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<document id="EE0CB57DA09E8429448266A2B9CF3BEE" ID-CLB-Dataset="6241" ID-DOI="10.11646/zootaxa.4973.1.1" ID-GBIF-Dataset="d0967b08-98d1-4ccd-b472-a678e0e57eda" ID-ISSN="1175-5326" ID-Zenodo-Dep="4771958" ID-ZooBank="42EB4BF2-A571-4894-9EEF-783649A27E4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1621409551899" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander &amp; Auffenberg, Kurt" docDate="2021" docId="038487E62F382423FF12B589FE5BFE74" docLanguage="en" docName="zootaxa.4973.1.1.pdf" docOrigin="Zootaxa 4973 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dicharax ataranensis" docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="FFBDFF9E2F20243AFF85B51EFF9BFFD8" masterDocTitle="Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)" masterLastPageNumber="61" masterPageNumber="1" pageNumber="25" updateTime="1698940034724" updateUser="ExternalLinkService">
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<mods:title id="860978EB8218F85DDC513B8770081167">Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)</mods:title>
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<mods:namePart id="870336F223B25A1B6AA2C8550E011971">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="19C6259B99847288F9AC8E3326459F9D">Hunyadi, András</mods:namePart>
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<mods:namePart id="83988F15099C3467190259DC71C1AEA0">Grego, Jozef</mods:namePart>
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<treatment id="038487E62F382423FF12B589FE5BFE74" ID-DOI="http://doi.org/10.5281/zenodo.4771770" ID-GBIF-Taxon="182640509" ID-Zenodo-Dep="4771770" LSID="urn:lsid:plazi:treatment:038487E62F382423FF12B589FE5BFE74" httpUri="http://treatment.plazi.org/id/038487E62F382423FF12B589FE5BFE74" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<paragraph id="8B9236F02F382422FF12B589FD5DFF69" blockId="24.[151,710,151,213]" box="[151,710,151,178]" pageId="24" pageNumber="25">
<heading id="D0DA819C2F382422FF12B589FD5DFF69" bold="true" box="[151,710,151,178]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="4">
<taxonomicName id="4C2D4D732F382422FF12B589FD5DFF69" authority="(Godwin-Austen, 1914)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[151,710,151,178]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ataranensis">
<emphasis id="B959EAE22F382422FF12B589FD5DFF69" bold="true" box="[151,710,151,178]" pageId="24" pageNumber="25">
<emphasis id="B959EAE22F382422FF12B589FE06FF69" bold="true" box="[151,413,151,178]" italics="true" pageId="24" pageNumber="25">Dicharax ataranensis</emphasis>
(Godwin-Austen, 1914)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C337657B2F382422FF12B5A5FF7CFF0D" box="[151,231,187,213]" pageId="24" pageNumber="25" type="description">
<paragraph id="8B9236F02F382422FF12B5A5FF7CFF0D" blockId="24.[151,710,151,213]" box="[151,231,187,213]" pageId="24" pageNumber="25">
<figureCitation id="13162A752F382422FF12B5A5FF7CFF0D" box="[151,231,187,213]" captionStart="FIGURE 18" captionStartId="24.[151,250,1859,1883]" captionTargetBox="[151,1436,545,1835]" captionTargetId="figure-122@24.[151,1436,545,1835]" captionTargetPageId="24" captionText="FIGURE 18. Dicharax ataranensis (Godwin-Austen, 1914). Myanmar, Kayin State, Mudon centre ESE ca. 51 km, Htimahto NE, NE point of Kwooprai Hill, 30 m a.s.l. 16°2.396N, 97°58.200E (locality code: 2018/47), leg. A. Hunyadi, K. Okubo &amp; J.U. Otani, 10 Oct. 2018, HA." figureDoi="http://doi.org/10.5281/zenodo.4771998" httpUri="https://zenodo.org/record/4771998/files/figure.png" pageId="24" pageNumber="25">Fig. 18</figureCitation>
</paragraph>
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<subSubSection id="C337657B2F382422FF12B41EFC92FE8D" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="8B9236F02F382422FF12B41EFCE3FEC0" blockId="24.[151,888,255,341]" box="[151,888,255,280]" pageId="24" pageNumber="25">
<taxonomicName id="4C2D4D732F382422FF12B41EFD1EFECF" authority="Godwin-Austen, 1914: 426" authorityName="Godwin-Austen" authorityPageNumber="426" authorityYear="1914" box="[151,645,255,280]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ataranensis">
<emphasis id="B959EAE22F382422FF12B41EFEF3FECF" box="[151,360,256,280]" italics="true" pageId="24" pageNumber="25">Alycaeus ataranensis</emphasis>
Godwin-Austen, 1914: 426
</taxonomicName>
, plate 148, figs 4, 4ab.
</paragraph>
<paragraph id="8B9236F02F382422FF12B400FD34FEEE" blockId="24.[151,888,255,341]" box="[151,687,286,310]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB3D11DE2F382422FF12B400FD34FEEE" box="[151,687,286,310]" pageId="24" pageNumber="25">
<taxonomicName id="4C2D4D732F382422FF12B400FE44FEEE" authorityName="Godwin-Austen" authorityYear="1914" box="[151,479,286,310]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ataranensis" subGenus="Dicharax">
<emphasis id="B959EAE22F382422FF12B400FF70FEEE" box="[151,235,286,310]" italics="true" pageId="24" pageNumber="25">Alycaeus</emphasis>
(
<emphasis id="B959EAE22F382422FF7FB400FEC1FEEE" box="[250,346,286,310]" italics="true" pageId="24" pageNumber="25">Dicharax</emphasis>
)
<emphasis id="B959EAE22F382422FEECB401FE44FEEE" box="[361,479,287,310]" italics="true" pageId="24" pageNumber="25">ataranensis</emphasis>
</taxonomicName>
<treatmentCitation id="0A8C10E12F382422FD80B400FD30FEEE" author="Gude, G. K." box="[517,683,286,310]" page="237" pageId="24" pageNumber="25" year="1921">
<bibRefCitation id="EFBC4B012F382422FD80B400FD30FEEE" author="Gude, G. K." box="[517,683,286,310]" pageId="24" pageNumber="25" refId="ref25109" refString="Gude, G. K. (1921) The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, 386 pp. https: // doi. org / 10.5962 / bhl. title. 12890" type="book" year="1921">Gude 1921: 237</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B9236F02F382422FF12B423FC92FE8D" blockId="24.[151,888,255,341]" box="[151,777,317,341]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB3D11DE2F382422FF12B423FC92FE8D" box="[151,777,317,341]" pageId="24" pageNumber="25">
<taxonomicName id="4C2D4D732F382422FF12B423FEE8FE8D" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[151,371,317,341]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ataranensis">
<emphasis id="B959EAE22F382422FF12B423FEE8FE8D" box="[151,371,317,341]" italics="true" pageId="24" pageNumber="25">Dicharax ataranensis</emphasis>
</taxonomicName>
<treatmentCitation id="0A8C10E12F382422FE1CB423FD2DFE8C" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J." box="[409,694,317,341]" httpUri="http://treatment.plazi.org/id/DE71AA9D2B5C5BA9A167FDBE28F59C6F" page="77" pageId="24" pageNumber="25" year="2020">
<bibRefCitation id="EFBC4B012F382422FE1CB423FD2DFE8C" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J." box="[409,694,317,341]" pageId="24" pageNumber="25" pagination="1 - 220" refId="ref26053" refString="Pall-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. &amp; Ablett, J. (2020) Genus-level revision of the Alycaeidae, with a catalog of the species (Gastropoda: Cyclophoroidea). ZooKeys, 981, 1 - 220. https: // doi. org / 10.3897 / zookeys. 981.53583" type="journal article" year="2020">
Páll-Gergely
<emphasis id="B959EAE22F382422FDA4B420FDD6FE8D" box="[545,589,317,341]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. 2020: 77
</bibRefCitation>
</treatmentCitation>
, fig. 18.
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</paragraph>
</subSubSection>
<subSubSection id="C337657B2F382423FF12B49DFE77FF68" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
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<emphasis id="B959EAE22F382422FF12B49DFE1FFE45" bold="true" box="[151,388,387,413]" pageId="24" pageNumber="25">Material examined.</emphasis>
<materialsCitation id="3B453CAD2F382422FE0BB49DFD0FFE19" ID-GBIF-Occurrence="3128117305" collectionCode="NZSI" collectorName="Dr. F. Stoliczka" country="Myanmar" location="Myanmar" pageId="24" pageNumber="25" specimenCount="2" typeStatus="holotype">
<emphasis id="B959EAE22F382422FE0BB49DFD9FFE45" bold="true" box="[398,516,387,413]" pageId="24" pageNumber="25">
<typeStatus id="549688522F382422FE0BB49DFE64FE45" box="[398,511,387,413]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F33A76602F382422FD8AB49AFDE5FE46" box="[527,638,388,414]" name="Myanmar" pageId="24" pageNumber="25">Myanmar</collectingCountry>
, Ataran, Burma, ex
<collectorName id="26D853262F382422FCEFB49AFBBAFE45" box="[874,1057,387,414]" pageId="24" pageNumber="25">Dr. F. Stoliczka</collectorName>
,
<collectionCode id="ED3CAE352F382422FBB5B49DFBF5FE45" box="[1072,1134,387,413]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="24" pageNumber="25">NZSI</collectionCode>
M.8073 (
<typeStatus id="549688522F382422FB67B49DFADDFE45" box="[1250,1350,387,413]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
[single specimen mentioned in original description])
</materialsCitation>
.
<emphasis id="B959EAE22F382422FD1BB4B9FCC0FE19" bold="true" box="[670,859,423,449]" pageId="24" pageNumber="25">Other material:</emphasis>
<materialsCitation id="3B453CAD2F382422FCE7B4B9FCAFFDD1" ID-GBIF-Occurrence="3128117376" collectingDate="2018-10-10" collectionCode="HA" collectorName="A. Hunyadi &amp; K. Okubo &amp; J. U. Otani" country="Myanmar" elevation="30" latitude="16.039934" location="Kwooprai Hill" longLatPrecision="1" longitude="97.97" municipality="Mudon" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Kayin">
1 shell,
<collectingCountry id="F33A76602F382422FC3EB4B6FBB1FE1A" box="[955,1066,424,450]" name="Myanmar" pageId="24" pageNumber="25">Myanmar</collectingCountry>
,
<collectingRegion id="49E9F8122F382422FBB0B4B9FB20FE19" box="[1077,1211,423,449]" country="Myanmar" name="Kayin" pageId="24" pageNumber="25">Kayin State</collectingRegion>
,
<collectingMunicipality id="6BF6AC8A2F382422FB42B4B9FA83FE19" box="[1223,1304,423,449]" pageId="24" pageNumber="25">Mudon</collectingMunicipality>
centre ESE ca.
<quantity id="4CD59B152F382422FF38B4D5FE9DFE3D" box="[189,262,459,486]" metricMagnitude="4" metricUnit="m" metricValue="5.1" pageId="24" pageNumber="25" unit="km" value="51.0">51 km</quantity>
,
<collectorName id="26D853262F382422FE8AB4D5FE3CFE3E" box="[271,423,459,486]" pageId="24" pageNumber="25">Htimahto NE</collectorName>
, NE point of
<location id="8EF2602B2F382422FDBCB4D5FD46FE3D" LSID="urn:lsid:plazi:treatment:038487E62F382423FF12B589FE5BFE74:8EF2602B2F382422FDBCB4D5FD46FE3D" box="[569,733,459,485]" country="Myanmar" latitude="16.039934" longLatPrecision="1" longitude="97.97" municipality="Mudon" name="Kwooprai Hill" pageId="24" pageNumber="25" stateProvince="Kayin">Kwooprai Hill</location>
,
<geoCoordinate id="EE1950372F382422FD62B4D5FCF6FE3D" box="[743,877,459,485]" degrees="16" direction="north" minutes="2.396" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="16.039934">16°2.396N</geoCoordinate>
,
<geoCoordinate id="EE1950372F382422FCFDB4D5FB92FE3E" box="[888,1033,459,486]" degrees="97" direction="east" minutes="58.200" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="97.97">97°58.200E</geoCoordinate>
,
<elevation id="0000D1C32F382422FB96B4D5FB1EFE3D" box="[1043,1157,459,486]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="25" unit="m" value="30.0">
<quantity id="4CD59B152F382422FB96B4D5FBD0FE3E" box="[1043,1099,459,486]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="25" unit="m" value="30.0">30 m</quantity>
a.s.l.
</elevation>
(locality code: 2018/47),
<date id="FF9310302F382422FF12B4F1FEB7FDD1" box="[151,300,495,521]" pageId="24" pageNumber="25" value="2018-10-10">
<collectingDate id="EFD7E9D82F382422FF12B4F1FEB7FDD1" box="[151,300,495,521]" pageId="24" pageNumber="25" value="2018-10-10">10 Oct. 2018</collectingDate>
</date>
,
<collectorName id="26D853262F382422FEB0B4F1FE21FDD1" box="[309,442,495,521]" pageId="24" pageNumber="25">A. Hunyadi</collectorName>
,
<collectorName id="26D853262F382422FE43B4EEFDAFFDD1" box="[454,564,495,522]" pageId="24" pageNumber="25">K. Okubo</collectorName>
&amp;
<collectorName id="26D853262F382422FDDDB4EEFD51FDD1" box="[600,714,495,522]" pageId="24" pageNumber="25">J.U. Otani</collectorName>
leg.,
<collectionCode id="ED3CAE352F382422FC8DB4F1FCAFFDD1" box="[776,820,495,521]" pageId="24" pageNumber="25">HA</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF5266782F382422FF12B25DFE5CF87C" ID-DOI="http://doi.org/10.5281/zenodo.4771998" ID-Zenodo-Dep="4771998" httpUri="https://zenodo.org/record/4771998/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1859,1883]" targetBox="[151,1436,545,1835]" targetPageId="24">
<paragraph id="8B9236F02F382422FF12B25DFE5CF87C" blockId="24.[151,1437,1859,1956]" pageId="24" pageNumber="25">
<emphasis id="B959EAE22F382422FF12B25DFEBAF883" bold="true" box="[151,289,1859,1883]" pageId="24" pageNumber="25">FIGURE 18.</emphasis>
<taxonomicName id="4C2D4D732F382422FEADB25DFC99F883" authority="(Godwin-Austen, 1914)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" box="[296,770,1859,1883]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ataranensis">
<emphasis id="B959EAE22F382422FEADB25DFD9EF883" box="[296,517,1859,1883]" italics="true" pageId="24" pageNumber="25">Dicharax ataranensis</emphasis>
(Godwin-Austen, 1914)
</taxonomicName>
. Myanmar, Kayin State, Mudon centre ESE ca. 51 km, Htimahto NE, NE point of Kwooprai Hill, 30 m a.s.l. 16°2.396N, 97°58.200E (locality code: 2018/47), leg. A. Hunyadi, K. Okubo &amp; J.U. Otani, 10 Oct. 2018, HA.
</paragraph>
</caption>
<paragraph id="8B9236F02F392423FF42B588FE77FF68" blockId="25.[151,1437,150,428]" box="[199,492,150,176]" pageId="25" pageNumber="26">
<emphasis id="B959EAE22F392423FF42B588FEFDFF68" bold="true" box="[199,358,150,176]" pageId="25" pageNumber="26">
<typeStatus id="549688522F392423FF42B588FE99FF68" box="[199,258,150,176]" pageId="25" pageNumber="26">Type</typeStatus>
locality.
</emphasis>
“1 Ataran”.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F392423FF42B5A4FCD0FE98" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8B9236F02F392423FF42B5A4FCD0FE98" blockId="25.[151,1437,150,428]" pageId="25" pageNumber="26">
<emphasis id="B959EAE22F392423FF42B5A4FCE0FF0C" bold="true" box="[199,891,186,212]" pageId="25" pageNumber="26">Diagnosis (emended based on a newly collected specimen).</emphasis>
Protoconch relatively low, glossy, without spiral striation; R1 glossy, widely-spaced ribs near the suture and inside the umbilicus, edge of R1 smooth; R2+R3 over a quarter whorl long, R2 ca. half as long as R3; R2 ribs typical for
<taxonomicName id="4C2D4D732F392423FC23B41CFB8BFEC4" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[934,1040,258,284]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F392423FC23B41CFB8BFEC4" box="[934,1040,258,284]" italics="true" pageId="25" pageNumber="26">Dicharax</emphasis>
</taxonomicName>
(curved towards the aperture); R3 smooth, with blunt, but large, conspicuous, elevated swelling.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F392423FF42B454FE5BFE74" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8B9236F02F392423FF42B454FE5BFE74" blockId="25.[151,1437,150,428]" pageId="25" pageNumber="26">
<emphasis id="B959EAE22F392423FF42B454FEA0FEBC" bold="true" box="[199,315,330,356]" pageId="25" pageNumber="26">Remarks.</emphasis>
Until recently the eroded
<typeStatus id="549688522F392423FDF4B454FD48FEBC" box="[625,723,330,356]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
was the only known specimen of the species making the assignment to
<taxonomicName id="4C2D4D732F392423FEA6B470FE16FE50" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[291,397,366,392]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F392423FEA6B470FE16FE50" box="[291,397,366,392]" italics="true" pageId="25" pageNumber="26">Dicharax</emphasis>
</taxonomicName>
questionable. The newly collected fresh shell allowed examination of shell sculpture which confirmed the assignment.
</paragraph>
</subSubSection>
</treatment>
</document>