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<document id="1A4499B1A048015BBE35740CE8EFE024" ID-CLB-Dataset="25600" ID-DOI="10.11646/zootaxa.4664.4.5" ID-GBIF-Dataset="844fafcf-8f84-4039-ade1-7a03eadaf1d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3386951" ID-ZooBank="B875497D-DF28-4639-880B-2B3F52E3631B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567672571258" checkinUser="plazi" docAuthor="Boll, Piter Kehoma, Amaral, Silvana Vargas Do &amp; Leal-Zanchet, Ana Maria" docDate="2019" docId="03848782FF84FFE9FF7EFE54FD7CFCE7" docLanguage="en" docName="zootaxa.4664.4.5.pdf.imf" docOrigin="Zootaxa 4664 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luteostriata subtilis Boll &amp; Amaral &amp; Leal-Zanchet 2019, sp. nov." docType="treatment" docUuid="F03CBF98-3ADB-4666-ACDB-07BA27EED07F" docUuidSource="ZooBank" docVersion="10" lastPageNumber="548" masterDocId="FFBDFFFAFF8CFFE4FFE9FF94FFC8FFD5" masterDocTitle="Two new land planarian species (Platyhelminthes: Tricladida: Geoplanidae) from protected areas in southern Brazil" masterLastPageNumber="550" masterPageNumber="535" pageNumber="543" updateTime="1698754694161" updateUser="ExternalLinkService">
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<mods:title id="EBD6CCCC9E74EB9CA0EB47E9E1F18888">Two new land planarian species (Platyhelminthes: Tricladida: Geoplanidae) from protected areas in southern Brazil</mods:title>
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<mods:namePart id="B72657CB9BBB990A41DABDABDB1AC8B0">Boll, Piter Kehoma</mods:namePart>
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<mods:namePart id="85025A93E509F19669283FA684A95A16">Amaral, Silvana Vargas Do</mods:namePart>
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<mods:namePart id="DEB9E6CE061B992033B80C8F25C5982F">Leal-Zanchet, Ana Maria</mods:namePart>
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<mods:title id="DD25A9E64F162F79F4079A0469EDE6BC">Zootaxa</mods:title>
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<mods:date id="9E099FBA09FAD27E6A9DA3F15ED76F5F">2019</mods:date>
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<subSubSection id="C337651FFF84FFECFF7EFE54FC29FE28" pageId="8" pageNumber="543" type="nomenclature">
<paragraph id="8B923694FF84FFECFF7EFE54FCA2FE0F" blockId="8.[151,993,447,509]" box="[151,874,447,474]" pageId="8" pageNumber="543">
<heading id="D0DA81F8FF84FFECFF7EFE54FCA2FE0F" bold="true" box="[151,874,447,474]" fontSize="11" level="1" pageId="8" pageNumber="543" reason="1">
<emphasis id="B959EA86FF84FFECFF7EFE54FCA2FE0F" bold="true" box="[151,874,447,474]" pageId="8" pageNumber="543">
<taxonomicName id="4C2D4D17FF84FFECFF7EFE54FCC8FE0F" authority="Boll, Amaral &amp; Leal-Zanchet" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[151,768,447,474]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF84FFECFF7EFE54FE4FFE0F" bold="true" box="[151,391,447,474]" italics="true" pageId="8" pageNumber="543">Luteostriata subtilis</emphasis>
Boll, Amaral &amp; Leal-Zanchet
</taxonomicName>
,
<taxonomicNameLabel id="A26A57FDFF84FFECFCE5FE54FCA2FE0F" box="[780,874,448,474]" pageId="8" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<uri id="FFBC3A96FF84FFECFF7EFE77FC29FE28" box="[151,993,483,509]" pageId="8" pageNumber="543">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<taxonomicName id="4C2D4D17FF84FFECFF7EFDBCFE89FD95" authorityName="Ogren &amp; Kawakatsu" authorityYear="1990" box="[151,321,552,576]" class="Rhabditophora" family="Geoplanidae" genus="Notogynaphallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF84FFECFF7EFDBCFE89FD95" box="[151,321,552,576]" italics="true" pageId="8" pageNumber="543">Notogynaphallia</emphasis>
</taxonomicName>
sp.
<quantity id="4CD59B71FF84FFECFE82FDB3FE43FD95" box="[363,395,551,576]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="543" unit="in" value="1.0">1in</quantity>
<bibRefCitation id="EFBC4B65FF84FFECFE7BFDB3FD9FFD95" author="Baptista, V. A. &amp; Matos, L. B. &amp; Fick, I. A. &amp; Leal-Zanchet, A. M." box="[402,599,551,576]" pageId="8" pageNumber="543" pagination="293 - 297" refId="ref10300" refString="Baptista, V. A., Matos, L. B., Fick, I. A. &amp; Leal-Zanchet, A. M. (2006) Composicao das comunidades de planarias terrestres (Platyhelminthes, Tricladida, Terricola) do Parque Nacional dos Aparados da Serra, Brasil. Iheringia. Serie Zoologia, 96, 293 - 297. https: // doi. org / 10.1590 / S 0073 - 4721200600030000 4" type="journal article" year="2006">
Baptista
<emphasis id="B959EA86FF84FFECFE02FDBDFDD0FD95" box="[491,536,552,576]" italics="true" pageId="8" pageNumber="543">et al</emphasis>
. 2006
</bibRefCitation>
</paragraph>
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<taxonomicName id="4C2D4D17FF84FFECFF7EFDD2FE89FD8B" authorityName="Ogren &amp; Kawakatsu" authorityYear="1990" box="[151,321,582,606]" class="Rhabditophora" family="Geoplanidae" genus="Notogynaphallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF84FFECFF7EFDD2FE89FD8B" box="[151,321,582,606]" italics="true" pageId="8" pageNumber="543">Notogynaphallia</emphasis>
</taxonomicName>
sp.
<quantity id="4CD59B71FF84FFECFE82FDD2FE5AFD8A" box="[363,402,582,607]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="543" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFBC4B65FF84FFECFE71FDD2FDFFFD8A" author="Fick, I. A. &amp; Leal-Zanchet, A. M. &amp; Vieira, E. M." box="[408,567,582,607]" pageId="8" pageNumber="543" pagination="306 - 313" refId="ref10733" refString="Fick, I. A., Leal-Zanchet, A. M. &amp; Vieira, E. M. (2006) Community structure of land flatworms (Platyhelminthes, Terricola): comparisons between Araucaria and Atlantic forest in Southern Brazil. Invertebrate Biology, 125, 306 - 313. https: // doi. org / 10.1111 / j. 1744 - 7410.2006.00062. x" type="journal article" year="2006">
Fick
<emphasis id="B959EA86FF84FFECFE22FDD3FE30FD8B" box="[459,504,582,606]" italics="true" pageId="8" pageNumber="543">et al</emphasis>
. 2006
</bibRefCitation>
</paragraph>
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<taxonomicName id="4C2D4D17FF84FFECFF7EFDF1FE89FDA8" authorityName="Ogren &amp; Kawakatsu" authorityYear="1990" box="[151,321,613,637]" class="Rhabditophora" family="Geoplanidae" genus="Notogynaphallia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF84FFECFF7EFDF1FE89FDA8" box="[151,321,613,637]" italics="true" pageId="8" pageNumber="543">Notogynaphallia</emphasis>
</taxonomicName>
sp.
<quantity id="4CD59B71FF84FFECFE82FDF1FE5AFDA8" box="[363,402,612,637]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="543" unit="in" value="1.0">1 in</quantity>
Leal-Zanchet &amp; Baptista 2009
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF84FFECFF7EFD38FBECFC14" pageId="8" pageNumber="543" type="etymology">
<paragraph id="8B923694FF84FFECFF7EFD38FCF1FCDB" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF7EFD38FEEAFD13" bold="true" box="[151,290,684,710]" pageId="8" pageNumber="543">Etymology:</emphasis>
The specific name is a Latin adjective and refers to the three subtle and discontinuous stripes present on the dorsum, usually not visible to the naked eye, as well as to the subtle internal differences between this species and
<taxonomicName id="4C2D4D17FF84FFECFF2EFD60FCFDFCDB" authority="(Froehlich &amp; Leal-Zanchet, 2003)" authorityName="Froehlich &amp; Leal-Zanchet" authorityYear="2003" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[199,821,755,782]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF84FFECFF2EFD60FE66FCD8" box="[199,430,755,781]" italics="true" pageId="8" pageNumber="543">Luteostriata ceciliae</emphasis>
(
<bibRefCitation id="EFBC4B65FF84FFECFE55FD67FCE5FCDB" author="Froehlich, E. M. &amp; Leal-Zanchet, A. M." box="[444,813,755,782]" pageId="8" pageNumber="543" pagination="745 - 753" refId="ref11039" refString="Froehlich, E. M. &amp; Leal-Zanchet, A. M. (2003) A new species of terrestrial planarian of the genus Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes, Tricladida, Terricola) from south Brazil and some comments on the genus. Revista Brasileira de Zoologia, 20, 745 - 753. https: // doi. org / 10.1590 / S 0101 - 8175200300040003 0" type="journal article" year="2003">Froehlich &amp; Leal-Zanchet, 2003</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B923694FF84FFECFF2EFC83FBECFC14" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EFC83FEBFFCE7" bold="true" box="[199,375,791,818]" pageId="8" pageNumber="543">Type-material.</emphasis>
<materialsCitation id="3B453CC9FF84FFECFE97FC8CFBECFC14" ID-GBIF-Occurrence="2382692859" collectingDate="2000-12-11" collectionCode="MZUSP, PL" collectorName="I. A. Fick" latitude="-29.25" location="Brazil-anterior" longLatPrecision="1228" longitude="-50.25" municipality="National Park of Aparados da Serra" pageId="8" pageNumber="543" specimenCount="1" stateProvince="Cambara do Sul, Rio Grande do Sul" typeStatus="holotype">
<emphasis id="B959EA86FF84FFECFE97FC8CFE3CFCE7" bold="true" box="[382,500,792,818]" pageId="8" pageNumber="543">
<typeStatus id="54968836FF84FFECFE97FC8CFE27FCE7" box="[382,495,792,818]" pageId="8" pageNumber="543" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED3CAE51FF84FFECFE12FC8CFD9FFCE7" box="[507,599,792,818]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="543">MZUSP</collectionCode>
<collectionCode id="ED3CAE51FF84FFECFDB4FC83FD49FCE7" box="[605,641,791,818]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="543" type="Museum">PL</collectionCode>
.2159:
<emphasis id="B959EA86FF84FFECFD25FC83FD3DFCE4" box="[716,757,791,817]" italics="true" pageId="8" pageNumber="543">leg.</emphasis>
<collectorName id="26D85342FF84FFECFD12FC8CFCAFFCE4" box="[763,871,791,818]" pageId="8" pageNumber="543">I. A. Fick</collectorName>
,
<date id="FF931054FF84FFECFC9BFC8CFB80FCE7" box="[882,1096,791,818]" pageId="8" pageNumber="543" value="2000-12-11">
<collectingDate id="EFD7E9BCFF84FFECFC9BFC8CFB80FCE7" box="[882,1096,791,818]" pageId="8" pageNumber="543" value="2000-12-11">11 December 2000</collectingDate>
</date>
,
<collectingMunicipality id="6BF6ACEEFF84FFECFBBAFC83FF1AFC83" pageId="8" pageNumber="543">National Park of Aparados da Serra</collectingMunicipality>
(29°05’ to
<geoCoordinate id="EE195053FF84FFECFEA7FCAFFE64FC83" box="[334,428,827,854]" degrees="29" direction="south" minutes="15" orientation="latitude" pageId="8" pageNumber="543" precision="925" value="-29.25">29°15’S</geoCoordinate>
, 50°00’ to
<geoCoordinate id="EE195053FF84FFECFDCDFCAFFD5AFC83" box="[548,658,827,854]" degrees="50" direction="west" minutes="15" orientation="longitude" pageId="8" pageNumber="543" precision="925" value="-50.25">50°15’ W</geoCoordinate>
),
<collectingRegion id="49E9F876FF84FFECFD4CFCAFFBFFFC80" box="[677,1079,827,853]" isEnumeration="true" pageId="8" pageNumber="543">Cambará do Sul, Rio Grande do Sul</collectingRegion>
,
<location id="8EF2604FFF84FFECFBA8FCAFFB31FC80" LSID="urn:lsid:plazi:treatment:03848782FF84FFE9FF7EFE54FD7CFCE7:8EF2604FFF84FFECFBA8FCAFFB31FC80" box="[1089,1273,827,853]" latitude="-29.25" longLatPrecision="1228" longitude="-50.25" municipality="National Park of Aparados da Serra" name="Brazil-anterior" pageId="8" pageNumber="543" stateProvince="Cambara do Sul, Rio Grande do Sul">Brazil—anterior</location>
tip: transverse sections on 23 slides; anterior region at the level of the ovaries: transverse sections on 25 slides; anterior region at the level of the testes: sagittal sections on 23 slides; pre-pharyngeal region: transverse sections on 5 slides; pharynx: sagittal sections on 12 slides; copulatory apparatus: sagittal sections on 15 slides.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF84FFECFF2EFC58FAA8FB84" pageId="8" pageNumber="543" type="description">
<paragraph id="8B923694FF84FFECFF2EFC58FAA8FB84" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<materialsCitation id="3B453CC9FF84FFECFF2EFC58FA94FB84" ID-GBIF-Occurrence="2382692860" collectingDate="1999-12-22" collectingDateMax="2004-08-21" collectingDateMin="1999-12-22" collectionCode="PL" collectorName="I. A. Fick &amp; L. M. Campos" pageId="8" pageNumber="543" specimenCount="2" typeStatus="paratype">
<emphasis id="B959EA86FF84FFECFF2EFC58FE8AFC33" bold="true" box="[199,322,972,998]" pageId="8" pageNumber="543">
<typeStatus id="54968836FF84FFECFF2EFC58FEF4FC33" box="[199,316,972,998]" pageId="8" pageNumber="543" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Two specimens collected in the same locality of the
<typeStatus id="54968836FF84FFECFC81FC5FFC00FC30" box="[872,968,971,997]" pageId="8" pageNumber="543" type="holotype">holotype</typeStatus>
. MZU
<collectionCode id="ED3CAE51FF84FFECFBFAFC58FBFDFC33" box="[1043,1077,972,998]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="543" type="Museum">PL</collectionCode>
. 273:
<emphasis id="B959EA86FF84FFECFB9AFC5FFB5BFC30" box="[1139,1171,971,997]" italics="true" pageId="8" pageNumber="543">leg</emphasis>
.
<collectorName id="26D85342FF84FFECFB76FC58FACBFC30" box="[1183,1283,971,998]" pageId="8" pageNumber="543">I. A. Fick</collectorName>
,
<date id="FF931054FF84FFECFAE7FC58FF01FBDC" pageId="8" pageNumber="543" value="1999-12-22">
<collectingDate id="EFD7E9BCFF84FFECFAE7FC58FF01FBDC" pageId="8" pageNumber="543" value="1999-12-22">22 December 1999</collectingDate>
</date>
—anterior tip: transverse sections on 15 slides; anterior region at the level of the ovaries: sagittal sections on 9 slides; pre-pharyngeal region: transversal sections on 9 slides; pharynx: sagittal sections on 14 slides; copulatory apparatus: sagittal sections on 18 slides; MZU
<collectionCode id="ED3CAE51FF84FFECFE56FBACFE29FB87" box="[447,481,1080,1106]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="543" type="Museum">PL</collectionCode>
. 274:
<emphasis id="B959EA86FF84FFECFDC9FBA3FD88FB84" box="[544,576,1079,1105]" italics="true" pageId="8" pageNumber="543">leg</emphasis>
.
<collectorName id="26D85342FF84FFECFDA4FBACFD21FB87" box="[589,745,1080,1106]" pageId="8" pageNumber="543">L. M. Campos</collectorName>
,
<date id="FF931054FF84FFECFD1AFBACFC52FB84" box="[755,922,1079,1106]" pageId="8" pageNumber="543" value="2004-08-21">
<collectingDate id="EFD7E9BCFF84FFECFD1AFBACFC52FB84" box="[755,922,1079,1106]" pageId="8" pageNumber="543" value="2004-08-21">21 August 2004</collectingDate>
</date>
—whole animal preserved in 70% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF84FFECFF2EFBCFFC7BFB68" pageId="8" pageNumber="543" type="diagnosis">
<paragraph id="8B923694FF84FFECFF2EFBCFFAF4FB4C" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EFBCFFE8CFBA3" bold="true" box="[199,324,1115,1142]" pageId="8" pageNumber="543">Diagnosis:</emphasis>
species of
<taxonomicName id="4C2D4D17FF84FFECFE2CFBC8FD86FBA0" authorityName="Carbayo" authorityYear="2010" box="[453,590,1116,1141]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="543" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF84FFECFE2CFBC8FD86FBA0" box="[453,590,1116,1141]" italics="true" pageId="8" pageNumber="543">Luteostriata</emphasis>
</taxonomicName>
with light-yellow dorsum and two well-marked longitudinal black stripes; pharynx bell-shaped; large fold in the male atrium located ventrally to the opening of the ejaculatory duct.
</paragraph>
<paragraph id="8B923694FF84FFECFF2EFB37FC7BFB68" blockId="8.[151,1437,683,2041]" box="[199,947,1187,1214]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EFB37FEA6FB6B" bold="true" box="[199,366,1187,1214]" pageId="8" pageNumber="543">Type-locality:</emphasis>
Cambará do Sul, state of
<collectingRegion id="49E9F876FF84FFECFD79FB37FCACFB68" box="[656,868,1187,1213]" country="Brazil" name="Rio Grande do Sul" pageId="8" pageNumber="543">Rio Grande do Sul</collectingRegion>
,
<collectingCountry id="F33A7604FF84FFECFC86FB37FC7BFB68" box="[879,947,1187,1213]" name="Brazil" pageId="8" pageNumber="543">Brazil</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF84FFECFF2EFB53FD3EFB34" box="[199,758,1223,1250]" pageId="8" pageNumber="543" type="distribution">
<paragraph id="8B923694FF84FFECFF2EFB53FD3EFB34" blockId="8.[151,1437,683,2041]" box="[199,758,1223,1250]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EFB53FEABFB37" bold="true" box="[199,355,1223,1250]" pageId="8" pageNumber="543">Distribution:</emphasis>
Known only from the type-locality.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF84FFE9FF2EFB7FFD7CFCE7" lastPageId="13" lastPageNumber="548" pageId="8" pageNumber="543" type="description">
<paragraph id="8B923694FF84FFECFF2EFB7FFD25FA9B" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EFB7FFDF4FAD3" bold="true" box="[199,572,1259,1286]" pageId="8" pageNumber="543">Description. External features:</emphasis>
Body elongate with parallel margins, anterior tip rounded and posterior tip pointed. The maximal length when creeping reaches
<quantity id="4CD59B71FF84FFECFD36FA9BFCF8FAFF" box="[735,816,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="543" unit="mm" value="62.0">62 mm</quantity>
;
<quantity id="4CD59B71FF84FFECFCD3FA84FC40FAFF" box="[826,904,1296,1322]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="543" unit="mm" value="47.0">47 mm</quantity>
after fixation. Mouth located at the middle third of the body; gonopore at the posterior third (
<tableCitation id="C6AF032FFF84FFECFD60FAA7FD17FA9B" box="[649,735,1331,1358]" captionStart="TABLE 3" captionStartId="9.[151,239,153,177]" captionTargetPageId="9" captionText="TABLE 3. Measurements, in mm, of specimens of Luteostriata subtilis sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF85FFEDFF7EFF0DFDDBFEC8" pageId="8" pageNumber="543" tableUuid="DF52661CFF85FFEDFF7EFF0DFDDBFEC8">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8B923694FF84FFECFF2EFAC3FD45F99C" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
Alive, dorsum light yellow with two well-marked black lateral stripes; cephalic region has an orange shade that gradually fades posteriorly into the yellow color of the dorsum. Under the stereomicroscope, three additional inconspicuous stripes can be seen, one median and two paramarginal stripes. These three stripes are very thin and discontinuous, usually not visible to the naked eye (
<figureCitation id="13162A11FF84FFECFD5CFA57FCC0FA0B" box="[693,776,1475,1502]" captionStart="FIGURES 13–14" captionStartId="9.[151,264,1975,2000]" captionTargetBox="[189,1420,1186,1956]" captionTargetId="figure@9.[167,1420,1167,1956]" captionTargetPageId="9" captionText="FIGURES 13–14. Luteostriata subtilis sp. nov. in dorsal view: (13) color pattern of holotype; (14) eye pattern at the anterior end of paratype MZU PL. 273." httpUri="https://zenodo.org/record/3386961/files/figure.png" pageId="8" pageNumber="543">Fig. 13</figureCitation>
). Stripes begin behind the cephalic region and end abruptly before the posterior tip in preserved specimens (
<figureCitation id="13162A11FF84FFECFD46FA73FCCAF9D7" box="[687,770,1511,1538]" captionStart="FIGURES 13–14" captionStartId="9.[151,264,1975,2000]" captionTargetBox="[189,1420,1186,1956]" captionTargetId="figure@9.[167,1420,1167,1956]" captionTargetPageId="9" captionText="FIGURES 13–14. Luteostriata subtilis sp. nov. in dorsal view: (13) color pattern of holotype; (14) eye pattern at the anterior end of paratype MZU PL. 273." httpUri="https://zenodo.org/record/3386961/files/figure.png" pageId="8" pageNumber="543">Fig. 13</figureCitation>
). Lateral stripes correspond to about 1/12 of the body width each. Ventral surface pale yellow.After fixation, the pigmentation becomes paler, almost white after several months, with the lateral stripes not very well marked.
</paragraph>
<paragraph id="8B923694FF84FFECFF2EF9C0FB31F960" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
Eyes monolobate (with pigment cups of about
<quantity id="4CD59B71FF84FFECFD33F9C0FC9BF9BB" box="[730,851,1620,1646]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="543" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 µm</quantity>
). They surround the anterior tip and are uniserial in the cephalic region (about 7% of the body length) (
<figureCitation id="13162A11FF84FFECFD3AF9E3FCEEF947" box="[723,806,1655,1682]" captionStart="FIGURES 13–14" captionStartId="9.[151,264,1975,2000]" captionTargetBox="[189,1420,1186,1956]" captionTargetId="figure@9.[167,1420,1167,1956]" captionTargetPageId="9" captionText="FIGURES 13–14. Luteostriata subtilis sp. nov. in dorsal view: (13) color pattern of holotype; (14) eye pattern at the anterior end of paratype MZU PL. 273." httpUri="https://zenodo.org/record/3386961/files/figure.png" pageId="8" pageNumber="543">Fig. 14</figureCitation>
). After that, eyes become pluriserial and spread dorsally up to the level of the lateral stripes. Towards posterior tip, they become less numerous and uniserial.
</paragraph>
<paragraph id="8B923694FF84FFECFF2EF92BFD5FF8F7" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">
<emphasis id="B959EA86FF84FFECFF2EF92BFCE1F90F" bold="true" box="[199,809,1727,1754]" pageId="8" pageNumber="543">Sensory organs, epidermis and body musculature.</emphasis>
Sensory pits as simple invaginations (about
<quantity id="4CD59B71FF84FFECFAD9F954FA54F90F" box="[1328,1436,1728,1754]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="543" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30µm</quantity>
deep) that contour the anterior tip and occur ventromarginally in an irregular, single row until slightly after the cephalic region (about 12% of the body length).
</paragraph>
<paragraph id="8B923694FF84FFECFF2EF8BFFA9DF893" blockId="8.[151,1437,683,2041]" box="[199,1365,1835,1862]" pageId="8" pageNumber="543">
Creeping sole occupying about 80% of the body width (
<tableCitation id="C6AF032FFF84FFECFCD0F8BFFC45F893" box="[825,909,1835,1862]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="8" pageNumber="543" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
). A glandular margin is absent (
<figureCitation id="13162A11FF84FFECFB1DF8BFFA8FF893" box="[1268,1351,1835,1862]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="8" pageNumber="543">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8B923694FF84FFECFF2EF8C4FE6BF82C" blockId="8.[151,1437,683,2041]" pageId="8" pageNumber="543">Secretory cells in the pre-pharyngeal region: abundant erythrophil glands with coarse granules, rhabditogen glands with xanthophil secretion (more abundant dorsally) and a few cyanophil glands with amorphous secretion (more abundant ventrally). Glands discharging through the cephalic region of the body are similar to those of the pre-pharyngeal region, but erythrophil glands with coarse granules greatly increase in number toward the anterior tip, especially ventrally.</paragraph>
<caption id="DF52661CFF85FFEDFF7EFF0DFDDBFEC8" ID-Table-UUID="DF52661CFF85FFEDFF7EFF0DFDDBFEC8" httpUri="http://table.plazi.org/id/DF52661CFF85FFEDFF7EFF0DFDDBFEC8" pageId="9" pageNumber="544" targetBox="[161,1403,297,1150]" targetIsTable="true" targetPageId="9">
<paragraph id="8B923694FF85FFEDFF7EFF0DFDDBFEC8" blockId="9.[151,1437,153,285]" pageId="9" pageNumber="544">
<emphasis id="B959EA86FF85FFEDFF7EFF0DFEC3FF64" bold="true" box="[151,267,153,177]" pageId="9" pageNumber="544">TABLE 3.</emphasis>
Measurements, in mm, of specimens of
<taxonomicName id="4C2D4D17FF85FFEDFD28FF0EFC50FF64" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[705,920,153,177]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="9" pageNumber="544" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF85FFEDFD28FF0EFC50FF64" box="[705,920,153,177]" italics="true" pageId="9" pageNumber="544">Luteostriata subtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57FDFF85FFEDFC76FF0DFC3FFF64" box="[927,1015,153,177]" pageId="9" pageNumber="544" rank="species">
<emphasis id="B959EA86FF85FFEDFC76FF0DFC27FF64" bold="true" box="[927,1007,153,177]" pageId="9" pageNumber="544">sp. nov</emphasis>
.
</taxonomicNameLabel>
–: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length.
</paragraph>
</caption>
<paragraph id="8B923694FF85FFEDFDB1FEBDFB52FBA8" pageId="9" pageNumber="544">
<table id="F92DC434FF85001BFF48FEBDFAB3FBAB" box="[161,1403,297,1150]" gridcols="4" gridrows="19" pageId="9" pageNumber="544">
<tr id="351D34D6FF85001BFF48FEBDFAB3FE97" box="[161,1403,297,322]" gridrow="0" pageId="9" pageNumber="544" rowspan-0="1">
<th id="76CC5DAAFF85001BFDB1FEBDFD3CFE97" box="[600,756,297,322]" gridcol="1" gridrow="0" pageId="9" pageNumber="544">Holotype MZU</th>
<th id="76CC5DAAFF85001BFCB6FEBDFB85FE97" box="[863,1101,297,322]" gridcol="2" gridrow="0" pageId="9" pageNumber="544">Paratype MZU PL. 273</th>
<th id="76CC5DAAFF85001BFB64FEBDFAB3FE97" box="[1165,1403,297,322]" gridcol="3" gridrow="0" pageId="9" pageNumber="544">Paratype MZU PL. 274</th>
</tr>
<tr id="351D34D6FF85001BFF48FED9FAB3FEB3" box="[161,1403,333,358]" gridrow="1" pageId="9" pageNumber="544" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76CC5DAAFF85001BFDB1FED9FD3CFEB3" box="[600,756,333,358]" gridcol="1" gridrow="1" pageId="9" pageNumber="544">PL.2159</td>
</tr>
<tr id="351D34D6FF85001BFF48FEFAFAB3FE52" box="[161,1403,366,391]" gridrow="2" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FEFAFE19FE52" box="[161,465,366,391]" gridcol="0" gridrow="2" pageId="9" pageNumber="544">Maximum length in extension</th>
<td id="76CC5DAAFF85001BFDB1FEFAFD3CFE52" box="[600,756,366,391]" gridcol="1" gridrow="2" pageId="9" pageNumber="544">62</td>
<td id="76CC5DAAFF85001BFCB6FEFAFB85FE52" box="[863,1101,366,391]" gridcol="2" gridrow="2" pageId="9" pageNumber="544">52</td>
<td id="76CC5DAAFF85001BFB64FEFAFAB3FE52" box="[1165,1403,366,391]" gridcol="3" gridrow="2" pageId="9" pageNumber="544">35</td>
</tr>
<tr id="351D34D6FF85001BFF48FE07FAB3FE79" box="[161,1403,403,428]" gridrow="3" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FE07FE19FE79" box="[161,465,403,428]" gridcol="0" gridrow="3" pageId="9" pageNumber="544">Maximum width in extension</th>
<td id="76CC5DAAFF85001BFDB1FE07FD3CFE79" box="[600,756,403,428]" gridcol="1" gridrow="3" pageId="9" pageNumber="544">2</td>
<td id="76CC5DAAFF85001BFCB6FE07FB85FE79" box="[863,1101,403,428]" gridcol="2" gridrow="3" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFB64FE07FAB3FE79" box="[1165,1403,403,428]" gridcol="3" gridrow="3" pageId="9" pageNumber="544">2</td>
</tr>
<tr id="351D34D6FF85001BFF48FE2DFAB3FE07" box="[161,1403,441,466]" gridrow="4" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FE2DFE19FE07" box="[161,465,441,466]" gridcol="0" gridrow="4" pageId="9" pageNumber="544">Length at rest</th>
<td id="76CC5DAAFF85001BFDB1FE2DFD3CFE07" box="[600,756,441,466]" gridcol="1" gridrow="4" pageId="9" pageNumber="544">31</td>
<td id="76CC5DAAFF85001BFCB6FE2DFB85FE07" box="[863,1101,441,466]" gridcol="2" gridrow="4" pageId="9" pageNumber="544">22</td>
<td id="76CC5DAAFF85001BFB64FE2DFAB3FE07" box="[1165,1403,441,466]" gridcol="3" gridrow="4" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FE4BFAB3FE2D" box="[161,1403,479,504]" gridrow="5" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FE4BFE19FE2D" box="[161,465,479,504]" gridcol="0" gridrow="5" pageId="9" pageNumber="544">Width at rest</th>
<td id="76CC5DAAFF85001BFDB1FE4BFD3CFE2D" box="[600,756,479,504]" gridcol="1" gridrow="5" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFCB6FE4BFB85FE2D" box="[863,1101,479,504]" gridcol="2" gridrow="5" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFB64FE4BFAB3FE2D" box="[1165,1403,479,504]" gridcol="3" gridrow="5" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FD91FAB3FDCB" box="[161,1403,517,542]" gridrow="6" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FD91FE19FDCB" box="[161,465,517,542]" gridcol="0" gridrow="6" pageId="9" pageNumber="544">Length*</th>
<td id="76CC5DAAFF85001BFDB1FD91FD3CFDCB" box="[600,756,517,542]" gridcol="1" gridrow="6" pageId="9" pageNumber="544">47</td>
<td id="76CC5DAAFF85001BFCB6FD91FB85FDCB" box="[863,1101,517,542]" gridcol="2" gridrow="6" pageId="9" pageNumber="544">40</td>
<td id="76CC5DAAFF85001BFB64FD91FAB3FDCB" box="[1165,1403,517,542]" gridcol="3" gridrow="6" pageId="9" pageNumber="544">23</td>
</tr>
<tr id="351D34D6FF85001BFF48FDBFFAB3FD91" box="[161,1403,555,580]" gridrow="7" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FDBFFE19FD91" box="[161,465,555,580]" gridcol="0" gridrow="7" pageId="9" pageNumber="544">Width*</th>
<td id="76CC5DAAFF85001BFDB1FDBFFD3CFD91" box="[600,756,555,580]" gridcol="1" gridrow="7" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFCB6FDBFFB85FD91" box="[863,1101,555,580]" gridcol="2" gridrow="7" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFB64FDBFFAB3FD91" box="[1165,1403,555,580]" gridcol="3" gridrow="7" pageId="9" pageNumber="544">2</td>
</tr>
<tr id="351D34D6FF85001BFF48FDC4FAB3FDBC" box="[161,1403,592,617]" gridrow="8" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FDC4FE19FDBC" box="[161,465,592,617]" gridcol="0" gridrow="8" pageId="9" pageNumber="544">DM*</th>
<td id="76CC5DAAFF85001BFDB1FDC4FD3CFDBC" box="[600,756,592,617]" gridcol="1" gridrow="8" pageId="9" pageNumber="544">25</td>
<td id="76CC5DAAFF85001BFCB6FDC4FB85FDBC" box="[863,1101,592,617]" gridcol="2" gridrow="8" pageId="9" pageNumber="544">25</td>
<td id="76CC5DAAFF85001BFB64FDC4FAB3FDBC" box="[1165,1403,592,617]" gridcol="3" gridrow="8" pageId="9" pageNumber="544">14.5</td>
</tr>
<tr id="351D34D6FF85001BFF48FDE0FAB3FD7B" box="[161,1403,628,686]" gridrow="9" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FDE0FE19FD7B" box="[161,465,628,686]" gridcol="0" gridrow="9" pageId="9" pageNumber="544">DG*</th>
<td id="76CC5DAAFF85001BFDB1FDE0FD3CFD7B" box="[600,756,628,686]" gridcol="1" gridrow="9" pageId="9" pageNumber="544">(53.2) 35</td>
<td id="76CC5DAAFF85001BFCB6FDE0FB85FD7B" box="[863,1101,628,686]" gridcol="2" gridrow="9" pageId="9" pageNumber="544">(62.5) 33</td>
<td id="76CC5DAAFF85001BFB64FDE0FAB3FD7B" box="[1165,1403,628,686]" gridcol="3" gridrow="9" pageId="9" pageNumber="544">(63.0) 17</td>
</tr>
<tr id="351D34D6FF85001BFF48FD2DFAB3FD26" box="[161,1403,697,755]" gridrow="10" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FD2DFE19FD26" box="[161,465,697,755]" gridcol="0" gridrow="10" pageId="9" pageNumber="544">DMG*</th>
<td id="76CC5DAAFF85001BFDB1FD2DFD3CFD26" box="[600,756,697,755]" gridcol="1" gridrow="10" pageId="9" pageNumber="544">(74.5) 10</td>
<td id="76CC5DAAFF85001BFCB6FD2DFB85FD26" box="[863,1101,697,755]" gridcol="2" gridrow="10" pageId="9" pageNumber="544">(82.5) 8</td>
<td id="76CC5DAAFF85001BFB64FD2DFAB3FD26" box="[1165,1403,697,755]" gridcol="3" gridrow="10" pageId="9" pageNumber="544">(73.9) 2.5</td>
</tr>
<tr id="351D34D6FF85001BFF48FC94FAB3FCCD" box="[161,1403,768,792]" gridrow="11" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FC94FE19FCCD" box="[161,465,768,792]" gridcol="0" gridrow="11" pageId="9" pageNumber="544">DPVP*</th>
<td id="76CC5DAAFF85001BFDB1FC94FD3CFCCD" box="[600,756,768,792]" gridcol="1" gridrow="11" pageId="9" pageNumber="544">0</td>
<td id="76CC5DAAFF85001BFCB6FC94FB85FCCD" box="[863,1101,768,792]" gridcol="2" gridrow="11" pageId="9" pageNumber="544">0</td>
<td id="76CC5DAAFF85001BFB64FC94FAB3FCCD" box="[1165,1403,768,792]" gridcol="3" gridrow="11" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FCB1FAB3FCEB" box="[161,1403,805,830]" gridrow="12" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FCB1FE19FCEB" box="[161,465,805,830]" gridcol="0" gridrow="12" pageId="9" pageNumber="544">Creeping sole %</th>
<td id="76CC5DAAFF85001BFDB1FCB1FD3CFCEB" box="[600,756,805,830]" gridcol="1" gridrow="12" pageId="9" pageNumber="544">78</td>
<td id="76CC5DAAFF85001BFCB6FCB1FB85FCEB" box="[863,1101,805,830]" gridcol="2" gridrow="12" pageId="9" pageNumber="544">79</td>
<td id="76CC5DAAFF85001BFB64FCB1FAB3FCEB" box="[1165,1403,805,830]" gridcol="3" gridrow="12" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FCDFFAB3FCB1" box="[161,1403,843,868]" gridrow="13" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FCDFFE19FCB1" box="[161,465,843,868]" gridcol="0" gridrow="13" pageId="9" pageNumber="544">Ovaries</th>
<td id="76CC5DAAFF85001BFDB1FCDFFD3CFCB1" box="[600,756,843,868]" gridcol="1" gridrow="13" pageId="9" pageNumber="544">6</td>
<td id="76CC5DAAFF85001BFCB6FCDFFB85FCB1" box="[863,1101,843,868]" gridcol="2" gridrow="13" pageId="9" pageNumber="544">7</td>
<td id="76CC5DAAFF85001BFB64FCDFFAB3FCB1" box="[1165,1403,843,868]" gridcol="3" gridrow="13" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FCFBFAB3FC7C" box="[161,1403,879,937]" gridrow="14" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FCFBFE19FC7C" box="[161,465,879,937]" gridcol="0" gridrow="14" pageId="9" pageNumber="544">Anteriormost testes</th>
<td id="76CC5DAAFF85001BFDB1FCFBFD3CFC7C" box="[600,756,879,937]" gridcol="1" gridrow="14" pageId="9" pageNumber="544">(13) 10</td>
<td id="76CC5DAAFF85001BFCB6FCFBFB85FC7C" box="[863,1101,879,937]" gridcol="2" gridrow="14" pageId="9" pageNumber="544">(17.5) 7.5</td>
<td id="76CC5DAAFF85001BFB64FCFBFAB3FC7C" box="[1165,1403,879,937]" gridcol="3" gridrow="14" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FC20FAB3FC3B" box="[161,1403,948,1006]" gridrow="15" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FC20FE19FC3B" box="[161,465,948,1006]" gridcol="0" gridrow="15" pageId="9" pageNumber="544">Posteriormost testes</th>
<td id="76CC5DAAFF85001BFDB1FC20FD3CFC3B" box="[600,756,948,1006]" gridcol="1" gridrow="15" pageId="9" pageNumber="544">(16) 25.5</td>
<td id="76CC5DAAFF85001BFCB6FC20FB85FC3B" box="[863,1101,948,1006]" gridcol="2" gridrow="15" pageId="9" pageNumber="544">(19) 21</td>
<td id="76CC5DAAFF85001BFB64FC20FAB3FC3B" box="[1165,1403,948,1006]" gridcol="3" gridrow="15" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FC6DFAB3FBE7" box="[161,1403,1017,1074]" gridrow="16" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FC6DFE19FBE7" box="[161,465,1017,1074]" gridcol="0" gridrow="16" pageId="9" pageNumber="544">Prostatic vesicle</th>
<td id="76CC5DAAFF85001BFDB1FC6DFD3CFBE7" box="[600,756,1017,1074]" gridcol="1" gridrow="16" pageId="9" pageNumber="544">(54) 5.3</td>
<td id="76CC5DAAFF85001BFCB6FC6DFB85FBE7" box="[863,1101,1017,1074]" gridcol="2" gridrow="16" pageId="9" pageNumber="544">(52.5) 3.4</td>
<td id="76CC5DAAFF85001BFB64FC6DFAB3FBE7" box="[1165,1403,1017,1074]" gridcol="3" gridrow="16" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FBABFAB3FB8D" box="[161,1403,1087,1112]" gridrow="17" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FBABFE19FB8D" box="[161,465,1087,1112]" gridcol="0" gridrow="17" pageId="9" pageNumber="544">Male atrium</th>
<td id="76CC5DAAFF85001BFDB1FBABFD3CFB8D" box="[600,756,1087,1112]" gridcol="1" gridrow="17" pageId="9" pageNumber="544">3.85</td>
<td id="76CC5DAAFF85001BFCB6FBABFB85FB8D" box="[863,1101,1087,1112]" gridcol="2" gridrow="17" pageId="9" pageNumber="544">3</td>
<td id="76CC5DAAFF85001BFB64FBABFAB3FB8D" box="[1165,1403,1087,1112]" gridcol="3" gridrow="17" pageId="9" pageNumber="544">–</td>
</tr>
<tr id="351D34D6FF85001BFF48FBF1FAB3FBAB" box="[161,1403,1125,1150]" gridrow="18" pageId="9" pageNumber="544">
<th id="76CC5DAAFF85001BFF48FBF1FE19FBAB" box="[161,465,1125,1150]" gridcol="0" gridrow="18" pageId="9" pageNumber="544">Female atrium</th>
<td id="76CC5DAAFF85001BFDB1FBF1FD3CFBAB" box="[600,756,1125,1150]" gridcol="1" gridrow="18" pageId="9" pageNumber="544">1.15</td>
<td id="76CC5DAAFF85001BFCB6FBF1FB85FBAB" box="[863,1101,1125,1150]" gridcol="2" gridrow="18" pageId="9" pageNumber="544">0.8</td>
<td id="76CC5DAAFF85001BFB64FBF1FAB3FBAB" box="[1165,1403,1125,1150]" gridcol="3" gridrow="18" pageId="9" pageNumber="544">–</td>
</tr>
</table>
</paragraph>
<caption id="DF52661CFF85FFEDFF7EF823FE19F821" httpUri="https://zenodo.org/record/3386961/files/figure.png" pageId="9" pageNumber="544" startId="9.[151,264,1975,2000]" targetBox="[189,1420,1186,1956]" targetPageId="9">
<paragraph id="8B923694FF85FFEDFF7EF823FE19F821" blockId="9.[151,1436,1975,2037]" pageId="9" pageNumber="544">
<emphasis id="B959EA86FF85FFEDFF7EF823FE9DF805" bold="true" box="[151,341,1975,2000]" pageId="9" pageNumber="544">FIGURES 13–14.</emphasis>
<taxonomicName id="4C2D4D17FF85FFEDFEB5F82DFDE0F805" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[348,552,1976,2000]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="9" pageNumber="544" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF85FFEDFEB5F82DFDE0F805" box="[348,552,1976,2000]" italics="true" pageId="9" pageNumber="544">Luteostriata subtilis</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF85FFEDFDC6F82CFD49F805" bold="true" box="[559,641,1976,2000]" pageId="9" pageNumber="544">
<taxonomicNameLabel id="A26A57FDFF85FFEDFDC6F82CFD49F805" box="[559,641,1976,2000]" pageId="9" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in dorsal view: (13) color pattern of holotype; (14) eye pattern at the anterior end of paratype MZU PL. 273.
</paragraph>
</caption>
<paragraph id="8B923694FF86FFEEFF2EFF0DFAB6FE06" blockId="10.[151,1437,153,467]" pageId="10" pageNumber="545">
Cutaneous musculature in the pre-pharyngeal region similar to other species of
<taxonomicName id="4C2D4D17FF86FFEEFBDDFF0EFB75FF66" authorityName="Carbayo" authorityYear="2010" box="[1076,1213,154,179]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="10" pageNumber="545" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF86FFEEFBDDFF0EFB75FF66" box="[1076,1213,154,179]" italics="true" pageId="10" pageNumber="545">Luteostriata</emphasis>
</taxonomicName>
, having a longitudinal layer with very thick bundles, thicker ventrally than dorsally (
<figureCitation id="13162A11FF86FFEEFC9AFF29FC32FF02" box="[883,1018,189,215]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 17–19</figureCitation>
) between four and eight times that of epidermis. Musculature with about the same thickness across the dorsum, becoming lower at the body margins; ventral and dorsal musculatures with similar thickness at the sagittal plane. Mc:h 17–19% (
<tableCitation id="C6AF032FFF86FFEEFB9CFE91FB05FECA" box="[1141,1229,261,287]" captionStart="TABLE 4" captionStartId="10.[151,239,514,538]" captionTargetPageId="10" captionText="TABLE 4. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of Luteostriata subtilis sp. nov." httpUri="http://table.plazi.org/id/DF52661CFF86FFEEFF7EFD96FDEDFDB6" pageId="10" pageNumber="545" tableUuid="DF52661CFF86FFEEFF7EFD96FDEDFDB6">Table 4</tableCitation>
). The longitudinal muscle fibers of the ventral sub-epidermal musculature concentrate progressively in the median region at about the anterior 1/9 of the body, forming a rounded to lens-shaped cephalic retractor muscle (
<figureCitation id="13162A11FF86FFEEFBA0FED9FB18FEBD" box="[1097,1232,332,360]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 15, 16</figureCitation>
). Next to the anterior tip, bundles of 2–8 fibers detach from the retractor muscle and run forward towards the body margins and dorsal surface. The visualization of the musculature is hampered by the large number of erythrophil cells. Mesenchymatic musculature well developed with a similar arrangement to that described for other species of the genus (
<figureCitation id="13162A11FF86FFEEFACDFE2DFAB9FE06" box="[1316,1393,441,467]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Fig 17</figureCitation>
).
</paragraph>
<caption id="DF52661CFF86FFEEFF7EFD96FDEDFDB6" ID-Table-UUID="DF52661CFF86FFEEFF7EFD96FDEDFDB6" httpUri="http://table.plazi.org/id/DF52661CFF86FFEEFF7EFD96FDEDFDB6" pageId="10" pageNumber="545" targetBox="[161,1171,622,905]" targetIsTable="true" targetPageId="10">
<paragraph id="8B923694FF86FFEEFF7EFD96FDEDFDB6" blockId="10.[151,1436,514,611]" pageId="10" pageNumber="545">
<emphasis id="B959EA86FF86FFEEFF7EFD96FEC3FDCE" bold="true" box="[151,267,514,539]" pageId="10" pageNumber="545">TABLE 4.</emphasis>
Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of
<taxonomicName id="4C2D4D17FF86FFEEFF1AFDDFFE01FDB6" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[243,457,587,611]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="10" pageNumber="545" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF86FFEEFF1AFDDFFE01FDB6" box="[243,457,587,611]" italics="true" pageId="10" pageNumber="545">Luteostriata subtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57FDFF86FFEEFE39FDDFFDEDFDB6" box="[464,549,587,611]" pageId="10" pageNumber="545" rank="species">
<emphasis id="B959EA86FF86FFEEFE39FDDFFDEDFDB6" bold="true" box="[464,549,587,611]" pageId="10" pageNumber="545">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B923694FF86FFEEFDC7FDFAFBD5FC5C" pageId="10" pageNumber="545">
<table id="F92DC434FF86001BFF48FDFAFB5BFC5C" box="[161,1171,622,905]" gridcols="3" gridrows="8" pageId="10" pageNumber="545">
<tr id="351D34D6FF86001BFF48FDFAFB5BFD52" box="[161,1171,622,647]" gridrow="0" pageId="10" pageNumber="545" rowspan-0="1">
<th id="76CC5DAAFF86001BFDC7FDFAFD16FD52" box="[558,734,622,647]" gridcol="1" gridrow="0" pageId="10" pageNumber="545">Holotype</th>
<th id="76CC5DAAFF86001BFBEDFDFAFB5BFD52" box="[1028,1171,622,647]" gridcol="2" gridrow="0" pageId="10" pageNumber="545">Paratype</th>
</tr>
<tr id="351D34D6FF86001BFF48FD06FB5BFD7E" box="[161,1171,658,683]" gridrow="1" pageId="10" pageNumber="545" rowspan-0="1">
<td id="76CC5DAAFF86001BFDC7FD06FD16FD7E" box="[558,734,658,683]" gridcol="1" gridrow="1" pageId="10" pageNumber="545">MZUSP PL.2159</td>
<td id="76CC5DAAFF86001BFBEDFD06FB5BFD7E" box="[1028,1171,658,683]" gridcol="2" gridrow="1" pageId="10" pageNumber="545">MZU PL. 273</td>
</tr>
<tr id="351D34D6FF86001BFF48FD27FB5BFD19" box="[161,1171,691,716]" gridrow="2" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FD27FEA4FD19" box="[161,364,691,716]" gridcol="0" gridrow="2" pageId="10" pageNumber="545">Dorsal epidermis</th>
<td id="76CC5DAAFF86001BFDC7FD27FD16FD19" box="[558,734,691,716]" gridcol="1" gridrow="2" pageId="10" pageNumber="545">9</td>
<td id="76CC5DAAFF86001BFBEDFD27FB5BFD19" box="[1028,1171,691,716]" gridcol="2" gridrow="2" pageId="10" pageNumber="545">9</td>
</tr>
<tr id="351D34D6FF86001BFF48FD4DFB5BFD27" box="[161,1171,729,754]" gridrow="3" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FD4DFEA4FD27" box="[161,364,729,754]" gridcol="0" gridrow="3" pageId="10" pageNumber="545">Ventral epidermis</th>
<td id="76CC5DAAFF86001BFDC7FD4DFD16FD27" box="[558,734,729,754]" gridcol="1" gridrow="3" pageId="10" pageNumber="545">9</td>
<td id="76CC5DAAFF86001BFBEDFD4DFB5BFD27" box="[1028,1171,729,754]" gridcol="2" gridrow="3" pageId="10" pageNumber="545">9</td>
</tr>
<tr id="351D34D6FF86001BFF48FD6BFB5BFCCD" box="[161,1171,767,792]" gridrow="4" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FD6BFEA4FCCD" box="[161,364,767,792]" gridcol="0" gridrow="4" pageId="10" pageNumber="545">Dorsal musculature</th>
<td id="76CC5DAAFF86001BFDC7FD6BFD16FCCD" box="[558,734,767,792]" gridcol="1" gridrow="4" pageId="10" pageNumber="545">64</td>
<td id="76CC5DAAFF86001BFBEDFD6BFB5BFCCD" box="[1028,1171,767,792]" gridcol="2" gridrow="4" pageId="10" pageNumber="545">94</td>
</tr>
<tr id="351D34D6FF86001BFF48FCB0FB5BFCE8" box="[161,1171,804,829]" gridrow="5" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FCB0FEA4FCE8" box="[161,364,804,829]" gridcol="0" gridrow="5" pageId="10" pageNumber="545">Ventral musculature</th>
<td id="76CC5DAAFF86001BFDC7FCB0FD16FCE8" box="[558,734,804,829]" gridcol="1" gridrow="5" pageId="10" pageNumber="545">112</td>
<td id="76CC5DAAFF86001BFBEDFCB0FB5BFCE8" box="[1028,1171,804,829]" gridcol="2" gridrow="5" pageId="10" pageNumber="545">112</td>
</tr>
<tr id="351D34D6FF86001BFF48FCDEFB5BFCB6" box="[161,1171,842,867]" gridrow="6" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FCDEFEA4FCB6" box="[161,364,842,867]" gridcol="0" gridrow="6" pageId="10" pageNumber="545">Body height</th>
<td id="76CC5DAAFF86001BFDC7FCDEFD16FCB6" box="[558,734,842,867]" gridcol="1" gridrow="6" pageId="10" pageNumber="545">936</td>
<td id="76CC5DAAFF86001BFBEDFCDEFB5BFCB6" box="[1028,1171,842,867]" gridcol="2" gridrow="6" pageId="10" pageNumber="545">1156</td>
</tr>
<tr id="351D34D6FF86001BFF48FCE4FB5BFC5C" box="[161,1171,880,905]" gridrow="7" pageId="10" pageNumber="545">
<th id="76CC5DAAFF86001BFF48FCE4FEA4FC5C" box="[161,364,880,905]" gridcol="0" gridrow="7" pageId="10" pageNumber="545">Mc:h (%)</th>
<td id="76CC5DAAFF86001BFDC7FCE4FD16FC5C" box="[558,734,880,905]" gridcol="1" gridrow="7" pageId="10" pageNumber="545">19</td>
<td id="76CC5DAAFF86001BFBEDFCE4FB5BFC5C" box="[1028,1171,880,905]" gridcol="2" gridrow="7" pageId="10" pageNumber="545">17</td>
</tr>
</table>
</paragraph>
<paragraph id="8B923694FF86FFEEFF2EFC2EFB79FBC9" blockId="10.[151,1437,953,1988]" pageId="10" pageNumber="545">
<emphasis id="B959EA86FF86FFEEFF2EFC2EFEFDFC01" bold="true" box="[199,309,954,980]" pageId="10" pageNumber="545">Pharynx.</emphasis>
Pharynx bell-shaped, about 5% of the body length, occupying the first three quarters of the pharyngeal pouch (
<figureCitation id="13162A11FF86FFEEFEC9FC4AFEA5FC2D" box="[288,365,990,1016]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Fig 20</figureCitation>
). Dorsal insertion approximately on the same transversal level as the mouth, at the beginning of the median third of the pharyngeal pouch; margins highly folded (
<figureCitation id="13162A11FF86FFEEFC93FB96FC06FBC9" box="[890,974,1026,1052]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Fig. 20</figureCitation>
). Esophagus absent.
</paragraph>
<paragraph id="8B923694FF86FFEEFF2EFBB1FB3DF9C1" blockId="10.[151,1437,953,1988]" pageId="10" pageNumber="545">
<emphasis id="B959EA86FF86FFEEFF2EFBB1FE02FB95" bold="true" box="[199,458,1061,1088]" pageId="10" pageNumber="545">Reproductive system.</emphasis>
The testes are arranged in one irregular row on either side of the body, beneath the dorsal transverse mesenchymal muscles and interstitially to the intestinal branches (
<figureCitation id="13162A11FF86FFEEFBD7FBDEFB0DFBB1" box="[1086,1221,1098,1124]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 17, 18</figureCitation>
). They begin in the anterior sixth of the body and extend to near the root of the pharynx (
<tableCitation id="C6AF032FFF86FFEEFC7CFBFAFC20FB5D" box="[917,1000,1134,1160]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="10" pageNumber="545" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
). Sperm ducts, sometimes divided into ductules, dorsomedial to ovovitelline ducts, beneath fibers of the subintestinal transverse mesenchymal musculature in pre-pharyngeal region (
<figureCitation id="13162A11FF86FFEEFE56FB22FDDCFB05" box="[447,532,1206,1232]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Fig. 19</figureCitation>
). They form spermiducal vesicles posteriorly to the pharynx and divide distally into five to seven branches which open separately in the second anteriormost quarter of the prostatic vesicle (
<figureCitation id="13162A11FF86FFEEFA86FB4EFF7FFACD" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Fig. 22</figureCitation>
). The prostatic vesicle is very long, extrabulbar, and laterally sinuous; its anterior end is forked and continues laterally to the pharyngeal pouch (
<figureCitation id="13162A11FF86FFEEFDFCFAB6FDA1FAE9" box="[533,617,1314,1340]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Fig. 22</figureCitation>
). The distal part of the prostatic vesicle is directed ventrally among fibers of the common muscle coat, and then curves upward and slightly backward, entering the penis bulb (
<figureCitation id="13162A11FF86FFEEFAEFFAD2FAACFAB5" box="[1286,1380,1350,1376]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Figs. 22</figureCitation>
,
<figureCitation id="13162A11FF86FFEEFA99FAD2FA58FAB5" box="[1392,1424,1350,1376]" captionStart="FIGURES 23–24" captionStartId="12.[151,264,1960,1985]" captionTargetBox="[151,1436,940,1940]" captionTargetId="figure@12.[151,1436,940,1940]" captionTargetPageId="12" captionText="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">24</figureCitation>
). Inside the penis bulb, it continues as an ejaculatory duct, which proceeds dorsally in a sinuous path. The ejaculatory duct opens into the dorso-anterior wall of the male atrium (
<figureCitation id="13162A11FF86FFEEFCC5FA1AFC7FFA7D" box="[812,951,1422,1448]" captionStart-0="FIGURE 22" captionStart-1="FIGURES 23–24" captionStartId-0="12.[151,250,865,890]" captionStartId-1="12.[151,264,1960,1985]" captionTargetBox-0="[151,1436,481,845]" captionTargetBox-1="[151,1436,940,1940]" captionTargetId-0="figure@12.[151,1436,481,845]" captionTargetId-1="figure@12.[151,1436,940,1940]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." captionText-1="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri-0="https://zenodo.org/record/3386965/files/figure.png" httpUri-1="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">Figs. 22–24</figureCitation>
). Male atrium folded and very long, having a main fold located ventrally to the opening of the ejaculatory duct (
<figureCitation id="13162A11FF86FFEEFC48FA26FBE5FA19" box="[929,1069,1458,1484]" captionStart-0="FIGURE 22" captionStart-1="FIGURES 23–24" captionStartId-0="12.[151,250,865,890]" captionStartId-1="12.[151,264,1960,1985]" captionTargetBox-0="[151,1436,481,845]" captionTargetBox-1="[151,1436,940,1940]" captionTargetId-0="figure@12.[151,1436,481,845]" captionTargetId-1="figure@12.[151,1436,940,1940]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." captionText-1="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri-0="https://zenodo.org/record/3386965/files/figure.png" httpUri-1="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">Figs. 22–24</figureCitation>
). Few cyanophil and erythrophil glands discharge their fine granular secretion into the ejaculatory duct. Other aspects of the epithelium, musculature and secretions of the male copulatory apparatus are similar to those of other species of
<taxonomicName id="4C2D4D17FF86FFEEFB8CFA6FFB26F9C1" authorityName="Carbayo" authorityYear="2010" box="[1125,1262,1531,1556]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="10" pageNumber="545" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF86FFEEFB8CFA6FFB26F9C1" box="[1125,1262,1531,1556]" italics="true" pageId="10" pageNumber="545">Luteostriata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B923694FF86FFEEFF2EF98AFB8FF88D" blockId="10.[151,1437,953,1988]" pageId="10" pageNumber="545">
Vitellaria, situated between intestinal branches, well-developed in all observed specimens (
<figureCitation id="13162A11FF86FFEEFB31F98AFA47F9ED" box="[1240,1423,1566,1592]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 17–19, 21</figureCitation>
). Ovaries oval-shaped, about 1.5 times longer than wide, measuring about 0.2 mm in their antero-posterior axis (
<figureCitation id="13162A11FF86FFEEFAD6F9D6FA58F989" box="[1343,1424,1602,1628]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Fig. 21</figureCitation>
). They are located dorsally to the ventral nerve plate, slightly anteriorly to the anteriormost testes, approximately in the anterior sixth of the body (
<tableCitation id="C6AF032FFF86FFEEFE54F91EFDDAF971" box="[445,530,1674,1700]" captionStart="TABLE 3" captionStartId="9.[151,239,153,177]" captionTargetPageId="9" captionText="TABLE 3. Measurements, in mm, of specimens of Luteostriata subtilis sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF85FFEDFF7EFF0DFDDBFEC8" pageId="10" pageNumber="545" tableUuid="DF52661CFF85FFEDFF7EFF0DFDDBFEC8">Table 3</tableCitation>
). Ovovitelline ducts emerge dorsally from the median third of the ovaries and run posteriorly immediately above the nerve plate (
<figureCitation id="13162A11FF86FFEEFD74F93AFC83F91D" box="[669,843,1710,1736]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 17, 19, 21</figureCitation>
). Close to the gonopore, the ovovitelline ducts ascend posteromedially and unite dorsally at the level of the posterior third of the female atrium, forming a long common glandular ovovitelline duct (
<figureCitation id="13162A11FF86FFEEFE49F962FE35F8C5" box="[416,509,1782,1808]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Figs. 22</figureCitation>
,
<figureCitation id="13162A11FF86FFEEFDE0F962FDE0F8C5" box="[521,552,1782,1808]" captionStart="FIGURES 23–24" captionStartId="12.[151,264,1960,1985]" captionTargetBox="[151,1436,940,1940]" captionTargetId="figure@12.[151,1436,940,1940]" captionTargetPageId="12" captionText="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">23</figureCitation>
). The common glandular ovovitelline duct opens into the female canal, dorsally oriented, that connects to the posterior end of the female atrium (
<figureCitation id="13162A11FF86FFEEFCBAF88EFC66F8E1" box="[851,942,1818,1844]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Figs. 22</figureCitation>
,
<figureCitation id="13162A11FF86FFEEFC51F88EFC1FF8E1" box="[952,983,1818,1844]" captionStart="FIGURES 23–24" captionStartId="12.[151,264,1960,1985]" captionTargetBox="[151,1436,940,1940]" captionTargetId="figure@12.[151,1436,940,1940]" captionTargetPageId="12" captionText="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">23</figureCitation>
). The female atrium is ample, with folded walls, and has one third to one fourth of the male atrium length (
<tableCitation id="C6AF032FFF86FFEEFCBFF8AAFC62F88D" box="[854,938,1854,1880]" captionStart="TABLE 3" captionStartId="9.[151,239,153,177]" captionTargetPageId="9" captionText="TABLE 3. Measurements, in mm, of specimens of Luteostriata subtilis sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF85FFEDFF7EFF0DFDDBFEC8" pageId="10" pageNumber="545" tableUuid="DF52661CFF85FFEDFF7EFF0DFDDBFEC8">Table 3</tableCitation>
,
<figureCitation id="13162A11FF86FFEEFC5FF8AAFBD9F88D" box="[950,1041,1854,1880]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="10" pageNumber="545">Figs. 22</figureCitation>
,
<figureCitation id="13162A11FF86FFEEFBF5F8AAFBF3F88D" box="[1052,1083,1854,1880]" captionStart="FIGURES 23–24" captionStartId="12.[151,264,1960,1985]" captionTargetBox="[151,1436,940,1940]" captionTargetId="figure@12.[151,1436,940,1940]" captionTargetPageId="12" captionText="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="10" pageNumber="545">23</figureCitation>
).
</paragraph>
<paragraph id="8B923694FF86FFEEFF2EF8F6FEC1F811" blockId="10.[151,1437,953,1988]" pageId="10" pageNumber="545">
Ovovitelline ducts lined with at least partially cyanophil epithelium (
<figureCitation id="13162A11FF86FFEEFC27F8F6FB90F8A9" box="[974,1112,1890,1916]" captionStart="FIGURES 15–21" captionStartId="11.[151,264,1895,1920]" captionTargetBox="[270,1316,181,1869]" captionTargetId="figure@11.[270,1317,181,1869]" captionTargetPageId="11" captionText="FIGURES 15–21. Luteostriata subtilis sp. nov., holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary." httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="10" pageNumber="545">Figs. 17, 19</figureCitation>
). Other characteristics of the epithelium, musculature and secretions of the female copulatory apparatus are similar to those of other species in the genus.
</paragraph>
<caption id="DF52661CFF87FFEFFF7EF8F3FC29F81D" httpUri="https://zenodo.org/record/3386963/files/figure.png" pageId="11" pageNumber="546" startId="11.[151,264,1895,1920]" targetBox="[270,1316,181,1869]" targetPageId="11">
<paragraph id="8B923694FF87FFEFFF7EF8F3FC29F81D" blockId="11.[151,1436,1895,1992]" pageId="11" pageNumber="546">
<emphasis id="B959EA86FF87FFEFFF7EF8F3FE85F855" bold="true" box="[151,333,1895,1920]" pageId="11" pageNumber="546">FIGURES 15–21</emphasis>
.
<taxonomicName id="4C2D4D17FF87FFEFFEB0F8FCFDEBF855" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[345,547,1896,1920]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="11" pageNumber="546" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF87FFEFFEB0F8FCFDEBF855" box="[345,547,1896,1920]" italics="true" pageId="11" pageNumber="546">Luteostriata subtilis</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF87FFEFFDC0F8FCFDB1F855" bold="true" box="[553,633,1896,1920]" pageId="11" pageNumber="546">
<taxonomicNameLabel id="A26A57FDFF87FFEFFDC0F8FCFDB1F855" box="[553,633,1896,1920]" pageId="11" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype in transverse (15–19) and sagittal sections with the anterior to the left (20, 21): (15, 16) cephalic region; (17) pre-pharyngeal region; (18) detail of the dorsal surface in the pre-pharyngeal region; (19) detail of the ventral surface in the pre-pharyngeal region; (20) pharynx; (21) ovary.
</paragraph>
</caption>
<paragraph id="8B923694FF80FFE8FF2EFF0DFE43FE96" blockId="12.[151,1437,153,467]" pageId="12" pageNumber="547">
Gonoduct slightly inclined backward. Male and female atria are separated at the level of the gonopore by oblique dorsal and ventral, laterally displaced folds (
<figureCitation id="13162A11FF80FFE8FD04FF29FC85FF02" box="[749,845,189,215]" captionStart="FIGURE 22" captionStartId="12.[151,250,865,890]" captionTargetBox="[151,1436,481,845]" captionTargetId="figure@12.[151,1436,481,845]" captionTargetPageId="12" captionText="FIGURE 22. Luteostriata subtilis sp. nov., composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore." httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="12" pageNumber="547">Figs. 22</figureCitation>
,
<figureCitation id="13162A11FF80FFE8FCB0FF29FCB1FF02" box="[857,889,189,215]" captionStart="FIGURES 23–24" captionStartId="12.[151,264,1960,1985]" captionTargetBox="[151,1436,940,1940]" captionTargetId="figure@12.[151,1436,940,1940]" captionTargetPageId="12" captionText="FIGURES 23–24. Luteostriata subtilis sp. nov.: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section." httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="12" pageNumber="547">24</figureCitation>
). Gonoduct lined with tall columnar epithelium, ciliated, receiving the opening of abundant glands with fine erythrophil secretion, as well as few glands with amor- phous cyanophil secretion. Muscularis of the gonoduct consisting of a layer of circular fibers followed by a layer of longitudinal fibers.
</paragraph>
<paragraph id="8B923694FF80FFE8FF2EFED8FDEBFE06" blockId="12.[151,1437,153,467]" pageId="12" pageNumber="547">
<emphasis id="B959EA86FF80FFE8FF2EFED8FE27FEB2" bold="true" box="[199,495,332,359]" pageId="12" pageNumber="547">Comparative discussion.</emphasis>
The external morphology, marked by monolobated eyes and yellow dorsum with dark longitudinal stripes, and internal characteristics, such as the general morphology of copulatory apparatus, as well as the presence of a cephalic retractor muscle associated with glands, strongly support the inclusion of the new species into the genus
<taxonomicName id="4C2D4D17FF80FFE8FE7AFE2EFDD4FE06" authorityName="Carbayo" authorityYear="2010" box="[403,540,442,467]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="12" pageNumber="547" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF80FFE8FE7AFE2EFDD4FE06" box="[403,540,442,467]" italics="true" pageId="12" pageNumber="547">Luteostriata</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF52661CFF80FFE8FF7EFCF5FC8EFC4B" httpUri="https://zenodo.org/record/3386965/files/figure.png" pageId="12" pageNumber="547" startId="12.[151,250,865,890]" targetBox="[151,1436,481,845]" targetPageId="12">
<paragraph id="8B923694FF80FFE8FF7EFCF5FC8EFC4B" blockId="12.[151,1436,865,926]" pageId="12" pageNumber="547">
<emphasis id="B959EA86FF80FFE8FF7EFCF5FEE8FCAF" bold="true" box="[151,288,865,890]" pageId="12" pageNumber="547">FIGURE 22.</emphasis>
<taxonomicName id="4C2D4D17FF80FFE8FECEFCF6FE3AFCAC" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[295,498,865,889]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="12" pageNumber="547" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF80FFE8FECEFCF6FE3AFCAC" box="[295,498,865,889]" italics="true" pageId="12" pageNumber="547">Luteostriata subtilis</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF80FFE8FE10FCF6FD82FCAF" bold="true" box="[505,586,866,890]" pageId="12" pageNumber="547">
<taxonomicNameLabel id="A26A57FDFF80FFE8FE10FCF6FD82FCAF" box="[505,586,866,890]" pageId="12" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, composite reconstruction of the copulatory apparatus of the holotype from sagittal sections. Arrows indicate oblique folds at the level of the gonopore.
</paragraph>
</caption>
<caption id="DF52661CFF80FFE8FF7EF83CFD23F830" httpUri="https://zenodo.org/record/3386967/files/figure.png" pageId="12" pageNumber="547" startId="12.[151,264,1960,1985]" targetBox="[151,1436,940,1940]" targetPageId="12">
<paragraph id="8B923694FF80FFE8FF7EF83CFD23F830" blockId="12.[151,1436,1960,2021]" pageId="12" pageNumber="547">
<emphasis id="B959EA86FF80FFE8FF7EF83CFE9DF814" bold="true" box="[151,341,1960,1985]" pageId="12" pageNumber="547">FIGURES 23–24.</emphasis>
<taxonomicName id="4C2D4D17FF80FFE8FEB5F83DFDE0F814" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[348,552,1961,1985]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="12" pageNumber="547" phylum="Platyhelminthes" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="B959EA86FF80FFE8FEB5F83DFDE0F814" box="[348,552,1961,1985]" italics="true" pageId="12" pageNumber="547">Luteostriata subtilis</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF80FFE8FDC6F83DFD48F814" bold="true" box="[559,640,1961,1985]" pageId="12" pageNumber="547">
<taxonomicNameLabel id="A26A57FDFF80FFE8FDC6F83DFD48F814" box="[559,640,1961,1985]" pageId="12" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (23) copulatory apparatus of the holotype in sagittal section; (24) copulatory apparatus of the paratype MZU PL. 273 in sagittal section.
</paragraph>
</caption>
<paragraph id="8B923694FF81FFE9FF2EFF0DFD28FDC3" blockId="13.[151,1437,153,818]" pageId="13" pageNumber="548">
Externally, the color pattern of
<taxonomicName id="4C2D4D17FF81FFE9FDC4FF0EFD55FF66" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[557,669,153,179]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="subtilis">
<emphasis id="B959EA86FF81FFE9FDC4FF0EFD55FF66" box="[557,669,153,179]" italics="true" pageId="13" pageNumber="548">L. subtilis</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C2D4D17FF81FFE9FC61FF0EFABAFF61" authority="(Leal-Zanchet &amp; Froehlich, 2006)" authorityName="Leal-Zanchet &amp; Froehlich" authorityYear="2006" baseAuthorityName="Leal-Zanchet &amp; Froehlich" baseAuthorityYear="2006" box="[904,1394,153,180]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="graffi">
<emphasis id="B959EA86FF81FFE9FC61FF0EFC2DFF66" box="[904,997,153,179]" italics="true" pageId="13" pageNumber="548">L. graffi</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FC1CFF0DFAA2FF61" author="Leal-Zanchet, A. M. &amp; Froehlich, E. M." box="[1013,1386,153,180]" pageId="13" pageNumber="548" pagination="81 - 100" refId="ref11249" refString="Leal-Zanchet, A. M. &amp; Froehlich, E. M. (2006) A species complex in the genus Notogynaphallia Ogren and Kawakatsu (Platyhelminthes: Tricladida: Terricola) with a taxonomic revision of homonyms of Geoplana marginata Schultze &amp; Muller and a reinterpretation of Notogynaphallia caissara (Froehlich) anatomy. Belgian Journal of Zoology, 136, 81 - 100." type="journal article" year="2006">Leal-Zanchet &amp; Froehlich, 2006</bibRefCitation>
)
</taxonomicName>
, although the median and paramarginal stripes are easily visible in the latter (
<bibRefCitation id="EFBC4B65FF81FFE9FC0CFF28FA9BFF02" author="Leal-Zanchet, A. M. &amp; Froehlich, E. M." box="[997,1363,188,215]" pageId="13" pageNumber="548" pagination="81 - 100" refId="ref11249" refString="Leal-Zanchet, A. M. &amp; Froehlich, E. M. (2006) A species complex in the genus Notogynaphallia Ogren and Kawakatsu (Platyhelminthes: Tricladida: Terricola) with a taxonomic revision of homonyms of Geoplana marginata Schultze &amp; Muller and a reinterpretation of Notogynaphallia caissara (Froehlich) anatomy. Belgian Journal of Zoology, 136, 81 - 100." type="journal article" year="2006">Leal-Zanchet &amp; Froehlich 2006</bibRefCitation>
). In
<taxonomicName id="4C2D4D17FF81FFE9FA6FFF29FF26FF2F" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FA6FFF29FF26FF2F" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
the median stripe is well marked and continuous, while the lateral and paramarginal ones are discontinuous but highly conspicuous (
<bibRefCitation id="EFBC4B65FF81FFE9FE46FE97FCD7FEC8" author="Froehlich, E. M. &amp; Leal-Zanchet, A. M." box="[431,799,259,285]" pageId="13" pageNumber="548" pagination="745 - 753" refId="ref11039" refString="Froehlich, E. M. &amp; Leal-Zanchet, A. M. (2003) A new species of terrestrial planarian of the genus Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes, Tricladida, Terricola) from south Brazil and some comments on the genus. Revista Brasileira de Zoologia, 20, 745 - 753. https: // doi. org / 10.1590 / S 0101 - 8175200300040003 0" type="journal article" year="2003">Froehlich &amp; Leal-Zanchet 2003</bibRefCitation>
).
<taxonomicName id="4C2D4D17FF81FFE9FCDAFE90FA55FECB" authority="(Leal-Zanchet &amp; Froehlich, 2006)" authorityName="Leal-Zanchet &amp; Froehlich" authorityYear="2006" baseAuthorityName="Leal-Zanchet &amp; Froehlich" baseAuthorityYear="2006" box="[819,1437,259,286]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ernesti">
<emphasis id="B959EA86FF81FFE9FCDAFE90FBD9FEC8" box="[819,1041,260,285]" italics="true" pageId="13" pageNumber="548">Luteostriata ernesti</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FBC8FE97FA5DFECB" author="Leal-Zanchet, A. M. &amp; Froehlich, E. M." box="[1057,1429,259,286]" pageId="13" pageNumber="548" pagination="81 - 100" refId="ref11249" refString="Leal-Zanchet, A. M. &amp; Froehlich, E. M. (2006) A species complex in the genus Notogynaphallia Ogren and Kawakatsu (Platyhelminthes: Tricladida: Terricola) with a taxonomic revision of homonyms of Geoplana marginata Schultze &amp; Muller and a reinterpretation of Notogynaphallia caissara (Froehlich) anatomy. Belgian Journal of Zoology, 136, 81 - 100." type="journal article" year="2006">Leal-Zanchet &amp; Froehlich, 2006</bibRefCitation>
)
</taxonomicName>
has two well-marked paramarginal stripes and three narrower internal lines, sometimes discontinuous, while both
<taxonomicName id="4C2D4D17FF81FFE9FF7EFEDFFE29FEB1" authority="(Froehlich, 1955)" authorityName="Froehlich" authorityYear="1955" baseAuthorityName="Froehlich" baseAuthorityYear="1955" box="[151,481,329,356]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="caissara">
<emphasis id="B959EA86FF81FFE9FF7EFEDFFEDDFEB1" box="[151,277,331,356]" italics="true" pageId="13" pageNumber="548">L. caissara</emphasis>
(Froehlich, 1955)
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF81FFE9FDF0FEDFFB8EFEB1" authority="(Lemos &amp; Leal-Zanchet, 2008)" authorityName="Lemos &amp; Leal-Zanchet" authorityYear="2008" baseAuthorityName="Lemos &amp; Leal-Zanchet" baseAuthorityYear="2008" box="[537,1094,330,356]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="pseudoceciliae">
<emphasis id="B959EA86FF81FFE9FDF0FEDFFD15FEB1" box="[537,733,330,356]" italics="true" pageId="13" pageNumber="548">L. pseudoceciliae</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FD04FEDEFBF7FEB1" author="Lemos, V. S. &amp; Leal-Zanchet, A. M." box="[749,1087,330,356]" pageId="13" pageNumber="548" pagination="28 - 46" refId="ref11383" refString="Lemos, V. S. &amp; Leal-Zanchet, A. M. (2008) Two new species of Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes: Tricladida: Terricola) from Southern Brazil. Zootaxa, 1907 (1), 28 - 46. https: // doi. org / 10.11646 / zootaxa. 1907.1. 2" type="journal article" year="2008">Lemos &amp; Leal-Zanchet, 2008</bibRefCitation>
)
</taxonomicName>
have five equally marked and continuous stripes (
<bibRefCitation id="EFBC4B65FF81FFE9FE98FEFAFD14FE5D" author="Leal-Zanchet, A. M. &amp; Froehlich, E. M." box="[369,732,366,392]" pageId="13" pageNumber="548" pagination="81 - 100" refId="ref11249" refString="Leal-Zanchet, A. M. &amp; Froehlich, E. M. (2006) A species complex in the genus Notogynaphallia Ogren and Kawakatsu (Platyhelminthes: Tricladida: Terricola) with a taxonomic revision of homonyms of Geoplana marginata Schultze &amp; Muller and a reinterpretation of Notogynaphallia caissara (Froehlich) anatomy. Belgian Journal of Zoology, 136, 81 - 100." type="journal article" year="2006">Leal-Zanchet &amp; Froehlich 2006</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF81FFE9FD0EFEFAFBFAFE5D" author="Lemos, V. S. &amp; Leal-Zanchet, A. M." box="[743,1074,366,392]" pageId="13" pageNumber="548" pagination="28 - 46" refId="ref11383" refString="Lemos, V. S. &amp; Leal-Zanchet, A. M. (2008) Two new species of Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes: Tricladida: Terricola) from Southern Brazil. Zootaxa, 1907 (1), 28 - 46. https: // doi. org / 10.11646 / zootaxa. 1907.1. 2" type="journal article" year="2008">Lemos &amp; Leal-Zanchet 2008</bibRefCitation>
). In
<taxonomicName id="4C2D4D17FF81FFE9FB8AFEFBFABBFE5D" authority="(Froehlich, 1959)" authorityName="Froehlich" authorityYear="1959" baseAuthorityName="Froehlich" baseAuthorityYear="1959" box="[1123,1395,365,392]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="fita">
<emphasis id="B959EA86FF81FFE9FB8AFEFBFB6EFE5D" box="[1123,1190,366,392]" italics="true" pageId="13" pageNumber="548">L. fita</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FB5FFEFAFAA2FE5D" author="Froehlich, C. G." box="[1206,1386,365,392]" pageId="13" pageNumber="548" pagination="201 - 265" refId="ref10922" refString="Froehlich, C. G. (1959) On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Serie Zoologia, 22, 201 - 265. https: // doi. org / 10.11606 / issn. 2526 - 3382. bffclzoologia. 1959.12032 6" type="journal article" year="1959">Froehlich, 1959</bibRefCitation>
)
</taxonomicName>
the median stripe is marked only in the first half of the body (
<bibRefCitation id="EFBC4B65FF81FFE9FCCDFE05FC1CFE7E" author="Froehlich, C. G." box="[804,980,400,427]" pageId="13" pageNumber="548" pagination="201 - 265" refId="ref10922" refString="Froehlich, C. G. (1959) On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Serie Zoologia, 22, 201 - 265. https: // doi. org / 10.11606 / issn. 2526 - 3382. bffclzoologia. 1959.12032 6" type="journal article" year="1959">Froehlich 1959</bibRefCitation>
). The remaining species have a different number of stripes:
<taxonomicName id="4C2D4D17FF81FFE9FE98FE22FD56FE1A" authority="(Graff, 1899)" authorityName="Graff" authorityYear="1899" baseAuthorityName="Graff" baseAuthorityYear="1899" box="[369,670,437,463]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="abundans">
<emphasis id="B959EA86FF81FFE9FE98FE22FE36FE1A" box="[369,510,437,463]" italics="true" pageId="13" pageNumber="548">L. abundans</emphasis>
(Graff, 1899)
</taxonomicName>
has seven,
<taxonomicName id="4C2D4D17FF81FFE9FCCFFE22FBA8FE05" authority="(Diesing, 1861)" authorityName="Diesing" authorityYear="1861" baseAuthorityName="Diesing" baseAuthorityYear="1861" box="[806,1120,437,464]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="muelleri">
<emphasis id="B959EA86FF81FFE9FCCFFE22FC6BFE1A" box="[806,931,437,463]" italics="true" pageId="13" pageNumber="548">L. muelleri</emphasis>
(Diesing, 1861)
</taxonomicName>
, one or three, and
<taxonomicName id="4C2D4D17FF81FFE9FAD2FE22FE37FE27" authority="(Lemos &amp; Leal-Zanchet, 2008)" authorityName="Lemos &amp; Leal-Zanchet" authorityYear="2008" baseAuthorityName="Lemos &amp; Leal-Zanchet" baseAuthorityYear="2008" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="arturi">
<emphasis id="B959EA86FF81FFE9FAD2FE22FA54FE1A" box="[1339,1436,438,463]" italics="true" pageId="13" pageNumber="548">L. arturi</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FF76FE4CFE3FFE27" author="Lemos, V. S. &amp; Leal-Zanchet, A. M." box="[159,503,472,498]" pageId="13" pageNumber="548" pagination="28 - 46" refId="ref11383" refString="Lemos, V. S. &amp; Leal-Zanchet, A. M. (2008) Two new species of Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes: Tricladida: Terricola) from Southern Brazil. Zootaxa, 1907 (1), 28 - 46. https: // doi. org / 10.11646 / zootaxa. 1907.1. 2" type="journal article" year="2008">Lemos &amp; Leal-Zanchet, 2008</bibRefCitation>
)
</taxonomicName>
, two lateral bands with dark inner and outer margins (
<bibRefCitation id="EFBC4B65FF81FFE9FB98FE4CFF1BFDC0" author="Leal-Zanchet, A. M. &amp; Froehlich, E. M." pageId="13" pageNumber="548" pagination="81 - 100" refId="ref11249" refString="Leal-Zanchet, A. M. &amp; Froehlich, E. M. (2006) A species complex in the genus Notogynaphallia Ogren and Kawakatsu (Platyhelminthes: Tricladida: Terricola) with a taxonomic revision of homonyms of Geoplana marginata Schultze &amp; Muller and a reinterpretation of Notogynaphallia caissara (Froehlich) anatomy. Belgian Journal of Zoology, 136, 81 - 100." type="journal article" year="2006">Leal-Zanchet &amp; Froehlich 2006</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF81FFE9FF37FE68FDEDFDC3" author="Lemos, V. S. &amp; Leal-Zanchet, A. M." box="[222,549,508,534]" pageId="13" pageNumber="548" pagination="28 - 46" refId="ref11383" refString="Lemos, V. S. &amp; Leal-Zanchet, A. M. (2008) Two new species of Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes: Tricladida: Terricola) from Southern Brazil. Zootaxa, 1907 (1), 28 - 46. https: // doi. org / 10.11646 / zootaxa. 1907.1. 2" type="journal article" year="2008">Lemos &amp; Leal-Zanchet 2008</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF81FFE9FDD8FE68FD1CFDC3" author="Carbayo, F." box="[561,724,508,534]" pageId="13" pageNumber="548" pagination="91 - 101" refId="ref10444" refString="Carbayo, F. (2010) A new genus for seven Brazilian land planarian species, split off from Notogynaphallia (Platyhelminthes, Tricladida). Belgian Journal of Zoology, 140, 91 - 101." type="journal article" year="2010">Carbayo 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8B923694FF81FFE9FF2EFD8BFC11FD3E" blockId="13.[151,1437,153,818]" pageId="13" pageNumber="548">
The body shape, as well as several internal characters, such as the pharynx, the proximally-forked prostatic vesicle, the branched sperm ducts, and the long and folded male atrium, are similar to those of
<taxonomicName id="4C2D4D17FF81FFE9FB5BFDD7FAEEFD88" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[1202,1318,579,605]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FB5BFDD7FAEEFD88" box="[1202,1318,579,605]" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
. The main differences are related to the opening of the ejaculatory duct, which is displaced ventrally in
<taxonomicName id="4C2D4D17FF81FFE9FB70FDF3FAC5FD55" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[1177,1293,614,640]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FB70FDF3FAC5FD55" box="[1177,1293,614,640]" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
and dorsally in
<taxonomicName id="4C2D4D17FF81FFE9FF5CFD1EFEEEFD76" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[181,294,649,675]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="subtilis">
<emphasis id="B959EA86FF81FFE9FF5CFD1EFEEEFD76" box="[181,294,649,675]" italics="true" pageId="13" pageNumber="548">L. subtilis</emphasis>
</taxonomicName>
, the position of the main male atrium fold, which is dorsal to the ejaculatory duct in
<taxonomicName id="4C2D4D17FF81FFE9FB1CFD1EFAA3FD76" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[1269,1387,649,675]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FB1CFD1EFAA3FD76" box="[1269,1387,649,675]" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
and ventral in
<taxonomicName id="4C2D4D17FF81FFE9FEEEFD3AFEBCFD12" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[263,372,685,711]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="subtilis">
<emphasis id="B959EA86FF81FFE9FEEEFD3AFEBCFD12" box="[263,372,685,711]" italics="true" pageId="13" pageNumber="548">L. subtilis</emphasis>
</taxonomicName>
, and the distal portion of the prostatic vesicle, which is directed dorsally against the muscle coat in
<taxonomicName id="4C2D4D17FF81FFE9FF5DFD45FEE1FD3F" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[180,297,720,746]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FF5DFD45FEE1FD3F" box="[180,297,720,746]" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
and ventrally in
<taxonomicName id="4C2D4D17FF81FFE9FE0FFD45FC1DFD3E" authority="(Froehlich &amp; Leal-Zanchet 2003)" authorityName="Froehlich &amp; Leal-Zanchet" authorityYear="2003" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[486,981,720,747]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="subtilis">
<emphasis id="B959EA86FF81FFE9FE0FFD45FD9DFD3F" box="[486,597,720,746]" italics="true" pageId="13" pageNumber="548">L. subtilis</emphasis>
(
<bibRefCitation id="EFBC4B65FF81FFE9FD8AFD44FC05FD3E" author="Froehlich, E. M. &amp; Leal-Zanchet, A. M." box="[611,973,720,747]" pageId="13" pageNumber="548" pagination="745 - 753" refId="ref11039" refString="Froehlich, E. M. &amp; Leal-Zanchet, A. M. (2003) A new species of terrestrial planarian of the genus Notogynaphallia Ogren &amp; Kawakatsu (Platyhelminthes, Tricladida, Terricola) from south Brazil and some comments on the genus. Revista Brasileira de Zoologia, 20, 745 - 753. https: // doi. org / 10.1590 / S 0101 - 8175200300040003 0" type="journal article" year="2003">Froehlich &amp; Leal-Zanchet 2003</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B923694FF81FFE9FF2EFD60FD7CFCE7" blockId="13.[151,1437,153,818]" pageId="13" pageNumber="548">
The combination of those characters of both external and internal morphology differentiates
<taxonomicName id="4C2D4D17FF81FFE9FB3AFD61FA89FCDB" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" box="[1235,1345,756,782]" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="subtilis">
<emphasis id="B959EA86FF81FFE9FB3AFD61FA89FCDB" box="[1235,1345,756,782]" italics="true" pageId="13" pageNumber="548">L. subtilis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C2D4D17FF81FFE9FA6FFD61FF26FCE4" baseAuthorityName="Froehlich &amp; Leal-Zanchet" baseAuthorityYear="2003" class="Rhabditophora" family="Geoplanidae" genus="Luteostriata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="13" pageNumber="548" phylum="Platyhelminthes" rank="species" species="ceciliae">
<emphasis id="B959EA86FF81FFE9FA6FFD61FF26FCE4" italics="true" pageId="13" pageNumber="548">L. ceciliae</emphasis>
</taxonomicName>
and from the other species of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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