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<document id="92C9A90AABC7CF2203F91556950D25A5" ID-DOI="10.1649/0010-065X-74.1.175" ID-GBIF-Dataset="28046c1b-405b-4b6e-b291-e88a770321e5" ID-ISSN="1938-4394" ID-Zenodo-Dep="3746907" ID-ZooBank="urn:lsid:zoobank.org:pub:F36FA2BB-C7F0-4A34-972A-C34572F9ED94" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586467962798" checkinUser="felipe" docAuthor="Powell, Gareth S." docDate="2020" docId="038487C2EE133244FD06FBF8FDF9FA41" docLanguage="en" docName="TheColeopteristsBulletin.74.1.175-180.pdf.imf" docOrigin="The Coleopterists Bulletin 74 (1)" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Carpophilus arizonicus Powell 2020, new species" docType="treatment" docUuid="2C03BEE8-0DE2-4127-B91E-15AF12CFCA06" docUuidSource="ZooBank" docVersion="5" lastPageNumber="180" masterDocId="FFBDFFBAEE163242FFDDFFC1FF85FFF6" masterDocTitle="Four New Species Of Carpophilus (Ecnomorphus) Motschulsky (Coleoptera: Nitidulidae: Carpophilinae) From The New World" masterLastPageNumber="180" masterPageNumber="175" pageNumber="179" updateTime="1698736860956" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="46B490918AC36E9BA37E39BEAD45D149">Four New Species Of Carpophilus (Ecnomorphus) Motschulsky (Coleoptera: Nitidulidae: Carpophilinae) From The New World</mods:title>
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<mods:name id="CEAE6D1374EF53B2B6E5F5AA9BC229A4" type="personal">
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<mods:namePart id="40E2B6D2570E3938D0CDD605FE4DE545">Powell, Gareth S.</mods:namePart>
<mods:affiliation id="A7E96E0696D85A29792CAF7295198357">Department of Biology Brigham Young University</mods:affiliation>
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<mods:title id="97B41ADE5301E5CFA2A4598B18FEFB64">The Coleopterists Bulletin</mods:title>
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<mods:date id="C4FEACAEE7F0E2236077D3C5E2DBD7D6">2020</mods:date>
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<treatment id="038487C2EE133244FD06FBF8FDF9FA41" ID-DOI="http://doi.org/10.5281/zenodo.3809094" ID-GBIF-Taxon="163544890" ID-Zenodo-Dep="3809094" LSID="urn:lsid:zoobank.org:act:2C03BEE8-0DE2-4127-B91E-15AF12CFCA06" httpUri="http://treatment.plazi.org/id/038487C2EE133244FD06FBF8FDF9FA41" lastPageId="6" lastPageNumber="180" pageId="5" pageNumber="179">
<subSubSection id="C337655FEE133247FD06FBF8FB15FBB9" box="[731,1168,1081,1103]" pageId="5" pageNumber="179" type="nomenclature">
<paragraph id="8B9236D4EE133247FD06FBF8FB15FBB9" blockId="5.[719,1180,1081,1183]" box="[731,1168,1081,1103]" pageId="5" pageNumber="179">
<heading id="D0DA81B8EE133247FD06FBF8FB15FBB9" bold="true" box="[731,1168,1081,1103]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="179" reason="2">
<emphasis id="B959EAC6EE133247FD06FBF8FB15FBB9" bold="true" box="[731,1168,1081,1103]" pageId="5" pageNumber="179">
<taxonomicName id="4C2D4D57EE133247FD06FBF8FB8BFBB9" authority="Powell" authorityName="Powell" authorityYear="2020" box="[731,1038,1081,1103]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="arizonicus" status="sp. nov.">
<emphasis id="B959EAC6EE133247FD06FBF8FC47FBB8" bold="true" box="[731,962,1081,1102]" italics="true" pageId="5" pageNumber="179">Carpophilus arizonicus</emphasis>
Powell
</taxonomicName>
,
<taxonomicNameLabel id="A26A57BDEE133247FBC6FBFBFB15FBB9" box="[1051,1168,1082,1103]" pageId="5" pageNumber="179" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C337655FEE133247FD22FB95FC66FB69" pageId="5" pageNumber="179" type="description">
<paragraph id="8B9236D4EE133247FD22FB95FC66FB69" blockId="5.[719,1180,1081,1183]" pageId="5" pageNumber="179">
Zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13162A51EE133247FC4DFB4BFC5EFB69" box="[912,987,1162,1183]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="5" pageNumber="179">Fig. 1D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE133247FD06FB75FC42FAB8" pageId="5" pageNumber="179" type="materials_examined">
<paragraph id="8B9236D4EE133247FD06FB75FC42FAB8" blockId="5.[707,1194,1204,1730]" pageId="5" pageNumber="179">
<emphasis id="B959EAC6EE133247FD06FB75FCF5FB3F" bold="true" box="[731,880,1204,1225]" pageId="5" pageNumber="179">Type Material.</emphasis>
<materialsCitation id="3B453C89EE133247FCA8FB75FC42FAB8" ID-GBIF-Occurrence="2597530338" collectingDate="2010-07" collectingMethod="intercept" collectionCode="ASU" collectorName="W. B. Warner" country="United States of America" county="Cochise Co." location="Cochise Co." pageId="5" pageNumber="179" specimenCount="1" typeStatus="holotype">
<emphasis id="B959EAC6EE133247FCA8FB75FB9FFB3F" bold="true" box="[885,1050,1204,1225]" pageId="5" pageNumber="179">
<typeStatus id="54968876EE133247FCA8FB75FC54FB3F" box="[885,977,1204,1225]" pageId="5" pageNumber="179" type="holotype">Holotype</typeStatus>
female
</emphasis>
labeled “
<collectingCountry id="F33A7644EE133247FBAEFB75FB23FB3F" box="[1139,1190,1204,1225]" name="United States of America" pageId="5" pageNumber="179">USA</collectingCountry>
: Ariz.:
<collectingCounty id="62F34E58EE133247FCDFFB0EFCFCFB12" box="[770,889,1231,1252]" pageId="5" pageNumber="179">Cochise Co.</collectingCounty>
| Huachuca Mtns.: Carr | Cyn. Below Carr house |
<geoCoordinate id="EE195013EE133247FC5AFB28FC65FB08" box="[903,992,1257,1278]" pageId="5" pageNumber="179">31.4447°</geoCoordinate>
,
<geoCoordinate id="EE195013EE133247FC31FB28FBD0FB08" box="[1004,1109,1257,1278]" pageId="5" pageNumber="179">-110.2864°</geoCoordinate>
|
<date id="FF931014EE133247FBB7FB28FB27FB08" box="[1130,1186,1257,1278]" pageId="5" pageNumber="179" value="2010-07">
<collectingDate id="EFD7E9FCEE133247FBB7FB28FB27FB08" box="[1130,1186,1257,1278]" pageId="5" pageNumber="179" value="2010-07">vii.10</collectingDate>
</date>
- 15.2018; V-flight |
<collectingMethod id="526C4EC3EE133247FCA8FAC5FC48FAEF" box="[885,973,1284,1305]" pageId="5" pageNumber="179">intercept</collectingMethod>
;
<collectorName id="26D85302EE133247FC06FAC5FBE6FAEF" box="[987,1123,1284,1305]" pageId="5" pageNumber="179">W. B. Warner</collectorName>
|| HO- LOTYPE: |
<taxonomicName id="4C2D4D57EE133247FCEAFADFFC2BFAC2" authorityName="Stephens" authorityYear="1829" box="[823,942,1310,1332]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAC6EE133247FCEAFADFFC2BFAC2" box="[823,942,1310,1332]" italics="true" pageId="5" pageNumber="179">Carpophilus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2D4D57EE133247FC1DFADFFBA6FAC5" authorityName="Powell" authorityYear="2020" box="[960,1059,1310,1331]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="arizonicus">
<emphasis id="B959EAC6EE133247FC1DFADFFBA6FAC5" box="[960,1059,1310,1331]" italics="true" pageId="5" pageNumber="179">arizonicus</emphasis>
</taxonomicName>
| Des.
<bibRefCitation id="EFBC4B25EE133247FBBAFADFFD76FAB8" author="Powell" firstAuthor="Powell" pageId="5" pageNumber="179" pagination="297 - 300" refId="ref4716" refString="Powell, G. S., and G. J. Martin. 2019. Type designations for sap beetles in the subfamily Carpophilinae Erichson (Coleoptera: Nitidulidae) housed in the Natural History Museum, London. Zootaxa 4590 (2): 297 - 300." type="journal article" year="2019">Powell 2019</bibRefCitation>
” (deposited in
<collectionCode id="ED3CAE11EE133247FC54FAF8FC39FAB8" box="[905,956,1337,1358]" country="USA" lsid="urn:lsid:biocol.org:col:15600" name="Arizona State University" pageId="5" pageNumber="179" type="Herbarium">ASU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE133247FD06FA95FBC8FA1B" pageId="5" pageNumber="179" type="diagnosis">
<paragraph id="8B9236D4EE133247FD06FA95FBC8FA1B" blockId="5.[707,1194,1204,1730]" pageId="5" pageNumber="179">
<emphasis id="B959EAC6EE133247FD06FA95FCC7FA9F" bold="true" box="[731,834,1364,1385]" pageId="5" pageNumber="179">Diagnosis.</emphasis>
<taxonomicName id="4C2D4D57EE133247FC8FFA92FBBEFA9F" authorityName="Powell" authorityYear="2020" box="[850,1083,1363,1385]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="arizonicus">
<emphasis id="B959EAC6EE133247FC8FFA92FBBEFA9F" box="[850,1083,1363,1385]" italics="true" pageId="5" pageNumber="179">Carpophilus arizonicus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C2D4D57EE133247FD1EFAACFC52FA75" authority="Murray" authorityName="Murray" authorityYear="1864" box="[707,983,1389,1411]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="ligneus">
<emphasis id="B959EAC6EE133247FD1EFAACFC02FA75" box="[707,903,1389,1411]" italics="true" pageId="5" pageNumber="179">Carpophilus ligneus</emphasis>
Murray
</taxonomicName>
(
<figureCitation id="13162A51EE133247FC3AFAAFFBB1FA75" box="[999,1076,1390,1411]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="5" pageNumber="179">Fig. 1H</figureCitation>
) but differs from the latter species in the shape of the scutellar shield (triangular in
<taxonomicName id="4C2D4D57EE133247FC52FA63FC71FA4E" authorityName="Murray" authorityYear="1864" box="[911,1012,1442,1464]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="ligneus">
<emphasis id="B959EAC6EE133247FC52FA63FC71FA4E" box="[911,1012,1442,1464]" italics="true" pageId="5" pageNumber="179">C. ligneus</emphasis>
</taxonomicName>
, pentagonal in
<taxonomicName id="4C2D4D57EE133247FB4EFA63FCA3FA25" authorityName="Powell" authorityYear="2020" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="arizonicus">
<emphasis id="B959EAC6EE133247FB4EFA63FCA3FA25" italics="true" pageId="5" pageNumber="179">C. arizonicus</emphasis>
</taxonomicName>
) and the more exaggerated slope of the elytral apices anteriorly in
<taxonomicName id="4C2D4D57EE133247FC1BFA19FBC2FA18" authorityName="Powell" authorityYear="2020" box="[966,1095,1496,1518]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="arizonicus">
<emphasis id="B959EAC6EE133247FC1BFA19FBC2FA18" box="[966,1095,1496,1518]" italics="true" pageId="5" pageNumber="179">C. arizonicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE133244FD06FA32FDFAFB0B" lastPageId="6" lastPageNumber="180" pageId="5" pageNumber="179" type="description">
<paragraph id="8B9236D4EE133244FD06FA32FDD7FB15" blockId="5.[707,1194,1204,1730]" lastBlockId="6.[151,639,194,1463]" lastPageId="6" lastPageNumber="180" pageId="5" pageNumber="179">
<emphasis id="B959EAC6EE133247FD06FA32FCD0F9FE" bold="true" box="[731,853,1523,1544]" pageId="5" pageNumber="179">Description.</emphasis>
Body weakly elongate, dorsoventrally flattened. Length
<quantity id="4CD59B31EE133247FC2EF9CCFBDCF9D4" box="[1011,1113,1549,1570]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="5" pageNumber="179" unit="mm" value="2.07">2.07 mm</quantity>
, width
<quantity id="4CD59B31EE133247FD1EF9E9FCA4F9CB" box="[707,801,1576,1597]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="5" pageNumber="179" unit="mm" value="0.81">0.81 mm</quantity>
. Color light brown, elytral apices and abdomen darker (
<figureCitation id="13162A51EE133247FCA8F982FC43F9AE" box="[885,966,1603,1624]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[154,1169,893,1645]" captionTargetId="figure@2.[152,1171,891,1647]" captionTargetPageId="2" captionText="Fig. 1. Carpophilus (Ecnomorphus) species, dorsal habitus. A) C. alexandrae, B) C. mellarius, C) C. taylori, D) C. arizonicus, E) C. veraepacis (lectotype in NHM), F) C. marginatus (holotype in MNHUB), G) C. discoideus (image from MCZ type database), H) C. ligneus (lectotype in NHM)." figureDoi="http://doi.org/10.5281/zenodo.3746909" httpUri="https://zenodo.org/record/3746909/files/figure.png" pageId="5" pageNumber="179">Fig. 1D</figureCitation>
). Ventral surface light brown. Dorsal sculpturing moderately shiny; dorsum with fine, golden pubescence.
<emphasis id="B959EAC6EE133247FBD1F9B9FBCCF97B" bold="true" box="[1036,1097,1656,1677]" pageId="5" pageNumber="179">Head:</emphasis>
Narrower than pronotum, finely punctured. Punctures uniform in size, becoming sparser at midline. Frontoclypeal region truncate, labrum weakly emarginate, mandibles toothed, light brown. Palpi fusiform, reaching about 2/3 length of mandibles. Eyes small, finely faceted, interocular distance
<quantity id="4CD59B31EE103244FE63FED0FD98FED0" box="[446,541,273,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="180" unit="mm" value="0.37">0.37 mm</quantity>
. Antenna 1.5× longer than head; antennomere 1 robust and curved, segment 2 as long as basal antennomere, antennomeres 2 and 3 subequal in length, antennomeres 28 expanding apically, each apical expansion conspicuously lighter in coloration, antennomeres 48 each about half length of antennomere 2, antennomeres 911 forming strong, compact club that is slightly longer than wide.
<emphasis id="B959EAC6EE103244FF4AFDC1FE80FDE3" bold="true" box="[151,261,512,533]" pageId="6" pageNumber="180">Pronotum:</emphasis>
1.5 times as wide as long, sides weakly rounded, anterior angles almost right angles, posterior angles clearly obtuse. Posterior margin sinuate, with well-developed marginal line. Disc flattened, moderately shiny, evenly punctured.
<emphasis id="B959EAC6EE103244FF4AFD44FEBBFD6C" bold="true" box="[151,318,645,666]" pageId="6" pageNumber="180">Scutellar shield:</emphasis>
Feebly pentagonal, reaching almost right angle posteriorly. Anteriorly finely granulated, gradually becoming glabrous.
<emphasis id="B959EAC6EE103244FDE8FD7BFDF8FD39" bold="true" box="[565,637,698,719]" pageId="6" pageNumber="180">Elytra:</emphasis>
Width subequal to pronotal width, as wide as long. Humeri weakly pronounced. Anterior angles almost right angles, sides parallel, posterior angles slightly acute, elytral apices retreating towards midline.
<emphasis id="B959EAC6EE103244FF4AFCFEFE85FCA2" bold="true" box="[151,256,831,852]" pageId="6" pageNumber="180">Abdomen:</emphasis>
Two tergites dorsally visible, goldishbrown setae denser than on rest of dorsum. Pygidium rounded with distinct, evenly spaced punctures.
<emphasis id="B959EAC6EE103244FF4AFC4EFF5AFC52" bold="true" box="[151,223,911,932]" pageId="6" pageNumber="180">Venter:</emphasis>
Overall paler, submentum narrow and transverse, antennal grooves well-developed. Prosternum punctate, punctures with fine, golden setae, process slightly convex in lateral profile, apically rounded. Mesothoracic ventrite finely punctured. Metathoracic ventrite setose, finely punctate throughout. Abdominal ventrite 1 almost as long as metathoracic ventrite, abdominal ventrites 23 small, finely, sparsely punctate, ventrites 4 and 5 large, granulated, more densely pubescent.
<emphasis id="B959EAC6EE103244FD9BFBBFFDF8FB65" bold="true" box="[582,637,1150,1171]" pageId="6" pageNumber="180">Legs:</emphasis>
Somewhat short, femora robust, covered with fine golden pubescence. Tibiae expanded apically, with 35 apical spines.
<emphasis id="B959EAC6EE103244FE91FB0FFE60FB15" bold="true" box="[332,485,1230,1251]" pageId="6" pageNumber="180">Male genitalia:</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9236D4EE103244FF72FB29FDFAFB0B" blockId="6.[151,639,194,1463]" box="[175,639,1256,1277]" pageId="6" pageNumber="180">
<emphasis id="B959EAC6EE103244FF72FB29FE90FB0B" bold="true" box="[175,277,1256,1277]" pageId="6" pageNumber="180">Variation.</emphasis>
Only known from a single specimen.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE103244FF72FAC2FE8DFABB" pageId="6" pageNumber="180" type="distribution">
<paragraph id="8B9236D4EE103244FF72FAC2FE8DFABB" blockId="6.[151,639,194,1463]" pageId="6" pageNumber="180">
<emphasis id="B959EAC6EE103244FF72FAC2FE37FAEE" bold="true" box="[175,434,1283,1304]" pageId="6" pageNumber="180">Geographic Distribution.</emphasis>
The species is currently known from a single specimen from
<collectingRegion id="49E9F836EE103244FD8AFADCFF43FABB" country="United States of America" name="Arizona" pageId="6" pageNumber="180">Arizona</collectingRegion>
,
<collectingCountry id="F33A7644EE103244FF0FFAF9FE80FABB" box="[210,261,1336,1357]" name="United States of America" pageId="6" pageNumber="180">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE103244FF72FA93FEF3FA74" pageId="6" pageNumber="180" type="biology_ecology">
<paragraph id="8B9236D4EE103244FF72FA93FEF3FA74" blockId="6.[151,639,194,1463]" pageId="6" pageNumber="180">
<emphasis id="B959EAC6EE103244FF72FA93FE84FA91" bold="true" box="[175,257,1362,1383]" pageId="6" pageNumber="180">Biology.</emphasis>
The specimen studied was captured using an intercept trap.
</paragraph>
</subSubSection>
<subSubSection id="C337655FEE103244FF72FA49FDF9FA41" pageId="6" pageNumber="180" type="etymology">
<paragraph id="8B9236D4EE103244FF72FA49FDF9FA41" blockId="6.[151,639,194,1463]" pageId="6" pageNumber="180">
<emphasis id="B959EAC6EE103244FF72FA49FEA7FA6B" bold="true" box="[175,290,1416,1437]" pageId="6" pageNumber="180">Etymology.</emphasis>
The specific epithet is in reference to the
<typeStatus id="54968876EE103244FF61FA63FF60FA41" box="[188,229,1442,1463]" pageId="6" pageNumber="180">type</typeStatus>
locality within the
<collectingCountry id="F33A7644EE103244FE79FA63FE51FA41" box="[420,468,1442,1463]" name="United States of America" pageId="6" pageNumber="180">USA</collectingCountry>
state of
<collectingRegion id="49E9F836EE103244FDF4FA63FDFDFA41" box="[553,632,1442,1463]" country="United States of America" name="Arizona" pageId="6" pageNumber="180">Arizona</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>