217 lines
26 KiB
XML
217 lines
26 KiB
XML
<document ID-CLB-Dataset="6286" ID-DOI="http://dx.doi.org/10.3897/zookeys.1042.66576" ID-GBIF-Dataset="598dbded-7da1-48b8-b8fa-2e648b22a198" ID-PMC="PMC8195948" ID-Pensoft-Pub="1313-2970-1042-1" ID-Pensoft-UUID="0930CA7A3EDB571AA10D026EF39B1739" ID-PubMed="34163286" ID-ZooBank="B0B0F76E89504A0AB45D701730486B56" ModsDocID="1313-2970-1042-1" checkinTime="1622838178301" checkinUser="pensoft" docAuthor="Zhang, Wen-Xuan, Hu, Fang-Shuo & Yin, Zi-Wei" docDate="2021" docId="230218FD121653099DAC3E7E6D46147E" docLanguage="en" docName="ZooKeys 1042: 1-22" docOrigin="ZooKeys 1042" docPubDate="2021-06-04" docSource="http://dx.doi.org/10.3897/zookeys.1042.66576" docTitle="Horniella nantouensis Zhang, Hu & Yin 2021, sp. nov." docType="treatment" docUuid="BAC47C36-A9D6-4D70-9A6D-2B5281C4904F" docUuidSource="ZooBank" docVersion="6" id="0930CA7A3EDB571AA10D026EF39B1739" lastPageNumber="1" masterDocId="0930CA7A3EDB571AA10D026EF39B1739" masterDocTitle="Six new species of Horniella Raffray from the Oriental region (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="1" updateTime="1720468530034" updateUser="ExternalLinkService">
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<mods:titleInfo id="010FE101786DFC1725376F4BCEECDD9E">
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<mods:title id="C0D5E31D1471687A99FFAD6F85EC7721">Six new species of Horniella Raffray from the Oriental region (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:name id="AF78F01BBED125B941D83D91E369AA6B" type="personal">
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<mods:roleTerm id="1C947D5969E51230EC1DE83694D5D0BC">Author</mods:roleTerm>
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<mods:namePart id="D4BC7CC9128AF9A206D47A6141F9BF3C">Zhang, Wen-Xuan</mods:namePart>
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<mods:affiliation id="9668DA1D3F38E778BB95C1546182D40A">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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<mods:roleTerm id="A2976B6C6B00B20651367B54EB110AEE">Author</mods:roleTerm>
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<mods:namePart id="E38D360AB469D43187CB34632EC4EF6E">Hu, Fang-Shuo</mods:namePart>
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<mods:nameIdentifier id="B549B89603DE06A7AA13BF1ECA61A6F5" type="ORCID">https://orcid.org/0000-0003-1269-8904</mods:nameIdentifier>
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<mods:affiliation id="6D520564474171DEEA090F3B145F5FE4">Department of Entomology, National Chung Hsing University 145 Xingda Rd., South District, Taichung City 402, Taiwan, China</mods:affiliation>
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<mods:name id="218E828C87324A0435C01538D119751D" type="personal">
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<mods:namePart id="075463DE16A0A6D38F230B83440E5406">Yin, Zi-Wei</mods:namePart>
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<mods:nameIdentifier id="91686A2E4B5203678C82DDC68CC377E6" type="ORCID">https://orcid.org/0000-0001-6659-9448</mods:nameIdentifier>
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<mods:affiliation id="FAD863C7BA3B2FC22902B9565E4C8C5F">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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<mods:nameIdentifier id="DB5D98D08BF670E0BF1890CA3BE02AC6" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:title id="75A327C6FE0E7FB6CE7A05B0104A9E86">ZooKeys</mods:title>
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<mods:part id="119A0863EE2B6C7F1BC84BAA31B25F4F">
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<mods:date id="6C158B67D3B96A837E07925980E63647">2021</mods:date>
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<mods:number id="13575086E06FF859276185F8E9C19883">2021-06-04</mods:number>
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<mods:number id="F8393DDAD63C192E09742441680C0171">1042</mods:number>
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<mods:classification id="8CB9C86A51B35D96918CD382EB72F426">journal article</mods:classification>
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<mods:identifier id="942D73BAC19D28FB956D36EBF50FB084" type="DOI">http://dx.doi.org/10.3897/zookeys.1042.66576</mods:identifier>
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<treatment id="230218FD121653099DAC3E7E6D46147E" ID-GBIF-Taxon="182494866" LSID="urn:lsid:zoobank.org:act:BAC47C36-A9D6-4D70-9A6D-2B5281C4904F" httpUri="http://treatment.plazi.org/id/230218FD121653099DAC3E7E6D46147E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="5F62F2B436A0E63F4FFE3019A4AAE785" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="47ABB6D8C62AEC3437A8FF9B32AD4051" pageId="0" pageNumber="1">
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<taxonomicName id="49ADDD092EF81A6E8EB5A3FEB74FB214" ID-CoL="8Y4MF" LSID="http://zoobank.org/BAC47C36-A9D6-4D70-9A6D-2B5281C4904F" authority="Zhang, Hu & Yin" authorityName="Zhang, Hu & Yin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Horniella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Horniella nantouensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nantouensis" status="sp. nov.">Horniella nantouensis Zhang, Hu & Yin</taxonomicName>
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<taxonomicNameLabel id="70273F37A30389B0F29E699AA2625B68" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="8E678832A870649579B0D92AE774624B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus of Horniella species A H. khasiensis sp. nov. B H. nantouensis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1048.66576.figure4" httpUri="https://binary.pensoft.net/fig/551737" pageId="0" pageNumber="1">Figures 4B</figureCitation>
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<figureCitation id="892BD1BBB3A057465A57518379FE05F1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">, 6</figureCitation>
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<figureCitation id="CCDBCE2C3D628ED98542C9A45AD8F775" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genitalia of Horniella species, in dorsal view A H. bifurca sp. nov. B H. nantouensis sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1048.66576.figure10" httpUri="https://binary.pensoft.net/fig/551743" pageId="0" pageNumber="1">, 10B</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="5EACA125E7E1EBEFE58AF19AE9CBFE5D" pageId="0" pageNumber="1" type="type material">
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||
<paragraph id="A9FE48D3EF0D91B1072FCECFFDDF9614" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="E3E8C10E1EAA027292989832116D4305" pageId="0" pageNumber="1">
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<emphasis id="D94627B08B7291D4F7D7F5370285BBF7" bold="true" pageId="0" pageNumber="1">
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<emphasis id="4A35FAA7E4C2479E967267DDC4454B5E" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: China
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</emphasis>
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: ♂, 'TAIWAN: Nantou County, Huisun Forest Reserve [惠荪林场], track to Xiaochushan Mt.,
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<geoCoordinate id="68E41B71B4BA38EFC0BB8D8F911A64A7" degrees="24.0745" direction="north" orientation="latitude" precision="5" value="24.0745">24.0745N</geoCoordinate>
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,
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<geoCoordinate id="125B7BAB703E9C12D067F24D29B6C9DB" degrees="121.0366" direction="east" orientation="longitude" precision="5" value="121.0366">121.0366E</geoCoordinate>
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; 1150 m, 4.v.2019;
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<normalizedToken id="59C72AB83016D17C52C19B13149B0770" originalValue="Damaška">Damaska</normalizedToken>
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,
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<normalizedToken id="9A1E8ABCC396E0C7DCFD802E2ED8F302" originalValue="Fikáček">Fikacek</normalizedToken>
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, Hu & Liu lgt., 2019-TW14 / primary forest on the slope with sparse understory; sifting of small accumulations of leaves / Huisun Leaf Litter Beetles Project, Additional specimen: HS1-034 /
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<taxonomicName id="17AC8A278D1726D98F4B18ECC80FD289" class="Insecta" family="Hornelliidae" genus="Horniella" kingdom="Animalia" lsidName="HORNIELLA" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="881A16E50CC486CB3D80A597221FB948" italics="true" pageId="0" pageNumber="1">HORNIELLA</emphasis>
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</taxonomicName>
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sp., P.
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<normalizedToken id="154577DC851E8229642E6EF16850CF24" originalValue="Hlaváč">Hlavac</normalizedToken>
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det., 2019' (NMNS).
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<emphasis id="71E0F0FD6DA734B23EF20F9D2FD16E8E" bold="true" pageId="0" pageNumber="1">
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<emphasis id="740CB5E5DEBC1DF218247FB586DB1156" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: China
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</emphasis>
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: 1 ♀, same data as holotype, except 'Huisun Leaf Litter Beetles Project, Additional specimen: HS1-035' (NMNS); 1 ♀, 'same locality and date, except '
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<geoCoordinate id="20D790F959443444C8B9E50CD1F1C306" degrees="24.0826" direction="north" orientation="latitude" precision="5" value="24.0826">24.0826N</geoCoordinate>
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,
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<geoCoordinate id="D53CC333B5870702C53FDF13900AF4AC" degrees="121.0316" direction="east" orientation="longitude" precision="5" value="121.0316">121.0316E</geoCoordinate>
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; 1050 m, 2019-TW15' / sparse secondary forest with dense understory incl. tree ferns on the margin of a tree plantation: sifting / Huisun Leaf Litter Beetles Project, Additional specimen: HS2-041' (NMNS); 1 ♂, 'Tehuashe (900 m), Nantou, Taiwan,
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<normalizedToken id="F05472E0C0C94A5A1E533C3BE29676D2" originalValue="南投县德化社">南投县德化社</normalizedToken>
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(Tehuashe), 14.xi.2000, Hiroshi Sugaya leg. (in the leaf litter)' (MHNG); 1 ♀, same data as previous, except
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<normalizedToken id="E22DF332DA8500FE6085E0E8E7369838" originalValue="‘4-5.v.(20)01’">'4-5.v.(20)01'</normalizedToken>
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(MHNG); 2 ♀♀, 'Aowanta (1400 m), Nantou, Taiwan,
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<normalizedToken id="760696C4B53F5D31BDC3DC5C46EFB8C8" originalValue="南投县奥万大">南投县奥万大</normalizedToken>
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, 15.xi.2000, Hiroshi Sugaya leg. (in the leaf litter)' (MHNG).
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</paragraph>
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</subSubSection>
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<subSubSection id="C236D69FD06ABD33F2D804A9BAC2F387" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="7B1421E6197DACB978297B36E24BC793" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="08EDB1141034FFB37F168CAEC3B03E5A" pageId="0" pageNumber="1">
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<emphasis id="32D40059E054E4E1B8CB30AC7E57DDE0" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Head longer than wide, with weakly indicated anterolateral genal projection, anterior margin of projection oblique; with short ocular canthus; lateral margin of scape straight, antennomeres 9-11 slightly enlarged. Pronotum rounded at anterolateral margins. Ventral margin of profemur with one short and acute, and one tiny spine at base; protibia with one small preapical denticle. Tergite 1 (IV) with median carina extending posteriorly for approximately 1/3 of tergal length, lacking discal carinae. Aedeagus with asymmetric median lobe, apical part of median lobe narrowed and protruding apicad, apex nearly rounded in dorsal view. Female. Similar to male in external morphology, profemur with two ventral spines near base; genital complex as in Fig.
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<figureCitation id="35F1E5B79B853B6C19E1A0F8B0C5AAE9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genitalia of Horniella species, in dorsal view A H. bifurca sp. nov. B H. nantouensis sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1048.66576.figure10" httpUri="https://binary.pensoft.net/fig/551743" pageId="0" pageNumber="1">10B</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9D6325D0AF4D2D7919E534A9AF98DDD0" pageId="0" pageNumber="1" type="description">
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<paragraph id="265ACE3093EC5CEE8004A0C4956FFD9D" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="28E2705D35585E6BFB91D54E303FB201" pageId="0" pageNumber="1">
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<emphasis id="CC3AAA58AFEE118C393FD30B2CBA63E1" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Body reddish-brown, length 3.68 mm. Head (Fig.
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<figureCitation id="A39EA25EDAB060EAA035636B84C13393" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6A</figureCitation>
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) slightly longer than wide, HL 0.74 mm, HW 0.63 mm; anterolateral genal projection weakly developed, anterior margin of projection oblique; antenna 2.03 mm long, scape lacking expansion at lateral margin, antennomeres 2-8 slightly elongate or moniliform, club loosely formed by apical three moderately enlarged antennomeres, antennomere 11 largest, as long as antennomeres 9 and 10 combined; indistinct lateral postantennal pits present; eyes prominent, each composed of approximately 45 large facets, with pair of short ocular canthi (Fig.
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<figureCitation id="2ED63D05C11B177F6457914E2217E30A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6B</figureCitation>
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).
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</paragraph>
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<caption id="368D75AB286418F8E506F87368B48D6F" doi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
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<paragraph id="2009DE159EECC464256C1EB4DABF1182" pageId="0" pageNumber="1">
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<emphasis id="F4C97490E9DCC8FBB745D689C30090C0" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
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Diagnostic characters of
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<taxonomicName id="29F384EB882724635EEE8C518F67B9FC" authorityName="Zhang, Hu & Yin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Horniella" kingdom="Animalia" lsidName="Horniella nantouensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nantouensis">
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<emphasis id="755703CF51BBB72C34D8CCB54A4B72BB" italics="true" pageId="0" pageNumber="1">Horniella nantouensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="93BEA08CF1E21B8AB654776C9BC0C785" bold="true" pageId="0" pageNumber="1">A</emphasis>
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left half of head, in dorsal view
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<emphasis id="A03F2BAC0FB820D7C5BA72A819A3FA6F" bold="true" pageId="0" pageNumber="1">B</emphasis>
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head, in lateral view
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<emphasis id="590ECB40E9CB9DD9DC202DA3D29CDC87" bold="true" pageId="0" pageNumber="1">C</emphasis>
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protrochanter and profemur
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||
<emphasis id="4B9F885204CBBB81003F2F79FAB3F591" bold="true" pageId="0" pageNumber="1">D</emphasis>
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protibia
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<emphasis id="2A605F5CE00D6AEC5F3A8DEB64B4467F" bold="true" pageId="0" pageNumber="1">E</emphasis>
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mesotrochanter and mesofemur
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||
<emphasis id="924B2C22A070132BB468A2864E6A8E04" bold="true" pageId="0" pageNumber="1">F</emphasis>
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mesotibia
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<emphasis id="1A0556960A0DC8EE6DF9C61AF81F3260" bold="true" pageId="0" pageNumber="1">G-I</emphasis>
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aedeagus, in dorsal (
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||
<emphasis id="EBAFBA9E86998E979D910FF690408B7C" bold="true" pageId="0" pageNumber="1">G</emphasis>
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), lateral (
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<emphasis id="8884E5D71E74BE4AC63E8BA9685ADF8C" bold="true" pageId="0" pageNumber="1">H</emphasis>
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), and ventral (
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<emphasis id="3BEA4FB23ADB8AF2A36435411CEDA3DE" bold="true" pageId="0" pageNumber="1">I</emphasis>
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) view. Scale bars: 0.2 mm (
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<emphasis id="319CA17CBFA0785F168D513755B77A85" bold="true" pageId="0" pageNumber="1">A, B, G-I</emphasis>
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); 0.3 mm in (
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<emphasis id="19007D3C8B68C5E58436629A0877F4F7" bold="true" pageId="0" pageNumber="1">C-F</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph id="BDE8736AB5620A2B09FE1716B4535AE7" pageId="0" pageNumber="1">Pronotum longer than wide, PL 0.77 mm, PW 0.71 mm; widest at apical 1/3; anterolateral margins rounded; disc moderately convex, finely punctate, with distinct median antebasal and lateral antebasal foveae connected by complete transverse antebasal sulcus.</paragraph>
|
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<paragraph id="053300EFEFCFAA1422D5FDEAE04E93E2" pageId="0" pageNumber="1">Elytra much wider than long, EL 0.85 mm, EW 1.33 mm; each elytron with two large, setose basal foveae; discal striae extending from outer basal foveae to middle of elytral length.</paragraph>
|
||
<paragraph id="4A2B7FF94F0067E5801AC3F44391D97F" pageId="0" pageNumber="1">
|
||
Legs elongate; protrochanter (Fig.
|
||
<figureCitation id="2F7FEF83547C031A18B4FC5E62640724" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6C</figureCitation>
|
||
) simple, profemur (Fig.
|
||
<figureCitation id="953E56AE867CC491B19E3294171933AB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6C</figureCitation>
|
||
) with one short and acute, and one tiny ventral spine at base; protibia (Fig.
|
||
<figureCitation id="8B3A523C0C00DDF26D7B1F8E7250C07E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6D</figureCitation>
|
||
) with small preapical spur; mesotrochanter (Fig.
|
||
<figureCitation id="C5A20BCFBE16B6854317619B9A5330DE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6E</figureCitation>
|
||
) and mesofemur (Fig.
|
||
<figureCitation id="4FD873C5DF1C9AD794FC0331D56DEB73" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6E</figureCitation>
|
||
) simple, mesotibia (Fig.
|
||
<figureCitation id="F7B617C5311DC7E186717273B7757F61" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6F</figureCitation>
|
||
) with thick apical setae.
|
||
</paragraph>
|
||
<paragraph id="31D4595F748E75951490486C7F072579" pageId="0" pageNumber="1">Abdomen slightly longer than broad, broadest at lateral margins of tergite 1 (IV), AL 1.32 mm, AW 1.28 mm; tergite 1 (IV) slightly longer than tergites 2 (V), with median carina extending to near basal 1/3 of tergal length, lacking discal carinae, tergite 2 (V) lacking carina, tergites 2-4 (V-VII) each with small basolateral foveae. Sternite 2 (IV) with broad basal sulcus, lacking mediobasal foveae, basolateral foveae developed as large cuticular pockets, with two pairs of antebasal nodules, sternites 3-5 (V-VII) with basolateral foveae, one median and two lateral nodules.</paragraph>
|
||
<paragraph id="058E1956710157FE6C59643F0B08FD43" pageId="0" pageNumber="1">
|
||
Aedeagus (Fig.
|
||
<figureCitation id="E5BC256589A6095670CF6BB27582F4EF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Horniella nantouensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G-I aedeagus, in dorsal (G), lateral (H), and ventral (I) view. Scale bars: 0.2 mm (A, B, G-I); 0.3 mm in (C-F)." figureDoi="10.3897/zookeys.1048.66576.figure6" httpUri="https://binary.pensoft.net/fig/551739" pageId="0" pageNumber="1">6G-I</figureCitation>
|
||
) 0.57 mm long, with asymmetric median lobe, apical portion of median lobe narrowed and greatly protruding apically, apex nearly rounded in dorsal view; endophallus composed of broad membranous part with single broad elongate sclerite at middle.
|
||
</paragraph>
|
||
<paragraph id="3977D7B187A4F2CDE696B7E84CF36282" pageId="0" pageNumber="1">
|
||
<emphasis id="BE5D6021B49B94D644B11233998D13B6" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
|
||
General morphology similar to male, each eye composed of approximately 40 facets; profemur each with two distinct ventral spines near base, protibia lacking spur. Measurements (as for male): BL 3.68-3.72 mm, HL 0.74 mm, HW 0.63-0.65 mm, PL 0.72-0.73 mm, PW 0.74-0.75 mm, EL 0.87-0.9 mm, EW 1.33 mm, AL 1.35 mm, AW 1.32-1.33 mm. Genital complex (Fig.
|
||
<figureCitation id="DC15215297F28330BC336A3858140165" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genitalia of Horniella species, in dorsal view A H. bifurca sp. nov. B H. nantouensis sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1048.66576.figure10" httpUri="https://binary.pensoft.net/fig/551743" pageId="0" pageNumber="1">10B</figureCitation>
|
||
) with moderately sclerotized central and membranous lateral parts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="771F63F6A9A50418A318DBEAD853E289" pageId="0" pageNumber="1" type="comparisons">
|
||
<paragraph id="CF2964B215C20540109CF10C794E920A" pageId="0" pageNumber="1">Comparative notes.</paragraph>
|
||
<paragraph id="7B7469E23BDB048623B6073591B39A03" pageId="0" pageNumber="1">
|
||
This species is placed as a member of the
|
||
<taxonomicName id="3C7DF34E92DBA88C3C5873073B7D0E5E" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. hirtella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hirtella">
|
||
<emphasis id="A5FE1FD2EDF59D8A1E97289AEF283E25" italics="true" pageId="0" pageNumber="1">H. hirtella</emphasis>
|
||
</taxonomicName>
|
||
group. The new species is similar to
|
||
<taxonomicName id="021B30A88F5A47BDDA01F615ABC30A06" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. simplaria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplaria">
|
||
<emphasis id="8DAEE80748C92372DC3DE2B46431B78D" italics="true" pageId="0" pageNumber="1">H. simplaria</emphasis>
|
||
</taxonomicName>
|
||
Yin & Li by the male having similar anterolateral genal projections, and presence of two ventral spines of profemur. They can be otherwise clearly separated by the larger body size (3.68 mm vs 3.23 mm), lack of a mesal hook-like spine of the protibia (present in
|
||
<taxonomicName id="D249DCB848944EF572D9E1B9AD3727DA" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. simplaria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="simplaria">
|
||
<emphasis id="772E6D1D79B00DAB04BF26844D4B86E1" italics="true" pageId="0" pageNumber="1">H. simplaria</emphasis>
|
||
</taxonomicName>
|
||
), and the different shape and structure of the aedeagus of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="152C8019FF4133C691BF4E34FFB4B9A7" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="E8A2FAC8216CCD6EFC60E0E9E473713F" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="23EC3F35B0F67A6C6D60599F1BDFCB4A" pageId="0" pageNumber="1">China: Taiwan.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28E0703FDEE7338C3AFA89BEB6BF7308" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="A6FB82AF08392B88C3074695ADBE7DB0" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="6E5344CF014716F9692D28A840CA89F2" pageId="0" pageNumber="1">The new specific is named after its type locality, Nantou County.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |