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<document id="7117C8C2A70738EAB5AFFCDFCC4BBC91" ID-CLB-Dataset="5940" ID-DOI="10.11646/zootaxa.3761.1.1" ID-GBIF-Dataset="5444e27e-3e2a-429b-931b-823a2086e3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909057" ID-ZooBank="82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623107451891" checkinUser="felipe" docAuthor="Luk, Stephen P. L. &amp; Marshall, Stephen A." docDate="2014" docId="038487F1FF8FFF98FDC7F8ECFE310A30" docLanguage="en" docName="zt03761p156.pdf" docOrigin="Zootaxa 3761 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aptilotella angela Luk &amp; Marshall 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="FFBDFF89FFA5FFB3FD50FE2BFFC70D18" masterDocTitle="A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="43" updateTime="1699002622211" updateUser="ExternalLinkService">
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<mods:title id="C59D3BD000293CDA4DB5C754BDD425BF">A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)</mods:title>
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<mods:namePart id="55802390F478DD471982990BDA1CD401">Luk, Stephen P. L.</mods:namePart>
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<mods:namePart id="07AF58FFBFC9BB2C0C8DEEF2199B964C">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="D3FFC4C4AB1940E447871E5483361092" type="email">luk@uoguelph.ca</mods:nameIdentifier>
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<mods:part id="23771BAEC3B68EF2DA7A91FA25E5C118">
<mods:date id="6E0B341B815DBC8B145FE9926E70C98E">2014</mods:date>
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<treatment id="038487F1FF8FFF98FDC7F8ECFE310A30" ID-DOI="http://doi.org/10.5281/zenodo.4909151" ID-GBIF-Taxon="183203084" ID-Zenodo-Dep="4909151" LSID="urn:lsid:plazi:treatment:038487F1FF8FFF98FDC7F8ECFE310A30" httpUri="http://treatment.plazi.org/id/038487F1FF8FFF98FDC7F8ECFE310A30" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="C337656CFF8FFF99FDC7F8ECFD550BFA" box="[151,658,1735,1762]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8B9236E7FF8FFF99FDC7F8ECFD550BFA" blockId="42.[151,658,1735,1795]" box="[151,658,1735,1762]" pageId="42" pageNumber="43">
<heading id="D0DA818BFF8FFF99FDC7F8ECFD550BFA" bold="true" box="[151,658,1735,1762]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="4">
<emphasis id="B959EAF5FF8FFF99FDC7F8ECFD550BFA" bold="true" box="[151,658,1735,1762]" pageId="42" pageNumber="43">
<taxonomicName id="4C2D4D64FF8FFF99FDC7F8ECFD840BFA" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,579,1735,1762]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="angela" status="sp. nov.">
<emphasis id="B959EAF5FF8FFF99FDC7F8ECFEAB0BF9" bold="true" box="[151,364,1735,1761]" italics="true" pageId="42" pageNumber="43">Aptilotella angela</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<taxonomicNameLabel id="A26A578EFF8FFF99FF00F8E3FD550BFA" box="[592,658,1736,1762]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8FFF98FDC7F8C1FC23084B" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8B9236E7FF8FFF99FDC7F8C1FE5E0A1B" blockId="42.[151,658,1735,1795]" box="[151,409,1770,1795]" pageId="42" pageNumber="43">
<figureCitation id="13162A62FF8FFF99FDC7F8C1FEC70A1B" box="[151,256,1770,1795]" captionStart="FIGURE 2" captionStartId="3.[151,249,1879,1901]" captionTargetId="figure-236@3.[503,1084,1404,1862]" captionTargetPageId="3" captionText="FIGURE 2. Aptilotella angela Luk &amp; Marshall, sp. n., paratype male." figureDoi="http://doi.org/10.5281/zenodo.5042237" httpUri="https://zenodo.org/record/5042237/files/figure.png" pageId="42" pageNumber="43">Figures 2</figureCitation>
and
<figureCitation id="13162A62FF8FFF99FC66F8C1FE5E0A1B" box="[310,409,1770,1795]" captionStart-0="FIGURES 306309" captionStart-1="FIGURES 310312" captionStart-2="FIGURES 313316" captionStartId-0="145.[151,264,1249,1272]" captionStartId-1="146.[151,264,1177,1200]" captionStartId-2="147.[151,264,1297,1320]" captionTargetBox-0="[391,1201,213,1213]" captionTargetBox-1="[420,1160,201,1151]" captionTargetBox-2="[385,1202,201,1272]" captionTargetId-0="figure-42@145.[376,1211,193,1228]" captionTargetId-1="figure-81@146.[415,1172,193,1157]" captionTargetId-2="figure-51@147.[377,1211,193,1276]" captionTargetPageId-0="145" captionTargetPageId-1="146" captionTargetPageId-2="147" captionText-0="FIGURES 306309. Aptilotella angela Luk &amp; Marshall, sp. n.: 306—male terminalia, posterior; 307—male terminalia, left lateral; 308—male synsternite 6+7; 309—male sternite 5." captionText-1="FIGURES 310312. Aptilotella angela Luk &amp; Marshall, sp. n.: 310—aedeagus with postgonites removed; 311—left postgonite; 312—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." captionText-2="FIGURES 313316. Aptilotella angela Luk &amp; Marshall, sp. n.: 313—female terminalia, dorsal; 314—female terminalia, left lateral; 315—female terminalia, ventral; 316—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct, S8 = sternite 8." figureDoi-0="http://doi.org/10.5281/zenodo.4909308" figureDoi-1="http://doi.org/10.5281/zenodo.4909310" figureDoi-2="http://doi.org/10.5281/zenodo.4909312" httpUri-0="https://zenodo.org/record/4909308/files/figure.png" httpUri-1="https://zenodo.org/record/4909310/files/figure.png" httpUri-2="https://zenodo.org/record/4909312/files/figure.png" pageId="42" pageNumber="43">306316</figureCitation>
</paragraph>
<paragraph id="8B9236E7FF8FFF98FDC7F919FB500CB5" blockId="42.[151,1436,1842,2011]" lastBlockId="43.[151,1436,151,1832]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FDC7F919FEE40A53" bold="true" box="[151,291,1842,1867]" italics="true" pageId="42" pageNumber="43">Description.</emphasis>
Habitus as in
<figureCitation id="13162A62FF8FFF99FC81F919FDF20A53" box="[465,565,1842,1867]" captionStart="FIGURE 2" captionStartId="3.[151,249,1879,1901]" captionTargetId="figure-236@3.[503,1084,1404,1862]" captionTargetPageId="3" captionText="FIGURE 2. Aptilotella angela Luk &amp; Marshall, sp. n., paratype male." figureDoi="http://doi.org/10.5281/zenodo.5042237" httpUri="https://zenodo.org/record/5042237/files/figure.png" pageId="42" pageNumber="43">Figure 2</figureCitation>
. Body length
<quantity id="4CD59B02FF8FFF99FF8BF919FCAF0A52" box="[731,872,1842,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="42" pageNumber="43" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.11.5 mm</quantity>
. Head ground color yellow-orange. Orbital plate shining, darkened behind antennal insertions, with a somewhat paler spot in ocular emargination; interfrontal plate finely rugose; brown medial stripes each approximately one-fifth the width of frons, diffuse along the anterior margin. Ocellar tubercle raised, shining; ocellar bristle subequal to the length of frons. Orbital bristle absent; orbital setulae small, in four pairs. Interfrontal setae long, in two pairs. Face shining; facial tubercle conical, excavation very shallow, with slightly infuscated margin; gena paler, weakly shining and finely rugose. Antenna black, first flagellomere brown. Occiput black. Scutum and scutellum black, shining. Scutum uniformly, sparsely setose. Scutellum bare; three times wider than long, two-thirds the width of scutum. Apical scutellar bristles 1.4 times as long as basal. Pleuron black. Legs bicoloured; coxa black; trochanter orange; femur black except for orange apex; fore and mid tibiae black, orange basally and apically; hind tibia black except for broad orange band basally and along mid-length; tarsus orange, darkening distally; mid tibia with a single distal anterodorsal bristle. Wing rudiment brown. Abdomen black, shining; tergites uniformly setose, basal margin densely microtrichose except for syntergite; sternites finely microtrichose, with rows of setae posteriorly. Cercus red-orange.
</paragraph>
<paragraph id="8B9236E7FF8EFF98FD97FF9CFCD80965" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FF9CFE4E0CC8" bold="true" box="[199,393,439,464]" italics="true" pageId="43" pageNumber="44">Male terminalia.</emphasis>
Sternite 5 (
<figureCitation id="13162A62FF8EFF98FF5FFF9CFDA80CC8" box="[527,623,439,464]" captionStart="FIGURES 306309" captionStartId="145.[151,264,1249,1272]" captionTargetBox="[391,1201,213,1213]" captionTargetId="figure-42@145.[376,1211,193,1228]" captionTargetPageId="145" captionText="FIGURES 306309. Aptilotella angela Luk &amp; Marshall, sp. n.: 306—male terminalia, posterior; 307—male terminalia, left lateral; 308—male synsternite 6+7; 309—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909308" httpUri="https://zenodo.org/record/4909308/files/figure.png" pageId="43" pageNumber="44">Fig. 309</figureCitation>
) medially with a field of minute spinules; posteromedial margin giving rise to a somewhat membranous, shallowly concave shelf, covered in rows of bifid or trifid spinules, flanked on each side by a setal cluster. Synsternite 6+7 (
<figureCitation id="13162A62FF8EFF98FFF3FFD4FCC10F00" box="[675,774,511,536]" captionStart="FIGURES 306309" captionStartId="145.[151,264,1249,1272]" captionTargetBox="[391,1201,213,1213]" captionTargetId="figure-42@145.[376,1211,193,1228]" captionTargetPageId="145" captionText="FIGURES 306309. Aptilotella angela Luk &amp; Marshall, sp. n.: 306—male terminalia, posterior; 307—male terminalia, left lateral; 308—male synsternite 6+7; 309—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909308" httpUri="https://zenodo.org/record/4909308/files/figure.png" pageId="43" pageNumber="44">Fig. 308</figureCitation>
) flanked on each side of medial bridge by a curved arm, posteromedially with pouch-like membrane attaching to the marginal shelf of sternite 5. Cercus (
<figureCitation id="13162A62FF8EFF98F9BDFC0FFA530F25" box="[1261,1428,548,573]" captionStart="FIGURES 306309" captionStartId="145.[151,264,1249,1272]" captionTargetBox="[391,1201,213,1213]" captionTargetId="figure-42@145.[376,1211,193,1228]" captionTargetPageId="145" captionText="FIGURES 306309. Aptilotella angela Luk &amp; Marshall, sp. n.: 306—male terminalia, posterior; 307—male terminalia, left lateral; 308—male synsternite 6+7; 309—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909308" httpUri="https://zenodo.org/record/4909308/files/figure.png" pageId="43" pageNumber="44">Figs. 306, 307</figureCitation>
) slender, curved, three times as long as basal width; base dilated, margin interiorly deeply excavated; inner basal margin with three long setae in a row and one long seta adjacent to the basal two, this seta two-thirds the length of cercus; apex bearing a stout, recurved seta. Surstylus (
<figureCitation id="13162A62FF8EFF98FFA3FCA4FC530FB0" box="[755,916,655,680]" captionStart="FIGURES 306309" captionStartId="145.[151,264,1249,1272]" captionTargetBox="[391,1201,213,1213]" captionTargetId="figure-42@145.[376,1211,193,1228]" captionTargetPageId="145" captionText="FIGURES 306309. Aptilotella angela Luk &amp; Marshall, sp. n.: 306—male terminalia, posterior; 307—male terminalia, left lateral; 308—male synsternite 6+7; 309—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909308" httpUri="https://zenodo.org/record/4909308/files/figure.png" pageId="43" pageNumber="44">Figs. 306, 307</figureCitation>
) conical; outer face anteriorly expanded into a rounded rectangular plate, about 1.5 times as long as the height of cone, bearing scattered sensory setae, and with a tuberculate posteroventral corner bearing numerous divergent setae. Postgonite (
<figureCitation id="13162A62FF8EFF98F94EFCFCFBBA0FE8" box="[1054,1149,727,752]" captionStart="FIGURES 310312" captionStartId="146.[151,264,1177,1200]" captionTargetBox="[420,1160,201,1151]" captionTargetId="figure-81@146.[415,1172,193,1157]" captionTargetPageId="146" captionText="FIGURES 310312. Aptilotella angela Luk &amp; Marshall, sp. n.: 310—aedeagus with postgonites removed; 311—left postgonite; 312—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909310" httpUri="https://zenodo.org/record/4909310/files/figure.png" pageId="43" pageNumber="44">Fig. 311</figureCitation>
) sinuate; descending arm gradually curving and tapering, with several marginal sensory setae; articulatory process for pregonite triangular and rounded; articulatory process for basiphallus short-stalked and knobbed. Hypandrium (
<figureCitation id="13162A62FF8EFF98F9F3FD34FAC30E20" box="[1187,1284,799,824]" captionStart="FIGURES 310312" captionStartId="146.[151,264,1177,1200]" captionTargetBox="[420,1160,201,1151]" captionTargetId="figure-81@146.[415,1172,193,1157]" captionTargetPageId="146" captionText="FIGURES 310312. Aptilotella angela Luk &amp; Marshall, sp. n.: 310—aedeagus with postgonites removed; 311—left postgonite; 312—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909310" httpUri="https://zenodo.org/record/4909310/files/figure.png" pageId="43" pageNumber="44">Fig. 312</figureCitation>
) with medial rod slightly pinched in distal third and apically dilated into a rounded fan; posteromedial fork indistinct but swollen; hypandrial arms irregularly triangular; pregonite minute and ovate. Aedeagus as in
<figureCitation id="13162A62FF8EFF98F842FD4CFA5F0E98" box="[1298,1432,871,896]" captionStart="FIGURES 310312" captionStartId="146.[151,264,1177,1200]" captionTargetBox="[420,1160,201,1151]" captionTargetId="figure-81@146.[415,1172,193,1157]" captionTargetPageId="146" captionText="FIGURES 310312. Aptilotella angela Luk &amp; Marshall, sp. n.: 310—aedeagus with postgonites removed; 311—left postgonite; 312—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909310" httpUri="https://zenodo.org/record/4909310/files/figure.png" pageId="43" pageNumber="44">Figure 310</figureCitation>
. Basiphallus cylindroid; articulatory process for postgonite pointed and anteriorly directed. Ventrobasal sclerite broad. Lateral flanking sclerite fused ventrobasally by a narrow bridge; dorsal margin rolled, straight and nearly parallel, then irregular diverging and ascending in distal third. Ventral flanking sclerites darker; the slender basal article fused along ventral margin of lateral flanking sclerite, preapically giving rise to a slender dorsal arm; the chevron-shaped medial article meeting the basal article interiorly; the rounded distal article with a darkened dorsal margin leading a triangular lobe inward and upward. Curved dorsal sclerite originating from inside distal ventral margin of lateral flanking sclerite and abruptly ascending.
</paragraph>
<paragraph id="8B9236E7FF8EFF98FD97FAACFE020828" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FAACFE6309B8" bold="true" box="[199,420,1159,1184]" italics="true" pageId="43" pageNumber="44">Female terminalia.</emphasis>
Epiproct (
<figureCitation id="13162A62FF8EFF98FF4AFAACFDBD09B8" box="[538,634,1159,1184]" captionStart="FIGURES 313316" captionStartId="147.[151,264,1297,1320]" captionTargetBox="[385,1202,201,1272]" captionTargetId="figure-51@147.[377,1211,193,1276]" captionTargetPageId="147" captionText="FIGURES 313316. Aptilotella angela Luk &amp; Marshall, sp. n.: 313—female terminalia, dorsal; 314—female terminalia, left lateral; 315—female terminalia, ventral; 316—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909312" httpUri="https://zenodo.org/record/4909312/files/figure.png" pageId="43" pageNumber="44">Fig. 313</figureCitation>
) very pale, subpentagonal; finely hairy, with four basal setae. Each half of tergite 8 (
<figureCitation id="13162A62FF8EFF98FC76FA87FE0C09DD" box="[294,459,1196,1221]" captionStart="FIGURES 313316" captionStartId="147.[151,264,1297,1320]" captionTargetBox="[385,1202,201,1272]" captionTargetId="figure-51@147.[377,1211,193,1276]" captionTargetPageId="147" captionText="FIGURES 313316. Aptilotella angela Luk &amp; Marshall, sp. n.: 313—female terminalia, dorsal; 314—female terminalia, left lateral; 315—female terminalia, ventral; 316—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909312" httpUri="https://zenodo.org/record/4909312/files/figure.png" pageId="43" pageNumber="44">Figs. 313315</figureCitation>
) strongly convex, its outer margin concave; with several setae. Cercus three times as long as wide; with two long apical setae. Hypoproct (
<figureCitation id="13162A62FF8EFF98FFBBFAE4FC8C09F0" box="[747,843,1231,1256]" captionStart="FIGURES 313316" captionStartId="147.[151,264,1297,1320]" captionTargetBox="[385,1202,201,1272]" captionTargetId="figure-51@147.[377,1211,193,1276]" captionTargetPageId="147" captionText="FIGURES 313316. Aptilotella angela Luk &amp; Marshall, sp. n.: 313—female terminalia, dorsal; 314—female terminalia, left lateral; 315—female terminalia, ventral; 316—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909312" httpUri="https://zenodo.org/record/4909312/files/figure.png" pageId="43" pageNumber="44">Fig. 315</figureCitation>
) triangular with rounded apex; finely hairy, with two preapical sensory setae. Spermathecae (
<figureCitation id="13162A62FF8EFF98FF09FADFFD7E0815" box="[601,697,1268,1293]" captionStart="FIGURES 313316" captionStartId="147.[151,264,1297,1320]" captionTargetBox="[385,1202,201,1272]" captionTargetId="figure-51@147.[377,1211,193,1276]" captionTargetPageId="147" captionText="FIGURES 313316. Aptilotella angela Luk &amp; Marshall, sp. n.: 313—female terminalia, dorsal; 314—female terminalia, left lateral; 315—female terminalia, ventral; 316—spermathecae. Abbreviations: EP = epiproct, HP = hypoproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909312" httpUri="https://zenodo.org/record/4909312/files/figure.png" pageId="43" pageNumber="44">Fig. 316</figureCitation>
) finely ridged; sclerotized ducts very long, four to five times the diameter of a spermatheca.
</paragraph>
<paragraph id="8B9236E7FF8EFF98FD97FB17FC23084B" blockId="43.[151,1436,151,1832]" box="[199,996,1340,1365]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FB17FEFE084D" bold="true" box="[199,313,1340,1365]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FB17FEF5084D" bold="true" box="[199,306,1340,1365]" italics="true" pageId="43" pageNumber="44">Variation</emphasis>
.
</emphasis>
Some specimens have a slightly reddish tinge in the scutum.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8EFF98FD97FB74FEBA0885" pageId="43" pageNumber="44" type="etymology">
<paragraph id="8B9236E7FF8EFF98FD97FB74FEBA0885" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FB74FE800860" bold="true" box="[199,327,1375,1400]" italics="true" pageId="43" pageNumber="44">Etymology.</emphasis>
The species epithet is a reference to the Ecuadorian city near the páramo habitat where the
<typeStatus id="54968845FF8EFF98F83BFB4BFA5C0860" box="[1387,1435,1376,1400]" pageId="43" pageNumber="44">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8EFF98FD97FB83FD0E0B6D" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8B9236E7FF8EFF98FD97FB83FD7F08FD" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97FB83FEA908D8" bold="true" box="[199,366,1447,1472]" italics="true" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="3B453CBAFF8EFF98FC26FB8CFD7F08FD" ID-GBIF-Occurrence="3311767440" collectingDate="1999-11-02" collectionCode="QCAZ" collectorName="Paramo El Angel &amp; R. S. Anderson." country="Ecuador" elevation="3450" location="El Angel" municipality="Paramo El Angel" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Carchi" typeStatus="holotype">
<typeStatus id="54968845FF8EFF98FC26FB8CFE1908D8" box="[374,478,1447,1472]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2BFD6EFF8EFF98FCB7FB83FE3908A7" box="[487,510,1448,1471]" pageId="43" pageNumber="44" type="male"></specimenCount>
,
<collectionCode id="ED3CAE22FF8EFF98FF5CFB8CFD9B08D8" box="[524,604,1447,1472]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="43" pageNumber="44" type="Museum">QCAZ</collectionCode>
.
<collectingCountry id="F33A7677FF8EFF98FF37FB8CFD3408D8" box="[615,755,1447,1472]" name="Ecuador" pageId="43" pageNumber="44">ECUADOR</collectingCountry>
:
<collectingRegion id="49E9F805FF8EFF98FFAFFB8CFC8B08D8" box="[767,844,1447,1472]" country="Ecuador" name="Carchi" pageId="43" pageNumber="44">Carchi</collectingRegion>
,
<collectorName id="26D85331FF8EFF98FE09FB8CFBD808D8" box="[857,1055,1447,1472]" pageId="43" pageNumber="44">
<collectingMunicipality id="6BF6AC9DFF8EFF98FE09FB8CFBD808D8" box="[857,1055,1447,1472]" pageId="43" pageNumber="44">
Páramo
<location id="8EF2603CFF8EFF98FEE7FB8CFBD808D8" LSID="urn:lsid:plazi:treatment:038487F1FF8FFF98FDC7F8ECFE310A30:8EF2603CFF8EFF98FEE7FB8CFBD808D8" box="[951,1055,1447,1472]" country="Ecuador" municipality="Paramo El Angel" name="El Angel" pageId="43" pageNumber="44" stateProvince="Carchi">El Ángel</location>
</collectingMunicipality>
</collectorName>
,
<locationDeviation id="9A202FC5FF8EFF98F97CFB8CFAF508D8" box="[1068,1330,1447,1472]" location="El Angel" pageId="43" pageNumber="44">
<quantity id="4CD59B02FF8EFF98F97CFB8CFB4D08D8" box="[1068,1162,1447,1472]" metricMagnitude="4" metricUnit="m" metricValue="1.41" pageId="43" pageNumber="44" unit="km" value="14.1">14.1 km</quantity>
NW El Ángel
</locationDeviation>
,
<quantity id="4CD59B02FF8EFF98F86FFB8CFA5008A7" box="[1343,1431,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="43" pageNumber="44" unit="m" value="3450.0">
<elevation id="0000D1D4FF8EFF98F86FFB8CFA5008A7" box="[1343,1431,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="43" pageNumber="44" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,
<date id="FF931027FF8EFF98FDC7FBE7FEC408FD" box="[151,259,1484,1509]" pageId="43" pageNumber="44" value="1999-11-02">
<collectingDate id="EFD7E9CFFF8EFF98FDC7FBE7FEC408FD" box="[151,259,1484,1509]" pageId="43" pageNumber="44" value="1999-11-02">2.xi.1999</collectingDate>
</date>
, mixed
<taxonomicName id="4C2D4D64FF8EFF98FC0CFBE7FE0308FC" box="[348,452,1484,1508]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FF8EFF98FC0CFBE7FE0308FC" box="[348,452,1484,1508]" italics="true" pageId="43" pageNumber="44">Polylepis</emphasis>
</taxonomicName>
litter,
<collectorName id="26D85331FF8EFF98FF5CFBE7FD7F08FD" box="[524,696,1484,1509]" pageId="43" pageNumber="44">R.S. Anderson.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9236E7FF8EFF98FD97FBDBFB630B35" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<materialsCitation id="3B453CBAFF8EFF98FD97FBDBFB670B35" ID-GBIF-Occurrence="3311767319" collectingDate="1999-11-01" collectionCode="DEBU" collectorName="R. S. Anderson" country="Ecuador" elevation="3300" location="Guandera Forest Reserve" pageId="43" pageNumber="44" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Carchi" typeStatus="paratype">
<emphasis id="B959EAF5FF8EFF98FD97FBDBFEF80B1F" bold="true" box="[199,319,1520,1543]" italics="true" pageId="43" pageNumber="44">
<typeStatus id="54968845FF8EFF98FD97FBDBFEFC0B1F" box="[199,315,1520,1543]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F33A7677FF8EFF98FC17FBC4FE140B10" box="[327,467,1519,1544]" name="Ecuador" pageId="43" pageNumber="44">ECUADOR</collectingCountry>
:
<collectingRegion id="49E9F805FF8EFF98FC8EFBC4FDEC0B10" box="[478,555,1519,1544]" country="Ecuador" name="Carchi" pageId="43" pageNumber="44">Carchi</collectingRegion>
, same label as
<typeStatus id="54968845FF8EFF98FF85FBC4FCF00B10" box="[725,823,1519,1544]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D2BFD6EFF8EFF98FE18FBC4FCA90B10" box="[840,878,1519,1544]" pageId="43" pageNumber="44" type="male">2♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8EFF98FE2AFBC4FC590B10" box="[890,926,1519,1544]" pageId="43" pageNumber="44" type="female">2♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8EFF98FEFCFBC4FC3A0B10" box="[940,1021,1519,1544]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="43" pageNumber="44">DEBU</collectionCode>
);
<location id="8EF2603CFF8EFF98F941FBC4FAF70B1F" LSID="urn:lsid:plazi:treatment:038487F1FF8FFF98FDC7F8ECFE310A30:8EF2603CFF8EFF98F941FBC4FAF70B1F" box="[1041,1328,1519,1544]" country="Ecuador" name="Guandera Forest Reserve" pageId="43" pageNumber="44" stateProvince="Carchi">Guandera Forest Reserve</location>
,
<locationDeviation id="9A202FC5FF8EFF98F86BFBC4FED80B35" location="San Gabriel" pageId="43" pageNumber="44">
<quantity id="4CD59B02FF8EFF98F86BFBC4FA440B10" box="[1339,1411,1519,1544]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="km" value="15.0">15 km</quantity>
E San Gabriel
</locationDeviation>
,
<quantity id="4CD59B02FF8EFF98FC7AF83FFE460B33" box="[298,385,1556,1581]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="44" unit="m" value="3300.0">
<elevation id="0000D1D4FF8EFF98FC7AF83FFE460B33" box="[298,385,1556,1581]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="44" unit="m" value="3300.0">3300 m</elevation>
</quantity>
,
<date id="FF931027FF8EFF98FCDCF83FFE3E0B35" box="[396,505,1556,1581]" pageId="43" pageNumber="44" value="1999-11-01">
<collectingDate id="EFD7E9CFFF8EFF98FCDCF83FFE3E0B35" box="[396,505,1556,1581]" pageId="43" pageNumber="44" value="1999-11-01">1.xi.1999</collectingDate>
</date>
, mixed riparian forest, leaf litter,
<collectorName id="26D85331FF8EFF98FE20F83FFBD30B35" box="[880,1044,1556,1581]" pageId="43" pageNumber="44">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8EFF98F976F83FFBFD0B35" box="[1062,1082,1556,1581]" pageId="43" pageNumber="44" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8EFF98F917F83FFB5F0B35" box="[1095,1176,1556,1581]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="43" pageNumber="44">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9236E7FF8EFF98FD97F81CFD0E0B6D" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97F81CFE280B48" bold="true" box="[199,495,1591,1616]" italics="true" pageId="43" pageNumber="44">Other material examined.</emphasis>
<materialsCitation id="3B453CBAFF8EFF98FCA7F81CFD020B6D" ID-GBIF-Occurrence="3311767403" collectingDate="1999-10-31" collectionCode="DEBU" collectorName="Paramo El Angel &amp; R. S. Anderson" country="Ecuador" elevation="3300" location="El Angel" municipality="Paramo El Angel" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
<collectingCountry id="F33A7677FF8EFF98FCA7F81CFD440B48" box="[503,643,1591,1616]" name="Ecuador" pageId="43" pageNumber="44">ECUADOR</collectingCountry>
:
<collectingRegion id="49E9F805FF8EFF98FFDDF81CFD1D0B48" box="[653,730,1591,1616]" country="Ecuador" name="Carchi" pageId="43" pageNumber="44">Carchi</collectingRegion>
,
<collectorName id="26D85331FF8EFF98FFB6F81CFC6D0B48" box="[742,938,1591,1616]" pageId="43" pageNumber="44">
<collectingMunicipality id="6BF6AC9DFF8EFF98FFB6F81CFC6D0B48" box="[742,938,1591,1616]" pageId="43" pageNumber="44">
Páramo
<location id="8EF2603CFF8EFF98FE13F81CFC6D0B48" LSID="urn:lsid:plazi:treatment:038487F1FF8FFF98FDC7F8ECFE310A30:8EF2603CFF8EFF98FE13F81CFC6D0B48" box="[835,938,1591,1616]" country="Ecuador" municipality="Paramo El Angel" name="El Angel" pageId="43" pageNumber="44" stateProvince="Carchi">El Ángel</location>
</collectingMunicipality>
</collectorName>
,
<locationDeviation id="9A202FC5FF8EFF98FEE6F81CFB710B48" box="[950,1206,1591,1616]" location="El Angel" pageId="43" pageNumber="44">
<quantity id="4CD59B02FF8EFF98FEE6F81CFBD50B48" box="[950,1042,1591,1616]" metricMagnitude="4" metricUnit="m" metricValue="1.88" pageId="43" pageNumber="44" unit="km" value="18.8">18.8 km</quantity>
NW El Ángel
</locationDeviation>
,
<quantity id="4CD59B02FF8EFF98F992F81CFADE0B57" box="[1218,1305,1591,1616]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="44" unit="m" value="3300.0">
<elevation id="0000D1D4FF8EFF98F992F81CFADE0B57" box="[1218,1305,1591,1616]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="44" unit="m" value="3300.0">3300 m</elevation>
</quantity>
,
<date id="FF931027FF8EFF98F875F81CFA500B48" box="[1317,1431,1591,1616]" pageId="43" pageNumber="44" value="1999-10-31">
<collectingDate id="EFD7E9CFFF8EFF98F875F81CFA500B48" box="[1317,1431,1591,1616]" pageId="43" pageNumber="44" value="1999-10-31">31.x.1999</collectingDate>
</date>
, mixed
<taxonomicName id="4C2D4D64FF8EFF98FDB4F877FE8B0B6C" box="[228,332,1628,1652]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAF5FF8EFF98FDB4F877FE8B0B6C" box="[228,332,1628,1652]" italics="true" pageId="43" pageNumber="44">Polylepis</emphasis>
</taxonomicName>
litter,
<collectorName id="26D85331FF8EFF98FCC5F877FDFE0B6D" box="[405,569,1628,1653]" pageId="43" pageNumber="44">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8EFF98FF1BF877FD980B6D" box="[587,607,1628,1653]" pageId="43" pageNumber="44" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8EFF98FF3CF877FD7A0B6D" box="[620,701,1628,1653]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="43" pageNumber="44">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8EFF98FD97F854FE310A30" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8B9236E7FF8EFF98FD97F854FE310A30" blockId="43.[151,1436,151,1832]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97F854FE8F0B80" bold="true" box="[199,328,1663,1688]" pageId="43" pageNumber="44">
<emphasis id="B959EAF5FF8EFF98FD97F854FE850B80" bold="true" box="[199,322,1663,1688]" italics="true" pageId="43" pageNumber="44">Comments</emphasis>
.
</emphasis>
<taxonomicName id="4C2D4D64FF8EFF98FC1FF8ABFDDF0B80" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[335,536,1664,1688]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="angela">
<emphasis id="B959EAF5FF8EFF98FC1FF8ABFDDF0B80" box="[335,536,1664,1688]" italics="true" pageId="43" pageNumber="44">Aptilotella angela</emphasis>
</taxonomicName>
and the closely related
<taxonomicName id="4C2D4D64FF8EFF98FE4EF8ABFC120B80" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[798,981,1664,1688]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pichinchensis">
<emphasis id="B959EAF5FF8EFF98FE4EF8ABFC120B80" box="[798,981,1664,1688]" italics="true" pageId="43" pageNumber="44">A. pichinchensis</emphasis>
</taxonomicName>
are somewhat rotund in appearance, and have banded legs and straight, narrow brown medial stripes on the frons. The distiphallus, as in other members of Clade 2, has curved dorsal sclerites and tripartite ventral flanking sclerites with a dorsally branched basal article.
<taxonomicName id="4C2D4D64FF8EFF98FDC7F8C7FEA10A1C" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,358,1772,1796]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="angela">
<emphasis id="B959EAF5FF8EFF98FDC7F8C7FEA10A1C" box="[151,358,1772,1796]" italics="true" pageId="43" pageNumber="44">Aptilotella angela</emphasis>
</taxonomicName>
is known from the northernmost slopes of the Ecuadorian Andes, and its distribution may possibly extend into
<collectingCountry id="F33A7677FF8EFF98FC2FF924FE360A30" box="[383,497,1807,1832]" name="Colombia" pageId="43" pageNumber="44">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>