1B91:FF7360613174D916467C6AFBFDE31D91" box="[601,639,1941,1967]" country="Ecuador" municipality="Tinalandia" name="Ca." pageId="9" pageNumber="410" stateProvince="Pichincha">Ca.</location>
<quantity id="3D549B5F3174D91646A06AFAFD411D91" box="[645,733,1940,1967]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="410" unit="ft" value="2500.0">
<elevation id="7181D1893174D91646A06AFAFD411D91" box="[645,733,1940,1967]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="410" unit="ft" value="2500.0">2500 ft.</elevation>
</quantity>
,
<date id="8E12107A3174D91646CD6AFBFC041D91" box="[744,920,1941,1967]" pageId="9" pageNumber="410" value="1986-05-11" valueMax="1986-05-17" valueMin="1986-05-11">
<collectingDate id="9E56E9923174D91646CD6AFBFC041D91" box="[744,920,1941,1967]" pageId="9" pageNumber="410" value="1986-05-11" valueMax="1986-05-17" valueMin="1986-05-11">11–17–V–1986</collectingDate>
</date>
,
<collectorName id="5759536C3174D91647876AFBFBD11D91" box="[930,1101,1941,1967]" pageId="9" pageNumber="410">J. E. Eger Coll.</collectorName>
,
<taxonomicName id="3DAC4D393174D91640726AFBFF7F1DED" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="C8D8EAA83174D91640726AFBFB6B1D91" box="[1111,1271,1941,1967]" italics="true" pageId="9" pageNumber="410">Edessa dolosa</emphasis>
<bibRefCitation id="9E3D4B4B3174D91640D86AFBFA001D91" author="Breddin, G." box="[1277,1436,1941,1967]" pageId="9" pageNumber="410" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w.
<typeStatus id="251788183174D916452B6AD7FEC81DED" box="[270,340,1977,2003]" pageId="9" pageNumber="410">TYPE</typeStatus>
<collectorName id="5759536C3174D91645786AD7FE4F1DED" box="[349,467,1977,2003]" pageId="9" pageNumber="410">Fernandes</collectorName>
, JAM 1999 (
<collectionCode id="9CBDAE7F3174D916464E6AD7FD021DED" box="[619,670,1977,2003]" pageId="9" pageNumber="410">JEE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AC43CE73174D91646916AD6FADB1DED" ID-GBIF-Occurrence="2570056023" box="[692,1351,1976,2003]" collectionCode="MZUSP" collectorName="Balzapamba. R. Haensch S." country="Ecuador" location="Bolivar" pageId="9" pageNumber="410" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar">
<emphasis id="C8D8EAA83174D91646916AD6FC8A1DED" bold="true" box="[692,790,1976,2003]" pageId="9" pageNumber="410">
<collectingRegion id="3868F8583174D91646916AD6FC8D1DED" box="[692,785,1976,2003]" country="Ecuador" name="Bolivar" pageId="9" pageNumber="410">Bolívar</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333174D916473A6AD7FCA81DED" box="[799,820,1977,2003]" pageId="9" pageNumber="410" type="male">♂</specimenCount>
<collectorName id="5759536C3174D91647186AD7FBE71DED" box="[829,1147,1977,2003]" pageId="9" pageNumber="410">Balzapamba. R. Haensch S.</collectorName>
72137 (
<collectionCode id="9CBDAE7F3174D91640FE6AD7FAA31DED" box="[1243,1343,1977,2003]" country="Brazil" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="410">MZUSP</collectionCode>
)
</materialsCitation>
; wrong locality data:
<materialsCitation id="4AC43CE73174D91645156AB3FC031DC9" ID-GBIF-Occurrence="2570056055" box="[304,927,2013,2039]" collectionCode="CMNH" collectorName="Coll. Nickerl Mus. Pragense" country="Ecuador" location="Zambesi" pageId="9" pageNumber="410" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar">
<specimenCount id="ECAAFD333174D91645156AB3FED91DC9" box="[304,325,2013,2039]" pageId="9" pageNumber="410" type="male">♂</specimenCount>
<location id="FF7360613174D91645696AB3FE301DC9" LSID="urn:lsid:plazi:treatment:720587AC3174D91344B26B2DFC0B1B91:FF7360613174D91645696AB3FE301DC9" box="[332,428,2013,2039]" country="Ecuador" name="Zambesi" pageId="9" pageNumber="410" stateProvince="Bolivar">Zambesi</location>
V/II,
<collectorName id="5759536C3174D91645CA6AB3FCB11DC9" box="[495,813,2013,2039]" pageId="9" pageNumber="410">Coll. Nickerl Mus. Pragense</collectorName>
(
<collectionCode id="9CBDAE7F3174D91647196AB3FC0A1DC9" box="[828,918,2013,2039]" country="USA" name="The Cleveland Museum of Natural History" pageId="9" pageNumber="410" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="AED366323177D91544B26AD1FBF51DC2" ID-DOI="http://doi.org/10.5281/zenodo.3677875" ID-Zenodo-Dep="3677875" httpUri="https://zenodo.org/record/3677875/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,250,1983,2008]" targetBox="[210,1383,200,1948]" targetPageId="10">
<paragraph id="FA1336BA3177D91544B26AD1FBF51DC2" blockId="10.[151,1436,1983,2044]" pageId="10" pageNumber="411">
<emphasis id="C8D8EAA83177D91544B26AD1FE881DE6" bold="true" box="[151,276,1983,2008]" pageId="10" pageNumber="411">FIGURE 3.</emphasis>
<taxonomicName id="3DAC4D393177D91545396AAEFE3D1DE9" authorityName="Breddin" authorityYear="1907" box="[284,417,1983,2007]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="claricolor">
<emphasis id="C8D8EAA83177D91545396AAEFE3D1DE9" box="[284,417,1983,2007]" italics="true" pageId="10" pageNumber="411">H. claricolor</emphasis>
</taxonomicName>
. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data (Scale = 1mm).
</paragraph>
</caption>
<caption id="AED366323176D91444B26B26FDEF1CBB" ID-DOI="http://doi.org/10.5281/zenodo.3677877" ID-Zenodo-Dep="3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="11" pageNumber="412" startId="11.[151,250,1608,1633]" targetBox="[162,1424,203,1547]" targetPageId="11">
<paragraph id="FA1336BA3176D91444B26B26FDEF1CBB" blockId="11.[151,1436,1608,1669]" pageId="11" pageNumber="412">
<emphasis id="C8D8EAA83176D91444B26B26FE8A1C5F" bold="true" box="[151,278,1608,1633]" pageId="11" pageNumber="412">FIGURE 4.</emphasis>
<taxonomicName id="3DAC4D393176D914453A6B27FE121C5F" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[287,398,1609,1633]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="C8D8EAA83176D914453A6B27FE121C5F" box="[287,398,1609,1633]" italics="true" pageId="11" pageNumber="412">H. dolosus</emphasis>
</taxonomicName>
. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm).
</paragraph>
</caption>
<subSubSection id="B2B665313176D91444E26BDDFAD61D2B" pageId="11" pageNumber="412" type="description">
<paragraph id="FA1336BA3176D91444E26BDDFAD61D2B" blockId="11.[151,1436,1714,2030]" pageId="11" pageNumber="412">
<emphasis id="C8D8EAA83176D91444E26BDDFEE21CF3" bold="true" box="[199,382,1715,1741]" pageId="11" pageNumber="412">Measurements.</emphasis>
total length: 15.9–17.4; head length: 1.9–2.1; head width: 3.0–3.4; pronotal width: 10.0–11.0; abdominal width: 8.8–9.7; length of antennal segments. I: 0.8–1.0; II: 1.7–1.9; III: 1.2–1.7; IV: 3.4–4.6; V: 4.4; interocular distance: 1.4–1.6; scutellum length: 7.2–8.3; scutellum width: 5.7–6.1; pronotal length: 3.3–3.7.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313176D91344E26A70FBFD1AC5" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412" type="diagnosis">
<paragraph id="FA1336BA3176D91344E26A70FBFD1AC5" blockId="11.[151,1436,1714,2030]" lastBlockId="12.[151,1437,153,431]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
<emphasis id="C8D8EAA83176D91444E26A70FEDE1D07" bold="true" box="[199,322,1822,1849]" pageId="11" pageNumber="412">Diagnosis.</emphasis>
Anterolateral margin of pronotum without solid black stripe. Cicatrices with black punctures delimiting anterior sulci (
<figureCitation id="62972A3F3176D91445536A2DFE431D63" box="[374,479,1859,1885]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="11" pageNumber="412">Fig. 18G</figureCitation>
). Anterolateral margin with few punctures black. Male (
<figureCitation id="62972A3F3176D914407C6A2DFB001D63" box="[1113,1180,1859,1885]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="11" pageNumber="412">Fig. 4</figureCitation>
A–C). Pygophore with dorsal rim widely excavated. Superior process of genital cup small, tongue-like with a dorsal transverse crest; near dorsal rim (
<figureCitation id="62972A3F3176D914453C6AE5FEED1D9B" box="[281,369,1931,1957]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="11" pageNumber="412">Fig. 4B</figureCitation>
). Paramere with a single lobe curved anteriorly, tapering and elongated, margin rounded; shank flat with a developed carina on inner side continuing in a low carina on anterior lobe (
<figureCitation id="62972A3F3176D91440476AC1FB271DF7" box="[1122,1211,1967,1993]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="11" pageNumber="412">Fig. 4B</figureCitation>
). Proctiger with laterodorsal tufts of setae separated by wide tumid ridge; posterior face ogival (
<figureCitation id="62972A3F3176D91447D06ABDFBD11DD3" box="[1013,1101,2003,2029]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="11" pageNumber="412">Fig. 4B</figureCitation>
). Ventral rim with expansions rounded, flat, level with posterolateral angles in ventral view (
<figureCitation id="62972A3F3171D913476A6DF7FC341A8D" box="[847,936,153,179]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="12" pageNumber="413">Fig. 4C</figureCitation>
); base of each expansion with a conspicuous transverse carina (
<figureCitation id="62972A3F3171D91345416DD3FE231AE9" box="[356,447,189,215]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="12" pageNumber="413">Fig. 4A</figureCitation>
). Female (
<figureCitation id="62972A3F3171D91346106DD3FD0C1AE9" box="[565,656,189,215]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="12" pageNumber="413">Fig. 4D</figureCitation>
). Gonocoxites 8 with posterior margin almost straight, slanting, inner angle acute; sutural border divergent from the base, gonapophyses 8 partially exposed.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313171D91344E26C6AFE6B1BB5" pageId="12" pageNumber="413" type="discussion">
<paragraph id="FA1336BA3171D91344E26C6AFE6B1BB5" blockId="12.[151,1437,153,431]" pageId="12" pageNumber="413">
<emphasis id="C8D8EAA83171D91344E26C6AFED31B21" bold="true" box="[199,335,260,287]" pageId="12" pageNumber="413">Comments.</emphasis>
<bibRefCitation id="9E3D4B4B3171D91345706C6BFD9C1B21" author="Breddin, G." box="[341,512,261,287]" pageId="12" pageNumber="413" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin (1907)</bibRefCitation>
considered
<taxonomicName id="3DAC4D393171D91346A06C6BFCB91B21" authorityName="Breddin" authorityYear="1907" box="[645,805,261,287]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="C8D8EAA83171D91346A06C6BFCB91B21" box="[645,805,261,287]" italics="true" pageId="12" pageNumber="413">Edessa dolosa</emphasis>
</taxonomicName>
very similar to
<taxonomicName id="3DAC4D393171D91347F46C68FBF11B21" authorityName=", Amyot &amp; Serville" authorityYear="1843" box="[977,1133,261,287]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA83171D91347F46C68FC7E1B21" box="[977,994,262,287]" italics="true" pageId="12" pageNumber="413">E</emphasis>
.
<emphasis id="C8D8EAA83171D91347CA6C6BFBF11B21" box="[1007,1133,261,287]" italics="true" pageId="12" pageNumber="413">quadridens</emphasis>
</taxonomicName>
. They can be differentiated by the shape of the paramere and superior process of the genital cup (
<figureCitation id="62972A3F3171D91347B26C47FC771B7D" box="[919,1003,297,323]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="12" pageNumber="413">Fig. 4B</figureCitation>
and
<figureCitation id="62972A3F3171D91340056C47FBDE1B7D" box="[1056,1090,297,323]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="12" pageNumber="413">1B</figureCitation>
); gonocoxites 8 more flat in
<taxonomicName id="3DAC4D393171D91341A46C44FF711B59" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="C8D8EAA83171D91341A46C44FF711B59" italics="true" pageId="12" pageNumber="413">H. dolosus</emphasis>
</taxonomicName>
(convex in
<emphasis id="C8D8EAA83171D91345556C23FD891B59" box="[368,533,332,359]" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="3DAC4D393171D91345556C23FD961B59" box="[368,522,332,359]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA83171D91345556C23FD961B59" bold="true" box="[368,522,332,359]" italics="true" pageId="12" pageNumber="413">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
), posterior margin of the gonocoxites 8 more straight in
<taxonomicName id="3DAC4D393171D91340AE6C20FA9F1B59" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[1163,1283,333,359]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis id="C8D8EAA83171D91340AE6C20FA9F1B59" box="[1163,1283,333,359]" italics="true" pageId="12" pageNumber="413">H. dolosus</emphasis>
</taxonomicName>
(arched in
<emphasis id="C8D8EAA83171D913415A6C23FE841BB5" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="3DAC4D393171D913415A6C23FE911BB5" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA83171D913415A6C23FE911BB5" bold="true" italics="true" pageId="12" pageNumber="413">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
), (
<figureCitation id="62972A3F3171D913451D6C1FFE121BB5" box="[312,398,369,395]" captionStart="FIGURE 4" captionStartId="11.[151,250,1608,1633]" captionTargetBox="[162,1424,203,1547]" captionTargetId="figure@11.[151,1436,181,1583]" captionTargetPageId="11" captionText="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677877" httpUri="https://zenodo.org/record/3677877/files/figure.png" pageId="12" pageNumber="413">Fig. 4D</figureCitation>
and
<figureCitation id="62972A3F3171D91345E06C1FFE761BB5" box="[453,490,369,395]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="12" pageNumber="413">1D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B665313171D91344E26CFAFC0B1B91" box="[199,919,404,431]" pageId="12" pageNumber="413" type="distribution">
<paragraph id="FA1336BA3171D91344E26CFAFC0B1B91" blockId="12.[151,1437,153,431]" box="[199,919,404,431]" pageId="12" pageNumber="413">
<emphasis id="C8D8EAA83171D91344E26CFAFE571B91" bold="true" box="[199,459,404,431]" pageId="12" pageNumber="413">
Distribution (
<figureCitation id="62972A3F3171D913454D6CFAFE211B91" box="[360,445,404,431]" captionStart="FIGURE 21" captionStartId="37.[151,250,1173,1198]" captionTargetBox="[159,1429,193,1137]" captionTargetId="figure@37.[151,1436,181,1148]" captionTargetPageId="37" captionText="FIGURE 21. Distributional map of the species of H. quadridens group." figureDoi="http://doi.org/10.5281/zenodo.3677911" httpUri="https://zenodo.org/record/3677911/files/figure.png" pageId="12" pageNumber="413">Fig. 21</figureCitation>
).
</emphasis>
<collectingCountry id="82BB762A3171D91345F76CFBFDC21B91" box="[466,606,405,431]" name="Ecuador" pageId="12" pageNumber="413">ECUADOR</collectingCountry>
:
<collectingRegion id="3868F8583171D913464D6CFBFD2A1B91" box="[616,694,405,431]" country="Ecuador" name="Carchi" pageId="12" pageNumber="413">Carchi</collectingRegion>
,
<collectingRegion id="3868F8583171D91346E46CFBFCAE1B91" box="[705,818,405,431]" country="Ecuador" name="Pichincha" pageId="12" pageNumber="413">Pichincha</collectingRegion>
, Bolívar.
</paragraph>
</subSubSection>
</treatment>
</document>
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