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<document id="64F5AC1789FA77E9AAE356522BABF787" ID-DOI="10.11646/phytotaxa.218.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13635877" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725333068986" checkinUser="felipe" docAuthor="Karremans, Adam P., Bogarín, Diego, Pupulin, Franco, Luer, Carlyle A. &amp; Gravendeel, Barbara" docDate="2015" docId="038487AAE475FFBBFF444585FC19F98A" docLanguage="en" docName="phytotaxa.218.2.1.pdf" docOrigin="Phytotaxa 218 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.218.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Specklinia glandulosa Pridgeon &amp; M. W. Chase. A. 2001" docType="treatment" docVersion="2" lastPageNumber="116" masterDocId="FFBDFFD2E47AFFB4FFCC451BFFEDFFD3" masterDocTitle="The glandulous Specklinia: morphological convergence versus phylogenetic divergence" masterLastPageNumber="127" masterPageNumber="101" pageNumber="116" updateTime="1725334363440" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DE2299AD359620C91CDCEC85C7BE24C5">The glandulous Specklinia: morphological convergence versus phylogenetic divergence</mods:title>
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<mods:namePart id="548155C6C35E845F622F4303C0916545">Karremans, Adam P.</mods:namePart>
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<mods:affiliation id="9559F6FEB2EE57F38F30A9A5B7EBD403">Herbario UCH, Universidad Autónoma de Chiriquí, 0427, David, Chiriquí, Panama.</mods:affiliation>
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<mods:namePart id="AAEC33453C917DC36FE45BE5DF575672">Pupulin, Franco</mods:namePart>
<mods:affiliation id="5363BECE8DD98E747BB85CEE52EF924B">Harvard University Herbaria, Cambridge, MA, U. S. A.; Marie Selby Botanical Gardens, Sarasota, FL, U. S. A.</mods:affiliation>
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<mods:namePart id="9B2F14ACA552C08C8D17BBFC8F0D55E8">Luer, Carlyle A.</mods:namePart>
<mods:affiliation id="38A56B26C53652F4F41DA686A46EC284">Curator emeritus: Missouri Botanical Garden, St. Louis, Missouri, U. S. A.</mods:affiliation>
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<mods:namePart id="2DD5C3F236CB75FE20BA3A6F5817DEEA">Gravendeel, Barbara</mods:namePart>
<mods:affiliation id="D58E809933F0ED6931E3685B125389A1">Naturalis Biodiversity Center-Leiden University, The Netherlands.</mods:affiliation>
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<treatmentCitation id="0A8C10ADE475FFBBFF444585FCA8FF6A" author="Pridgeon, A. M. &amp; Chase, M. W." box="[136,837,158,185]" page=")" pageId="15" pageNumber="116" year="2001">
<taxonomicName id="4C2D4D3FE475FFBBFF444585FCA8FF6A" ID-CoL="4YGWR" authority="(Ames) Pridgeon &amp; Chase (2001: 257)" authorityName="Pridgeon &amp; M. W. Chase. A." authorityPageNumber="257" authorityYear="2001" baseAuthorityName="Ames" baseAuthorityYear="1923" box="[136,837,158,185]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="glandulosa">
<emphasis id="B959EAAEE475FFBBFF444585FE65FF6A" bold="true" box="[136,392,158,185]" italics="true" pageId="15" pageNumber="116">Specklinia glandulosa</emphasis>
(Ames)
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.
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Basionym:
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<emphasis id="B959EAAEE475FFBBFEC445DFFDF0FF0E" box="[264,541,196,221]" italics="true" pageId="15" pageNumber="116">Pleurothallis glandulosa</emphasis>
<bibRefCitation id="EFBC4B4DE475FFBBFDEE45DFFD09FF0E" author="Ames, O." box="[546,740,196,221]" pageId="15" pageNumber="116" pagination="60 - 65" refId="ref13049" refString="Ames, O. (1923) New or noteworthy orchids. Schedulae Orchidianae 6: 60 - 65." type="journal article" year="1923">Ames (1923: 60)</bibRefCitation>
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.
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:—
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.
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range, sea level,
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.
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.
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306
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(
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,
<collectionCode id="ED3CAE79E475FFBBFCB84415FC53FEF0" box="[884,958,270,291]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15408" lsid="urn:lsid:biocol.org:col:15408" name="Harvard University - Oakes Ames Orchid Herbarium" pageId="15" pageNumber="116" type="Herbarium">AMES</collectionCode>
!;
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,
<collectionCode id="ED3CAE79E475FFBBFBE94415FB82FEF0" box="[1061,1135,270,291]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15408" lsid="urn:lsid:biocol.org:col:15408" name="Harvard University - Oakes Ames Orchid Herbarium" pageId="15" pageNumber="116" type="Herbarium">AMES</collectionCode>
!,
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!;
<figureCitation id="13162A39E475FFBBFB7B4415FB1CFEF0" box="[1207,1265,270,291]" captionStart="FIGURE 9" captionStartId="13.[136,229,2028,2049]" captionTargetBox="[301,1304,343,1950]" captionTargetId="figure-76@13.[238,1349,317,2002]" captionTargetPageId="13" captionText="FIGURE 9. Type illustration of Specklinia glandulosa (Ames) Pridgeon &amp; M.W.Chase, published by Ames (1923). Reproduced with the kind permission of the Orchid Herbarium of Oakes Ames, the Harvard University Herbaria." figureDoi="http://doi.org/10.5281/zenodo.13635901" httpUri="https://zenodo.org/record/13635901/files/figure.png" pageId="15" pageNumber="116">Fig. 9</figureCitation>
)
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.
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Epiphytic, caespitose, ascending, erect herb to
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tall (excluding the inflorescence). Roots fibrous, flexuous, glabrous. Stem abbreviated, terete, to
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long, completely concealed by papyraceous, subancipitous, acute sheaths. Leaf narrowly linear, up to 1821 × 1.52.0 mm, gradually tapering toward the base into a deeply conduplicate petiole, subcoriaceous. Inflorescence borne laterally from the apex of the stem, without an annulus, successively single flowered, up to
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long, glandular; peduncle terete, with 1 distant, glandular, terete bract. Floral bracts infundibuliform, glandular, broadly ovate, acute. Pedicel terete, glandular,
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long, persistent. Ovary terete. Flowers 1 (probably up to a few with time) per inflorescence, Pompeian red. Sepals fleshy, densely glandulose in the outer surface; dorsal sepal narrowly lanceolate, 3-veined, acute, 8.0 ×
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; lateral sepals linear-lanceolate, subfalcate, 3-veined, 8.0 × 1.0 mm, connate to below the middle, the base saccate, the apex acute, the veins strongly carinate abaxially. Petals small, ligulate-falcate, acute, 3 ×
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, 2-veined. Lip small, longitudinally arched-convex in natural position, thinly articulate with the column foot by a hyaline claw, subpandurate from a cordate-sagittate base when expanded, obtuse, 3.5 ×
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, provided with a pair of acute, triangular lateral lobes from just below to just above the middle, margin dentate-erose, especially apically. Column arched, terete and slender at the base,
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long without the foot, provided with membranous wings, margins irregular, the apex prolonged into a deeply cucullate, dentate clinandrium; column foot, stout, fleshy. *
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: Description adapted from the protologue.
</paragraph>
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<emphasis id="B959EAAEE475FFBBFF71466BFE55FC5A" bold="true" box="[189,440,880,905]" pageId="15" pageNumber="116">Additional material:</emphasis>
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.
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:
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fork (known locally as “Quebrada Mona”) of
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,
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,
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<geoCoordinate id="EE19507BE475FFBBFD92468FFD2FFC7E" box="[606,706,916,941]" degrees="84" direction="west" minutes="36" orientation="longitude" pageId="15" pageNumber="116" precision="925" value="-84.6">84º36W</geoCoordinate>
, elev.
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, epiphytic on branch of large fallen tree in primary forest,
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<collectingDate id="EFD7E994E475FFBBFF1B46A3FE60FC02" box="[215,397,952,977]" pageId="15" pageNumber="116" value="1985-08-31">31 August 1985</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFE5546A3FC85FC02" box="[409,872,952,977]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFE5546A3FE15FC02" box="[409,504,952,977]" pageId="15" pageNumber="116">Grayum</collectorName>
,
<collectorName id="26D8536AE475FFBBFDC846A3FDBAFC02" box="[516,599,952,977]" pageId="15" pageNumber="116">Warner</collectorName>
,
<collectorName id="26D8536AE475FFBBFDAE46A3FD5BFC02" box="[610,694,952,977]" pageId="15" pageNumber="116">Sleeper</collectorName>
&amp;
<collectorName id="26D8536AE475FFBBFD1046A3FCC5FC02" box="[732,808,952,977]" pageId="15" pageNumber="116">Phelps</collectorName>
5939
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFCB546A3FC5DFC02" box="[889,944,952,977]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="116" type="Herbarium">MO</collectionCode>
!).
<collectorName id="26D8536AE475FFBBFC0B46A3FBCFFC02" box="[967,1058,952,977]" pageId="15" pageNumber="116">Without</collectorName>
specific collection data, cultivated by
<collectorName id="26D8536AE475FFBBFF6746C7FE95FC26" box="[171,376,988,1013]" pageId="15" pageNumber="116">Gerson Villalobos</collectorName>
, flowered
<date id="FF93107CE475FFBBFE2046C7FD4DFC26" box="[492,672,988,1013]" pageId="15" pageNumber="116" value="2014-08-29">
<collectingDate id="EFD7E994E475FFBBFE2046C7FD4DFC26" box="[492,672,988,1013]" pageId="15" pageNumber="116" value="2014-08-29">29 August 2014</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFD6646C7FC8BFC26" box="[682,870,988,1013]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFD6646C7FCCAFC26" box="[682,807,988,1013]" pageId="15" pageNumber="116">Karremans</collectorName>
6306
</emphasis>
(JBL-spirit!;
<figureCitation id="13162A39E475FFBBFC3346C7FBA0FC26" box="[1023,1101,988,1013]" captionStart="FIGURE 1" captionStartId="3.[136,229,1989,2010]" captionTargetBox="[204,1384,190,1964]" captionTargetId="figure-14@3.[204,1384,190,1964]" captionTargetPageId="3" captionText="FIGURE 1. The glandulous Specklinia species. A. S. alajuelensis 1 (Karremans 3265). B. S. alajuelensis 2 (Bogarín 2895). C. S. chontalensis (Pupulin 6543). D. S. gersonii (Karremans 6025). E. S. glandulosa (Karremans 6306). F. S. vittariifolia (Karremans 2945). Photographs by A.P. Karremans." figureDoi="http://doi.org/10.5281/zenodo.13635879" httpUri="https://zenodo.org/record/13635879/files/figure.png" pageId="15" pageNumber="116">Fig. 1e</figureCitation>
,
<figureCitation id="13162A39E475FFBBFB9646C7FB99FC26" box="[1114,1140,988,1013]" captionStart="FIGURE 2" captionStartId="4.[136,229,1062,1083]" captionTargetBox="[438,1149,281,1037]" captionTargetId="figure-54@4.[437,1149,281,1037]" captionTargetPageId="4" captionText="FIGURE 2. Comparison of the habits and flowers of species of the glandulous species of Specklinia. A. S. alajuelensis 1 (Karremans 3265). B. S. gersonii (Karremans 6025). C. S. glandulosa (Karremans 6306). D. S. vittariifolia (Karremans 2945). Scale bar = 1 cm. Photographs by A.P. Karremans." figureDoi="http://doi.org/10.5281/zenodo.13635881" httpUri="https://zenodo.org/record/13635881/files/figure.png" pageId="15" pageNumber="116">2c</figureCitation>
&amp;
<figureCitation id="13162A39E475FFBBFB5B46C7FB5BFC26" box="[1175,1206,988,1013]" captionStart="FIGURE 10" captionStartId="14.[136,229,1994,2015]" captionTargetBox="[163,1436,220,1951]" captionTargetId="figure-14@14.[151,1436,187,1966]" captionTargetPageId="14" captionText="FIGURE 10. Specklinia glandulosa (Ames) Pridgeon &amp; M.W.Chase.A. habit. B. flower. C. dissected perianth. D. column and lip, lateral view. E. lip. F. anther and pollinaria. Drawn by A.P. Karremans &amp; L. Oses from Karremans 6306 (JBL-spirit)." figureDoi="http://doi.org/10.5281/zenodo.13635905" httpUri="https://zenodo.org/record/13635905/files/figure.png" pageId="15" pageNumber="116">10</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B453CE1E475FFBBFB0646C7FE8FFB56" collectingDate="1980-04-29" collectingDateMax="1994-09-29" collectingDateMin="1980-04-29" collectionCode="PMA, SEL, W, N" collectorName="de Chepo &amp; Galdames &amp; Montenegro &amp; Chung &amp; Valdez &amp; Chepo &amp; Minas &amp; Walter &amp; Luer &amp; Cerro Alto Higo" country="Panama" county="Distrito de Las Minas" elevation="900" latitude="7.7166667" location="Las Minas" longLatPrecision="1303" longitude="80.833336" municipality="Ciclo de Chepo" pageId="15" pageNumber="116" specimenCount="1" stateProvince="Herrera">
<collectingCountry id="F33A762CE475FFBBFB0646C7FAA8FC26" box="[1226,1349,988,1013]" name="Panama" pageId="15" pageNumber="116">PANAMA</collectingCountry>
.
<collectingRegion id="49E9F85EE475FFBBFA8246C7FA45FC26" box="[1358,1448,988,1013]" country="Panama" name="Herrera" pageId="15" pageNumber="116">Herrera</collectingRegion>
:
<collectingCounty id="62F34E30E475FFBBFF44411BFE96FBCA" box="[136,379,1024,1049]" pageId="15" pageNumber="116">Distrito de Las Minas</collectingCounty>
, alrededor del primer
<collectingMunicipality id="6BF6ACC6E475FFBBFDA1411BFCF1FBCA" box="[621,796,1024,1049]" pageId="15" pageNumber="116">
Ciclo
<collectorName id="26D8536AE475FFBBFD63411BFCF1FBCA" box="[687,796,1024,1049]" pageId="15" pageNumber="116">de Chepo</collectorName>
</collectingMunicipality>
, ca.
<quantity id="4CD59B59E475FFBBFC87411BFC7EFBCA" box="[843,915,1024,1049]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="116" unit="m" value="900.0">
<elevation id="0000D18FE475FFBBFC87411BFC7EFBCA" box="[843,915,1024,1049]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="116" unit="m" value="900.0">900 m</elevation>
</quantity>
,
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<geoCoordinate id="EE19507BE475FFBBFC38411BFBB9FBCA" box="[1012,1108,1024,1049]" degrees="80" direction="east" minutes="50" orientation="longitude" pageId="15" pageNumber="116" precision="925" value="80.833336">80º50O</geoCoordinate>
, bosque nuboso,
<date id="FF93107CE475FFBBFADE411BFF2EFBEE" pageId="15" pageNumber="116" value="1994-09-29">
<collectingDate id="EFD7E994E475FFBBFADE411BFF2EFBEE" pageId="15" pageNumber="116" value="1994-09-29">29 September 1994</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFF1D413FFD15FBEE" box="[209,760,1060,1085]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFF1D413FFEA8FBEE" box="[209,325,1060,1085]" pageId="15" pageNumber="116">Galdames</collectorName>
,
<collectorName id="26D8536AE475FFBBFE98413FFE32FBEE" box="[340,479,1060,1085]" pageId="15" pageNumber="116">Montenegro</collectorName>
,
<collectorName id="26D8536AE475FFBBFE21413FFDD5FBEE" box="[493,568,1060,1085]" pageId="15" pageNumber="116">Chung</collectorName>
&amp;
<collectorName id="26D8536AE475FFBBFDA1413FFD5BFBEE" box="[621,694,1060,1085]" pageId="15" pageNumber="116">Valdez</collectorName>
1758
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFCC7413FFCBFFBEE" box="[779,850,1060,1085]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="15" pageNumber="116">PMA</collectionCode>
!).
<collectorName id="26D8536AE475FFBBFCA0413FFC58FBEE" box="[876,949,1060,1085]" pageId="15" pageNumber="116">Chepo</collectorName>
de las
<collectorName id="26D8536AE475FFBBFBC2413FFBBAFBEE" box="[1038,1111,1060,1085]" pageId="15" pageNumber="116">Minas</collectorName>
,
<emphasis id="B959EAAEE475FFBBFBA9413FFAFAFBEE" box="[1125,1303,1060,1085]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFBA9413FFB42FBEE" box="[1125,1199,1060,1085]" pageId="15" pageNumber="116">Walter</collectorName>
78-1518
</emphasis>
, flowered in cultivation at
<collectionCode id="ED3CAE79E475FFBBFEE94153FEB7FBB2" box="[293,346,1096,1121]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15632" lsid="urn:lsid:biocol.org:col:15632" name="Marie Selby Botanical Gardens" pageId="15" pageNumber="116" type="Herbarium">SEL</collectionCode>
,
<date id="FF93107CE475FFBBFEAB4153FDE4FBB2" box="[359,521,1096,1121]" pageId="15" pageNumber="116" value="1980-04-29">
<collectingDate id="EFD7E994E475FFBBFEAB4153FDE4FBB2" box="[359,521,1096,1121]" pageId="15" pageNumber="116" value="1980-04-29">29 April 1980</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFDD94153FD66FBB2" box="[533,651,1096,1121]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFDD94153FDA7FBB2" box="[533,586,1096,1121]" pageId="15" pageNumber="116">Luer</collectorName>
5237
</emphasis>
[
<collectionCode id="ED3CAE79E475FFBBFD574153FD3AFBB2" box="[667,727,1096,1121]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15632" lsid="urn:lsid:biocol.org:col:15632" name="Marie Selby Botanical Gardens" pageId="15" pageNumber="116" type="Herbarium">SEL</collectionCode>
!; illustration by
<bibRefCitation id="EFBC4B4DE475FFBBFC434153FBCFFBB2" author="Luer, C. A." box="[911,1058,1096,1121]" pageId="15" pageNumber="116" pagination="1 - 274" refId="ref13635" refString="Luer, C. A. (2006) A Reconsideration of Masdevallia. Systematics of Specklinia and vegetatively similar taxa (Orchidaceae). Icones Pleurothallidinarum XXVIII. Monographs in Systematic Botany from the Missouri Botanical Garden 105: 1 - 274." type="journal article" year="2006">Luer (2006)</bibRefCitation>
!].
<quantity id="4CD59B59E475FFBBFBF54153FB6FFBB2" box="[1081,1154,1096,1121]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="116" unit="km" value="18.0">18 km</quantity>
<collectionCode id="ED3CAE79E475FFBBFB464153FB49FBB2" box="[1162,1188,1096,1121]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="15" pageNumber="116" type="Herbarium">W</collectionCode>
of
<location id="8EF26067E475FFBBFB014153FAAAFBB2" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFB014153FAAAFBB2" box="[1229,1351,1096,1121]" country="Panama" county="Distrito de Las Minas" latitude="7.7166667" longLatPrecision="1303" longitude="80.833336" municipality="Ciclo de Chepo" name="Las Minas" pageId="15" pageNumber="116" stateProvince="Herrera">Las Minas</location>
,
<collectionCode id="ED3CAE79E475FFBBFA9F4153FA8AFBB2" box="[1363,1383,1096,1121]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="15" pageNumber="116" type="Herbarium">N</collectionCode>
slope of
<collectorName id="26D8536AE475FFBBFF6B4177FE8FFB56" box="[167,354,1132,1157]" pageId="15" pageNumber="116">Cerro Alto Higo</collectorName>
</materialsCitation>
;
<quantity id="4CD59B59E475FFBBFEA24177FDF6FB56" box="[366,539,1132,1157]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" metricValueMax="9.144" metricValueMin="7.3152" pageId="15" pageNumber="116" unit="ft" value="2700.0" valueMax="3000.0" valueMin="2400.0">
<geoCoordinate id="EE19507BE475FFBBFEA24177FE40FB56" box="[366,429,1132,1157]" degrees="2,400" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="55" value="2.4">2,400</geoCoordinate>
<geoCoordinate id="EE19507BE475FFBBFE614177FE16FB56" box="[429,507,1132,1157]" degrees="3,000" direction="west" orientation="longitude" pageId="15" pageNumber="116" precision="55" value="-3.0">3,000</geoCoordinate>
ft.
</quantity>
(known locally as el Montoso);
<materialsCitation id="3B453CE1E475FFBBFC474177FA4CFB22" collectingDate="1960-08-24" collectingDateMax="2005-07-24" collectingDateMin="1960-08-24" collectionCode="MO, PMA" collectorName="Ibanez &amp; Nunez &amp; Lewis &amp; Baker &amp; MacBryde &amp; Oliver" country="Panama" county="Hammel" elevation="792" location="Parque Nacional Coiba" municipality="Coiba" pageId="15" pageNumber="116" specimenCount="1" stateProvince="Veraguas">
<date id="FF93107CE475FFBBFC474177FBDEFB56" box="[907,1075,1132,1157]" pageId="15" pageNumber="116" value="1978-08-06">
<collectingDate id="EFD7E994E475FFBBFC474177FBDEFB56" box="[907,1075,1132,1157]" pageId="15" pageNumber="116" value="1978-08-06">6 August 1978</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFBF34177FB33FB56" box="[1087,1246,1132,1157]" italics="true" pageId="15" pageNumber="116">
<collectingCounty id="62F34E30E475FFBBFBF34177FB73FB56" box="[1087,1182,1132,1157]" pageId="15" pageNumber="116">Hammel</collectingCounty>
4289
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFB224177FAC8FB56" box="[1262,1317,1132,1157]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="116" type="Herbarium">MO</collectionCode>
!).
<collectingRegion id="49E9F85EE475FFBBFAF14177FA45FB56" box="[1341,1448,1132,1157]" country="Panama" name="Veraguas" pageId="15" pageNumber="116">Veraguas</collectingRegion>
:
<collectingMunicipality id="6BF6ACC6E475FFBBFF44418BFF23FB7A" box="[136,206,1168,1193]" pageId="15" pageNumber="116">Coiba</collectingMunicipality>
,
<location id="8EF26067E475FFBBFF14418BFE30FB7A" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFF14418BFE30FB7A" box="[216,477,1168,1193]" country="Panama" county="Hammel" municipality="Coiba" name="Parque Nacional Coiba" pageId="15" pageNumber="116" stateProvince="Veraguas">Parque Nacional Coiba</location>
, afluente del
<location id="8EF26067E475FFBBFDBC418BFCF5FB7A" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFDBC418BFCF5FB7A" box="[624,792,1168,1193]" country="Panama" county="Hammel" municipality="Coiba" name="rio Santa Clara" pageId="15" pageNumber="116" stateProvince="Veraguas">río Santa Clara</location>
, orilla del río,
<date id="FF93107CE475FFBBFC75418BFBA6FB7A" box="[953,1099,1168,1193]" pageId="15" pageNumber="116" value="2005-07-24">
<collectingDate id="EFD7E994E475FFBBFC75418BFBA6FB7A" box="[953,1099,1168,1193]" pageId="15" pageNumber="116" value="2005-07-24">24 July 2005</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFB99418BFAABFB7A" box="[1109,1350,1168,1193]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFB99418BFB4DFB7A" box="[1109,1184,1168,1193]" pageId="15" pageNumber="116">Ibáñez</collectorName>
&amp;
<collectorName id="26D8536AE475FFBBFB0D418BFAEAFB7A" box="[1217,1287,1168,1193]" pageId="15" pageNumber="116">Núñez</collectorName>
4342
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFA98418BFA76FB7A" box="[1364,1435,1168,1193]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="15" pageNumber="116">PMA</collectionCode>
!).
<collectingRegion id="49E9F85EE475FFBBFF4441AFFF08FB1E" box="[136,229,1204,1229]" country="Panama" name="Panama" pageId="15" pageNumber="116">Panama</collectingRegion>
:
<location id="8EF26067E475FFBBFF2341AFFE80FB1E" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFF2341AFFE80FB1E" box="[239,365,1204,1229]" country="Panama" county="Hammel" municipality="Coiba" name="Cerro Azul" pageId="15" pageNumber="116" stateProvince="Veraguas">Cerro Azul</location>
, near
<location id="8EF26067E475FFBBFE6341AFFDD4FB1E" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFE6341AFFDD4FB1E" box="[431,569,1204,1229]" country="Panama" county="Hammel" municipality="Coiba" name="Goofy Lake" pageId="15" pageNumber="116" stateProvince="Veraguas">Goofy Lake</location>
,
<date id="FF93107CE475FFBBFD8F41AFFD1AFB1E" box="[579,759,1204,1229]" pageId="15" pageNumber="116" value="1960-08-24">
<collectingDate id="EFD7E994E475FFBBFD8F41AFFD1AFB1E" box="[579,759,1204,1229]" pageId="15" pageNumber="116" value="1960-08-24">24 August 1960</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFCCE41AFFC60FB1E" box="[770,909,1204,1229]" italics="true" pageId="15" pageNumber="116">
<location id="8EF26067E475FFBBFCCE41AFFCB1FB1E" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFCCE41AFFCB1FB1E" box="[770,860,1204,1229]" country="Panama" county="Hammel" municipality="Coiba" name="Ebinger" pageId="15" pageNumber="116" stateProvince="Veraguas">Ebinger</location>
984
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFC5141AFFC39FB1E" box="[925,980,1204,1229]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="116" type="Herbarium">MO</collectionCode>
!).
<collectingRegion id="49E9F85EE475FFBBFC2641AFFB87FB1E" box="[1002,1130,1204,1229]" country="Panama" name="Los Santos" pageId="15" pageNumber="116">Los Santos</collectingRegion>
:
<location id="8EF26067E475FFBBFBB841AFFAEDFB1E" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFBB841AFFAEDFB1E" box="[1140,1280,1204,1229]" country="Panama" county="Hammel" municipality="Coiba" name="Loma Prieta" pageId="15" pageNumber="116" stateProvince="Veraguas">Loma Prieta</location>
,
<location id="8EF26067E475FFBBFAC741AFFA4AFB1E" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFAC741AFFA4AFB1E" box="[1291,1447,1204,1229]" country="Panama" county="Hammel" municipality="Coiba" name="Cerro Grande" pageId="15" pageNumber="116" stateProvince="Veraguas">Cerro Grande</location>
, alt.
<quantity id="4CD59B59E475FFBBFF7E41C3FEB3FB22" box="[178,350,1240,1265]" metricMagnitude="2" metricUnit="m" metricValue="7.9248" metricValueMax="8.5344" metricValueMin="7.3152" pageId="15" pageNumber="116" unit="ft" value="2600.0" valueMax="2800.0" valueMin="2400.0">
<elevation id="0000D18FE475FFBBFF7E41C3FEB3FB22" box="[178,350,1240,1265]" metricMagnitude="2" metricUnit="m" metricValue="7.9248" metricValueMax="8.5344" metricValueMin="7.3152" pageId="15" pageNumber="116" unit="ft" value="2600.0" valueMax="2800.0" valueMin="2400.0">2,4002,800 ft.</elevation>
</quantity>
<location id="8EF26067E475FFBBFEA841C3FE1DFB22" LSID="urn:lsid:plazi:treatment:038487AAE475FFBBFF444585FC19F98A:8EF26067E475FFBBFEA841C3FE1DFB22" box="[356,496,1240,1265]" country="Panama" county="Hammel" municipality="Coiba" name="Cloud forest" pageId="15" pageNumber="116" stateProvince="Veraguas">Cloud forest</location>
and disturbed margins,
<date id="FF93107CE475FFBBFD3141C3FC6AFB22" box="[765,903,1240,1265]" pageId="15" pageNumber="116" value="1967-06-08">
<collectingDate id="EFD7E994E475FFBBFD3141C3FC6AFB22" box="[765,903,1240,1265]" pageId="15" pageNumber="116" value="1967-06-08">8 June 1967</collectingDate>
</date>
,
<emphasis id="B959EAAEE475FFBBFC5E41C3FAA3FB22" box="[914,1358,1240,1265]" italics="true" pageId="15" pageNumber="116">
<collectorName id="26D8536AE475FFBBFC5E41C3FC3BFB22" box="[914,982,1240,1265]" pageId="15" pageNumber="116">Lewis</collectorName>
,
<collectorName id="26D8536AE475FFBBFC2D41C3FBC9FB22" box="[993,1060,1240,1265]" pageId="15" pageNumber="116">Baker</collectorName>
,
<collectorName id="26D8536AE475FFBBFBE341C3FB49FB22" box="[1071,1188,1240,1265]" pageId="15" pageNumber="116">MacBryde</collectorName>
&amp;
<collectorName id="26D8536AE475FFBBFB0441C3FAFDFB22" box="[1224,1296,1240,1265]" pageId="15" pageNumber="116">Oliver</collectorName>
2214
</emphasis>
(
<collectionCode id="ED3CAE79E475FFBBFA9241C3FA78FB22" box="[1374,1429,1240,1265]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="116" type="Herbarium">MO</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376537E475FFBBFF7141E7FBC4FAEA" pageId="15" pageNumber="116" type="etymology">
<paragraph id="8B9236BCE475FFBBFF7141E7FBC4FAEA" blockId="15.[136,1452,340,1625]" pageId="15" pageNumber="116">
<emphasis id="B959EAAEE475FFBBFF7141E7FEA5FAC6" bold="true" box="[189,328,1276,1301]" pageId="15" pageNumber="116">Etymology:</emphasis>
—The name comes from the Latin
<emphasis id="B959EAAEE475FFBBFD3041E7FC8DFAC6" box="[764,864,1276,1301]" italics="true" pageId="15" pageNumber="116">glandula</emphasis>
meaning “diminutive of gland”, referring to the conspicuous presence of small glands covering most floral parts and inflorescence.
</paragraph>
</subSubSection>
<subSubSection id="C3376537E475FFBBFF71405FFC19F98A" pageId="15" pageNumber="116" type="distribution">
<paragraph id="8B9236BCE475FFBBFF71405FFA87FA8E" blockId="15.[136,1452,340,1625]" box="[189,1386,1348,1373]" pageId="15" pageNumber="116">
<emphasis id="B959EAAEE475FFBBFF71405FFEB4FA8E" bold="true" box="[189,345,1348,1373]" pageId="15" pageNumber="116">Distribution:</emphasis>
—Known only from
<collectingCountry id="F33A762CE475FFBBFD89405FFD53FA8E" box="[581,702,1348,1373]" name="Costa Rica" pageId="15" pageNumber="116">Costa Rica</collectingCountry>
and
<collectingCountry id="F33A762CE475FFBBFD39405FFCBCFA8E" box="[757,849,1348,1373]" name="Panama" pageId="15" pageNumber="116">Panama</collectingCountry>
, from around sea level to up to
<quantity id="4CD59B59E475FFBBFB7F405FFB17FA8E" box="[1203,1274,1348,1373]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="116" unit="m" value="900.0">900 m</quantity>
(
<figureCitation id="13162A39E475FFBBFAC5405FFAB1FA8E" box="[1289,1372,1348,1373]" captionStart="FIGURE 13" captionStartId="18.[136,229,1935,1956]" captionTargetBox="[272,1315,1325,1909]" captionTargetId="figure-504@18.[272,1315,1325,1909]" captionTargetPageId="18" captionText="FIGURE 13. Extract of the distribution map of the glandulous species of Specklinia, with emphasis on Nicaragua, Costa Rica and Panama. Edited by D. Bogarín." figureDoi="http://doi.org/10.5281/zenodo.13635913" httpUri="https://zenodo.org/record/13635913/files/figure.png" pageId="15" pageNumber="116">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8B9236BCE475FFBBFF714073FCB6F9C2" blockId="15.[136,1452,340,1625]" pageId="15" pageNumber="116">
<emphasis id="B959EAAEE475FFBBFF714073FEE4FA52" bold="true" box="[189,265,1384,1409]" pageId="15" pageNumber="116">Notes:</emphasis>
—The name
<taxonomicName id="4C2D4D3FE475FFBBFE6D4073FD71FA52" authorityName="Pridgeon &amp; M. W. Chase. A." authorityYear="2001" baseAuthorityName="Ames" baseAuthorityYear="1923" box="[417,668,1384,1409]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="glandulosa">
<emphasis id="B959EAAEE475FFBBFE6D4073FD71FA52" box="[417,668,1384,1409]" italics="true" pageId="15" pageNumber="116">Specklinia glandulosa</emphasis>
</taxonomicName>
has been applied to most of the species treated here at some point. Nevertheless, it can be recognized by the extremely long inflorescence in relation to the leaf (close to twice the length). The sepals are relatively long and narrow in comparison with
<taxonomicName id="4C2D4D3FE475FFBBFCF940ABFC3BFA1A" box="[821,982,1456,1481]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="alajuelensis">
<emphasis id="B959EAAEE475FFBBFCF940ABFC3BFA1A" box="[821,982,1456,1481]" italics="true" pageId="15" pageNumber="116">S. alajuelensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2D4D3FE475FFBBFC2F40ABFB8BFA1A" authorityName="Schweinfurth" authorityYear="1935" box="[995,1126,1456,1481]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="pertenuis">
<emphasis id="B959EAAEE475FFBBFC2F40ABFB8BFA1A" box="[995,1126,1456,1481]" italics="true" pageId="15" pageNumber="116">S. pertenuis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D3FE475FFBBFB5740ABFADBFA1A" baseAuthorityName="Karremans" baseAuthorityYear="2945" box="[1179,1334,1456,1481]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="vittariifolia">
<emphasis id="B959EAAEE475FFBBFB5740ABFADBFA1A" box="[1179,1334,1456,1481]" italics="true" pageId="15" pageNumber="116">S. vittariifolia</emphasis>
</taxonomicName>
. Although it can produce more than a single flower per inflorescence with time, these are closely placed (sub-fascicled), and the species normally appear to have a single flowered inflorescence.
</paragraph>
<paragraph id="8B9236BCE475FFBBFF714307FC19F98A" blockId="15.[136,1452,340,1625]" pageId="15" pageNumber="116">
In
<collectingCountry id="F33A762CE475FFBBFF2C4307FEB0F9E6" box="[224,349,1564,1589]" name="Costa Rica" pageId="15" pageNumber="116">Costa Rica</collectingCountry>
this species seems to be restricted to the lowlands of the Central and South Pacific, and it is apparently seen very rarely. Most known specimens are from Central
<collectingCountry id="F33A762CE475FFBBFC58435BFC1DF98A" box="[916,1008,1600,1625]" name="Panama" pageId="15" pageNumber="116">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>