109 lines
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109 lines
15 KiB
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<document id="447502861039BCCC8E6560205D33E18B" ID-DOI="10.26879/1323" ID-ISSN="1094-8074" ID-Zenodo-Dep="10962296" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712859958310" checkinUser="felipe" docAuthor="Villa, Andrea, Macaluso, Loredana & Mörs, Thomas" docDate="2024" docId="03F52665D074FFA8FE309AC3FA7F74B3" docLanguage="en" docName="PalaeontolElectron.27.1.a3.pdf" docOrigin="Palaeontologia Electronica (a 3) 27 (1)" docSource="http://dx.doi.org/10.26879/1323" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Ranidae Batsch 1796" docType="treatment" docVersion="1" lastPageNumber="33" masterDocId="FFCC5E1DD054FF88FFA69832FFDE760C" masterDocTitle="Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe" masterLastPageNumber="56" masterPageNumber="1" pageNumber="33" updateTime="1713792755732" updateUser="felipe" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="0366A50E1217E2E6EAB70D5F280E7F81">Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe</mods:title>
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<mods:namePart id="789395B68C41038C173A76AF94518C08">Villa, Andrea</mods:namePart>
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<mods:namePart id="EEB1CE622C6A9C205D3511956E3F6D4E">Macaluso, Loredana</mods:namePart>
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<mods:namePart id="AA993828A6C36287C6596B31D164062A">Mörs, Thomas</mods:namePart>
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<mods:title id="3A997154E3EF1403E57D708EFAB53009">Palaeontologia Electronica</mods:title>
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<mods:part id="578C576D2B722A540A5E8516466A9A49">
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<mods:date id="C40FBFFC3D9ED2D44D70B64947EF01C2">2024</mods:date>
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<mods:title id="ABD49427BA3B3B86C019A6F922E65BD8">a 3</mods:title>
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<mods:number id="9C5A9DC7B5DBA526303B56231841959C">27</mods:number>
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<mods:url id="4ED175595B016821CC905101525A7422">http://dx.doi.org/10.26879/1323</mods:url>
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<mods:identifier id="5605197C6C8920D046E5FCFB50A213CB" type="ISSN">1094-8074</mods:identifier>
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<treatment id="03F52665D074FFA8FE309AC3FA7F74B3" LSID="urn:lsid:plazi:treatment:03F52665D074FFA8FE309AC3FA7F74B3" httpUri="http://treatment.plazi.org/id/03F52665D074FFA8FE309AC3FA7F74B3" lastPageNumber="33" pageId="32" pageNumber="33">
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<subSubSection id="C346C4F8D074FFA8FE309AC3FD9D7506" box="[406,579,753,778]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BE39773D074FFA8FE309AC3FD9D7506" blockId="32.[406,579,753,810]" box="[406,579,753,778]" pageId="32" pageNumber="33">
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<heading id="D0AB201FD074FFA8FE309AC3FD9D7506" box="[406,579,753,778]" centered="true" fontSize="10" level="2" pageId="32" pageNumber="33" reason="8">
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<taxonomicName id="4C5CECF0D074FFA8FE309AC3FE277506" ID-CoL="FJM" authorityName="Batsch" authorityYear="1796" box="[406,505,753,778]" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="family">Ranidae</taxonomicName>
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indet.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D074FFA8FE3D9B23FDE37526" box="[411,573,784,810]" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BE39773D074FFA8FE3D9B23FDE37526" blockId="32.[406,579,753,810]" box="[411,573,784,810]" pageId="32" pageNumber="33">
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<figureCitation id="13678BF6D074FFA8FE3D9B23FDD27526" box="[411,524,784,810]" captionStart="FIGURE 22" captionStartId="31.[195,286,1181,1204]" captionTargetBox="[193,1431,172,1154]" captionTargetId="figure-206@31.[187,1437,165,1161]" captionTargetPageId="31" captionText="FIGURE 22. Ranids from Hambach: right ilium (IPB-HaH 2307) of Pelophylax sp. in lateral (A) and medial (B) views; left ilium (IPB-HaH 2387) of Pelophylax sp. in lateral (C) and medial (D) views; left ilium (IPB-HaH 2321) of Rana sp. in lateral (E) and medial (F) views; sacral vertebra (IPB-HaR 2018) of Ranidae indet. in dorsal (G) and ventral (H) views; sacral vertebra (IPB-HaR 2019) of Ranidae indet. in dorsal (I) and ventral (J) views; left scapula (IPB-HaH 2323) of Ranidae indet. in ventral (K) and dorsal (L) views; left ilium (IPB-HaR 2087) of Ranidae indet. in lateral (M) and medial (N) views. Scale bars equal 1 mm. Abbreviations: dc, dorsal crest; dt, dorsal tubercle." figureDoi="http://doi.org/10.5281/zenodo.10962341" httpUri="https://zenodo.org/record/10962341/files/figure.png" pageId="32" pageNumber="33">Figure 22</figureCitation>
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G-N
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D074FFA8FF669B09FE9775F8" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BE39773D074FFA8FF669B09FE9775F8" blockId="32.[192,793,827,1908]" pageId="32" pageNumber="33">
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<emphasis id="B9284B61D074FFA8FF669B09FEF27558" bold="true" box="[192,300,827,852]" pageId="32" pageNumber="33">Material.</emphasis>
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Hambach 6C: nine sacral vertebrae (IPB-HaH 2193/2194, IPB-HaH 2209/2215); one scapula (IPB-HaH 2323). Hambach 11: one trunk vertebra (IPB-HaR 2184); three sacral vertebrae (IPB-HaR 2018/2019; IPB-HaR 2031); one ilium (IPB-HaR 2087).
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D074FFA8FF669BC9FDFF7118" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BE39773D074FFA8FF669BC9FD207258" blockId="32.[192,793,827,1908]" pageId="32" pageNumber="33">
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<emphasis id="B9284B61D074FFA8FF669BC9FE847218" bold="true" box="[192,346,1019,1044]" pageId="32" pageNumber="33">Description.</emphasis>
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IPB-HaR 2184 is a small and amphicoelous trunk vertebra, with a short neural arch and laterally directed transverse processes.
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</paragraph>
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<paragraph id="8BE39773D074FFA8FF5C9C69FCCD72B8" blockId="32.[192,793,827,1908]" pageId="32" pageNumber="33">
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Sacral vertebrae (
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<figureCitation id="13678BF6D074FFA8FE6C9C69FDE17279" box="[458,575,1115,1141]" captionStart="FIGURE 22" captionStartId="31.[195,286,1181,1204]" captionTargetBox="[193,1431,172,1154]" captionTargetId="figure-206@31.[187,1437,165,1161]" captionTargetPageId="31" captionText="FIGURE 22. Ranids from Hambach: right ilium (IPB-HaH 2307) of Pelophylax sp. in lateral (A) and medial (B) views; left ilium (IPB-HaH 2387) of Pelophylax sp. in lateral (C) and medial (D) views; left ilium (IPB-HaH 2321) of Rana sp. in lateral (E) and medial (F) views; sacral vertebra (IPB-HaR 2018) of Ranidae indet. in dorsal (G) and ventral (H) views; sacral vertebra (IPB-HaR 2019) of Ranidae indet. in dorsal (I) and ventral (J) views; left scapula (IPB-HaH 2323) of Ranidae indet. in ventral (K) and dorsal (L) views; left ilium (IPB-HaR 2087) of Ranidae indet. in lateral (M) and medial (N) views. Scale bars equal 1 mm. Abbreviations: dc, dorsal crest; dt, dorsal tubercle." figureDoi="http://doi.org/10.5281/zenodo.10962341" httpUri="https://zenodo.org/record/10962341/files/figure.png" pageId="32" pageNumber="33">Figure 22</figureCitation>
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G-J) are small and provided with one anterior and two posterior condyles. They have cylindrical transverse processes.
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</paragraph>
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<paragraph id="8BE39773D074FFA8FF5C9C89FE8A7378" blockId="32.[192,793,827,1908]" pageId="32" pageNumber="33">
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IPB-HaH 2323 (
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<figureCitation id="13678BF6D074FFA8FE139C89FDF772D8" box="[437,553,1211,1236]" captionStart="FIGURE 22" captionStartId="31.[195,286,1181,1204]" captionTargetBox="[193,1431,172,1154]" captionTargetId="figure-206@31.[187,1437,165,1161]" captionTargetPageId="31" captionText="FIGURE 22. Ranids from Hambach: right ilium (IPB-HaH 2307) of Pelophylax sp. in lateral (A) and medial (B) views; left ilium (IPB-HaH 2387) of Pelophylax sp. in lateral (C) and medial (D) views; left ilium (IPB-HaH 2321) of Rana sp. in lateral (E) and medial (F) views; sacral vertebra (IPB-HaR 2018) of Ranidae indet. in dorsal (G) and ventral (H) views; sacral vertebra (IPB-HaR 2019) of Ranidae indet. in dorsal (I) and ventral (J) views; left scapula (IPB-HaH 2323) of Ranidae indet. in ventral (K) and dorsal (L) views; left ilium (IPB-HaR 2087) of Ranidae indet. in lateral (M) and medial (N) views. Scale bars equal 1 mm. Abbreviations: dc, dorsal crest; dt, dorsal tubercle." figureDoi="http://doi.org/10.5281/zenodo.10962341" httpUri="https://zenodo.org/record/10962341/files/figure.png" pageId="32" pageNumber="33">Figure 22</figureCitation>
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K-L) is an elongated scapula. It has no ridges on the anterior margin, but it is provided with a ridge on the inner surface. The cavitas glenoidalis opens in posterior direction and is partially hidden by the pars acromialis in ventral view.
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</paragraph>
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<paragraph id="8BE39773D074FFA8FF5C9D49FDFF7118" blockId="32.[192,793,827,1908]" pageId="32" pageNumber="33">
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The ilium (
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<figureCitation id="13678BF6D074FFA8FEDF9D49FE317398" box="[377,495,1403,1428]" captionStart="FIGURE 22" captionStartId="31.[195,286,1181,1204]" captionTargetBox="[193,1431,172,1154]" captionTargetId="figure-206@31.[187,1437,165,1161]" captionTargetPageId="31" captionText="FIGURE 22. Ranids from Hambach: right ilium (IPB-HaH 2307) of Pelophylax sp. in lateral (A) and medial (B) views; left ilium (IPB-HaH 2387) of Pelophylax sp. in lateral (C) and medial (D) views; left ilium (IPB-HaH 2321) of Rana sp. in lateral (E) and medial (F) views; sacral vertebra (IPB-HaR 2018) of Ranidae indet. in dorsal (G) and ventral (H) views; sacral vertebra (IPB-HaR 2019) of Ranidae indet. in dorsal (I) and ventral (J) views; left scapula (IPB-HaH 2323) of Ranidae indet. in ventral (K) and dorsal (L) views; left ilium (IPB-HaR 2087) of Ranidae indet. in lateral (M) and medial (N) views. Scale bars equal 1 mm. Abbreviations: dc, dorsal crest; dt, dorsal tubercle." figureDoi="http://doi.org/10.5281/zenodo.10962341" httpUri="https://zenodo.org/record/10962341/files/figure.png" pageId="32" pageNumber="33">Figure 22</figureCitation>
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M-N) is small and poorly preserved, missing completely the shaft. It has a large and subcircular acetabular fossa. Dorsal to the latter, a deep supraacetabular fossa is present. The dorsal tubercle is also present, even though mostly broken away. Nevertheless, it appears moderately distinct. Anteriorly, it is in continuation with a dorsal crest. Both the dorsal and ventral acetabular expansions are broken, but the former appears well developed based on what is preserved. Medially, there is no interiliac groove or tubercle. The ilioischiatic juncture seems rather low and large, but it is not completely preserved.
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D074FFA8FF669F29FA7F74B3" pageId="32" pageNumber="33" type="discussion">
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<paragraph id="8BE39773D074FFA8FF669F29FA7F74B3" blockId="32.[192,793,827,1908]" lastBlockId="32.[840,1441,166,703]" pageId="32" pageNumber="33">
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<emphasis id="B9284B61D074FFA8FF669F29FEE67138" bold="true" box="[192,312,1819,1844]" pageId="32" pageNumber="33">Remarks.</emphasis>
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All these specimens present a combination of features supporting attribution to indeterminate ranids according to the criteria presented by
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<bibRefCitation id="EFCDEA82D074FFA8FCEE9894FC3476B3" author="Bailon, S." box="[840,1002,166,191]" pageId="32" pageNumber="33" refId="ref29899" refString="Bailon, S. 1999. Differenciation osteologique des anoures (Amphibia, Anura) de France, p. 1 - 41. In Desse, J. and Desse-Berset, N. (eds.), Fiches d'osteologie animale pour l'Archeologie, Serie C: Varia, 1. APDCA, Antibes." type="journal volume" year="1999">Bailon (1999)</bibRefCitation>
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. For the trunk vertebra, this combination includes amphicoely, the short neural arch, and the laterally directed processes; this further suggests that it represents the eighth vertebra in the column (
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<bibRefCitation id="EFCDEA82D074FFA8FC799914FBA17733" author="Bailon, S." box="[991,1151,294,319]" pageId="32" pageNumber="33" refId="ref29899" refString="Bailon, S. 1999. Differenciation osteologique des anoures (Amphibia, Anura) de France, p. 1 - 41. In Desse, J. and Desse-Berset, N. (eds.), Fiches d'osteologie animale pour l'Archeologie, Serie C: Varia, 1. APDCA, Antibes." type="journal volume" year="1999">Bailon, 1999</bibRefCitation>
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). In the sacral vertebra, the diagnostic combination consists of the anterior condyle, the two posterior condyles, and the cylindrical processes. For the scapula, significant are the elongation, the absence of crista anterior, the cavitas glenoidalis hidden in ventral view, as well as the presence of an inner ridge on scapula. The attribution of the ilium is supported by the presence of the dorsal crest and the absence of interiliac tubercles and grooves. The low and large juncture could hint at green frogs for the ilium (
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<bibRefCitation id="EFCDEA82D074FFA8FAA79A54FC587493" author="Gleed-Owen, C. P." pageId="32" pageNumber="33" refId="ref32637" refString="Gleed-Owen, C. P. 1998. Quaternary herpetofaunas of the British Isles: taxonomic descriptions, palaeoenvironmental reconstructions, and biostratigraphic implications. Unpublished PhD Thesis, Coventry University, Coventry, UK." type="book" year="1998">Gleed-Owen, 1998</bibRefCitation>
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), but it is not clear how much this apparent morphology may be influenced by the preservation.
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</paragraph>
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</subSubSection>
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</treatment>
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