296 lines
35 KiB
XML
296 lines
35 KiB
XML
<document id="859CA91FFE8AFDDA1BA8F57BF097E681" ID-CLB-Dataset="20762" ID-DOI="10.11646/zootaxa.3365.1.1" ID-GBIF-Dataset="1db9bc74-46dd-4263-9784-88a59bd05d59" ID-ISSN="11755334" ID-Zenodo-Dep="6038900" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787146833" checkinUser="felipe" docAuthor="Garcia, Francisco Hita & Fisher, Brian L." docDate="2012" docId="03EF6217BF49FFB00AC0FC6F9C9FAE93" docLanguage="en" docName="zt03365p123.pdf" docOrigin="Zootaxa 3365" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tetramorium tsingy Garcia & Fisher 2012, sp. n." docType="treatment" docVersion="8" lastPageNumber="115" masterDocId="FFD61A6FBF39FFC20A57FF889C5BAB77" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups" masterLastPageNumber="123" masterPageNumber="1" pageNumber="113" updateTime="1699142484625" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="840825E7571EA6785F3E8C7BAA0912B6">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups</mods:title>
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<mods:namePart id="A54B8F2438C4C6A19820229434196ECD">Garcia, Francisco Hita</mods:namePart>
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<mods:namePart id="371231C7D38FD69DA542E26DB17D0BA9">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="BCCF422EA65744EAFFDD9CE04E5CD272">text</mods:typeOfResource>
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<mods:title id="C09B14B7DD81099C625AC062F138A6DA">Zootaxa</mods:title>
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<mods:part id="F34204292882B55B665E9EDE7E84958D">
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<mods:date id="5206DEDCC71E639E83A916CC992674F9">2012</mods:date>
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<mods:number id="17259C592C297AEB88BF1321F79BE7A4">2012-07-04</mods:number>
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<mods:detail id="C35933E83054B4968AC39601BE899617" type="volume">
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<mods:number id="F8A9CF44C3C87C7293D70C12BB285D89">3365</mods:number>
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<mods:classification id="F0B5A6748CC49C702AD676AE7AEE0C13">journal article</mods:classification>
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<mods:identifier id="56F4F79AF040508C0E4F2DC05FB829E7" type="CLB-Dataset">20762</mods:identifier>
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<mods:identifier id="81B6C86CE0446DC00A92A32FA9227AF8" type="DOI">10.11646/zootaxa.3365.1.1</mods:identifier>
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<treatment id="03EF6217BF49FFB00AC0FC6F9C9FAE93" ID-DOI="http://doi.org/10.5281/zenodo.5253696" ID-GBIF-Taxon="193281052" ID-Zenodo-Dep="5253696" LSID="urn:lsid:plazi:treatment:03EF6217BF49FFB00AC0FC6F9C9FAE93" httpUri="http://treatment.plazi.org/id/03EF6217BF49FFB00AC0FC6F9C9FAE93" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113">
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<subSubSection id="C35C808ABF49FFB20AC0FC6F9D96AF76" box="[151,461,999,1025]" pageId="112" pageNumber="113" type="nomenclature">
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<paragraph id="8BF9D301BF49FFB20AC0FC6F9D96AF76" blockId="112.[151,461,999,1058]" box="[151,461,999,1025]" pageId="112" pageNumber="113">
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<heading id="D0B1646DBF49FFB20AC0FC6F9D96AF76" bold="true" box="[151,461,999,1025]" fontSize="11" level="1" pageId="112" pageNumber="113" reason="1">
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<emphasis id="B9320F13BF49FFB20AC0FC6F9D96AF76" bold="true" box="[151,461,999,1025]" pageId="112" pageNumber="113">
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<taxonomicName id="4C46A882BF49FFB20AC0FC6F9DDFAF76" authority="Garcia & Fisher, 2012" authorityName="Garcia & Fisher" authorityYear="2012" box="[151,388,999,1025]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">
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<emphasis id="B9320F13BF49FFB20AC0FC6F9DDFAF76" bold="true" box="[151,388,999,1025]" italics="true" pageId="112" pageNumber="113">Tetramorium tsingy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A201B268BF49FFB20BDCFC6F9D96AF76" box="[395,461,999,1025]" pageId="112" pageNumber="113" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C808ABF49FFB20AC0FB829DE9AF55" box="[151,434,1034,1058]" pageId="112" pageNumber="113" type="description">
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<paragraph id="8BF9D301BF49FFB20AC0FB829DE9AF55" blockId="112.[151,461,999,1058]" box="[151,434,1034,1058]" pageId="112" pageNumber="113">
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(
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<figureCitation id="137DCF84BF49FFB20AC8FB829D58AF55" box="[159,259,1034,1058]" captionStart="FIGURES 156 & 157. 156" captionStartId="112.[151,263,791,814]" captionTargetBox="[151,1436,319,767]" captionTargetId="figure-588@112.[151,1436,319,767]" captionTargetPageId="112" captionText="FIGURES 156 & 157. 156. Body of T.tyrion in profile showing the minute propodeal teeth and short, fine erect pilosity on the first gastral tergite—CASENT0249085 (Estella Ortega 2011). 157. Body of T. tsingy in profile showing the moderately short propodeal spines and short and strongly appressed pubescence on the first gastral tergite—CASENT0426807 (Shannon Hartman 2011)." figureDoi="http://doi.org/10.5281/zenodo.6039067" httpUri="https://zenodo.org/record/6039067/files/figure.png" pageId="112" pageNumber="113">Figs 157</figureCitation>
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,
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<figureCitation id="137DCF84BF49FFB20B58FB829DF1AF55" box="[271,426,1034,1058]" captionStart="FIGURES 158–160" captionStartId="113.[151,264,1974,1997]" captionTargetBox="[181,1405,193,1953]" captionTargetId="figure-41@113.[181,1405,193,1953]" captionTargetPageId="113" captionText="FIGURES 158–160. T. tsingy, holotype—CASENT0426807 (Shannon Hartman 2011). 158. Body in profile. 159. Body in dorsal view. 160. Head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.6039071" httpUri="https://zenodo.org/record/6039071/files/figure.png" pageId="112" pageNumber="113">158, 159, 160</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C808ABF49FFB20AC0FBDA9E1BAFA1" pageId="112" pageNumber="113" type="materials_examined">
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<paragraph id="8BF9D301BF49FFB20AC0FBDA9E1BAFA1" blockId="112.[151,1436,1106,1238]" pageId="112" pageNumber="113">
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<materialsCitation id="3B2ED95CBF49FFB20AC0FBDA9F88AFC5" ID-GBIF-Occurrence="3462705523" accessionNumber="BLF04392" collectingDate="2001-11-11" collectingDateMax="2001-11-15" collectingDateMin="2001-11-11" collectionCode="CASC" collectorName="B. L. Fisher & C. Griswold" country="Madagascar" elevation="100" latitude="-19.13222" location="Ankidrodroa River" longLatPrecision="1" longitude="44.81467" municipality="Parc National Tsingy de Bemaraha" pageId="112" pageNumber="113" specimenCode="CASENT0426807" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
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<typeStatus id="54FD6DA3BF49FFB20AC0FBDA9CA4AF1D" box="[151,255,1106,1130]" pageId="112" pageNumber="113" type="holotype">Holotype</typeStatus>
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worker,
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<collectingCountry id="F3519391BF49FFB20B33FBDA9E75AF1D" box="[356,558,1106,1130]" name="Madagascar" pageId="112" pageNumber="113">MADAGASCAR</collectingCountry>
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,
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<collectingRegion id="49821DE3BF49FFB2086CFBDA9EE1AF1D" box="[571,698,1106,1130]" country="Madagascar" name="Mahajanga" pageId="112" pageNumber="113">Mahajanga</collectingRegion>
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,
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<collectingMunicipality id="6B9D497BBF49FFB20891FBDB980FAF1D" box="[710,1108,1106,1130]" pageId="112" pageNumber="113">Parc National Tsingy de Bemaraha</collectingMunicipality>
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,
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<locationDeviation id="9A4BCA23BF49FFB20E37FBDA99C3AF1D" box="[1120,1432,1106,1130]" location="Bekopaka" pageId="112" pageNumber="113">
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<quantity id="4CBE7EE4BF49FFB20E37FBDA98F5AF1D" box="[1120,1198,1106,1130]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="112" pageNumber="113" unit="km" value="2.5">2.5 km</quantity>
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62° ENE Bekopaka
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</locationDeviation>
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,
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<location id="8E9985DABF49FFB20AC0FBFE9D34AFF9" LSID="urn:lsid:plazi:treatment:03EF6217BF49FFB00AC0FC6F9C9FAE93:8E9985DABF49FFB20AC0FBFE9D34AFF9" box="[151,367,1142,1166]" country="Madagascar" latitude="-19.13222" longLatPrecision="1" longitude="44.81467" municipality="Parc National Tsingy de Bemaraha" name="Ankidrodroa River" pageId="112" pageNumber="113" stateProvince="Mahajanga">Ankidrodroa River</location>
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,
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<geoCoordinate id="EE72B5C6BF49FFB20B2AFBFE9E5BAFF9" box="[381,512,1142,1166]" degrees="19.13222" direction="south" orientation="latitude" pageId="112" pageNumber="113" precision="1" value="-19.13222">19.13222 S</geoCoordinate>
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,
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<geoCoordinate id="EE72B5C6BF49FFB20858FBFE9ECEAFF9" box="[527,661,1142,1166]" degrees="44.81467" direction="east" orientation="longitude" pageId="112" pageNumber="113" precision="1" value="44.81467">44.81467 E</geoCoordinate>
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,
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||
<quantity id="4CBE7EE4BF49FFB208F4FBFE9EB5AFF9" box="[675,750,1142,1166]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="113" unit="m" value="100.0">
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||
<elevation id="006B3432BF49FFB208F4FBFE9EB5AFF9" box="[675,750,1142,1166]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="113" unit="m" value="100.0">100 m</elevation>
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</quantity>
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, tropical dry forest on tsingy, ground nest, BLF04392,
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<date id="FFF8F5C1BF49FFB20F3BFBFE9D4DAFC5" pageId="112" pageNumber="113" value="2001-11-11" valueMax="2001-11-15" valueMin="2001-11-11">
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||
<collectingDate id="EFBC0C29BF49FFB20F3BFBFE9D4DAFC5" pageId="112" pageNumber="113" value="2001-11-11" valueMax="2001-11-15" valueMin="2001-11-11">11.– 15.XI.2001</collectingDate>
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||
</date>
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||
(
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<emphasis id="B9320F13BF49FFB20B7FFB139EDBAFC4" box="[296,640,1179,1203]" italics="true" pageId="112" pageNumber="113">
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<collectorName id="26B3B6D7BF49FFB20B7FFB139DFCAFC4" box="[296,423,1179,1203]" pageId="112" pageNumber="113">B.L. Fisher</collectorName>
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,
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<collectorName id="26B3B6D7BF49FFB20BE4FB139E66AFC4" box="[435,573,1179,1203]" pageId="112" pageNumber="113">C. Griswold</collectorName>
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et al.
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</emphasis>
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) (
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<collectionCode id="ED574BC4BF49FFB208CEFB129EB3AFC5" box="[665,744,1178,1202]" pageId="112" pageNumber="113">CASC</collectionCode>
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:
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<specimenCode id="DBE07B7ABF49FFB208A4FB129F9CAFC5" box="[755,967,1178,1202]" collectionCode="CASC" pageId="112" pageNumber="113">CASENT0426807</specimenCode>
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).
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</materialsCitation>
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||
<materialsCitation id="3B2ED95CBF49FFB2098CFB129E1BAFA1" ID-GBIF-Occurrence="3462705948" accessionNumber="BLF04392" collectingDate="2001-11-11" collectingDateMax="2001-11-15" collectingDateMin="2001-11-11" collectionCode="CASC" collectorName="B. L. Fisher & C. Griswold" country="Madagascar" elevation="100" latitude="-19.13222" location="Ankidrodroa River" longLatPrecision="1" longitude="44.81467" municipality="Parc National Tsingy de Bemaraha" pageId="112" pageNumber="113" specimenCode="CASENT0247113" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
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<specimenCount id="9D401888BF49FFB2098CFB129892AFC5" box="[987,1225,1178,1203]" pageId="112" pageNumber="113">One paratype worker</specimenCount>
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with same data as holotype (
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<collectionCode id="ED574BC4BF49FFB20B5FFB369D0CAFA1" box="[264,343,1214,1238]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="112" pageNumber="113">CASC</collectionCode>
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||
:
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<specimenCode id="DBE07B7ABF49FFB20B36FB369E6FAFA1" box="[353,564,1214,1238]" collectionCode="CASC" pageId="112" pageNumber="113">CASENT0247113</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35C808ABF49FFB20AC0FA8E9F17AE11" pageId="112" pageNumber="113" type="diagnosis">
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<paragraph id="8BF9D301BF49FFB20AC0FA8E9D51AE68" blockId="112.[151,1436,1286,1382]" box="[151,266,1286,1311]" pageId="112" pageNumber="113">
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<heading id="D0B1646DBF49FFB20AC0FA8E9D51AE68" bold="true" box="[151,266,1286,1311]" fontSize="10" level="2" pageId="112" pageNumber="113" reason="2">
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<emphasis id="B9320F13BF49FFB20AC0FA8E9D51AE68" bold="true" box="[151,266,1286,1311]" pageId="112" pageNumber="113">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF9D301BF49FFB20AC0FAA39F17AE11" blockId="112.[151,1436,1286,1382]" pageId="112" pageNumber="113">
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<taxonomicName id="4C46A882BF49FFB20AC0FAA39D28AE35" authorityName="Garcia & Fisher" authorityYear="2012" box="[151,371,1323,1346]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="tsingy">
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<emphasis id="B9320F13BF49FFB20AC0FAA39D28AE35" box="[151,371,1323,1346]" italics="true" pageId="112" pageNumber="113">Tetramorium tsingy</emphasis>
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</taxonomicName>
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is easily identifiable within its species group due to its strongly appressed pubescence on the first gastral tergite and the moderately short propodeal spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C808ABF49FFB00AC0FA1E9F53AAB8" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113" type="description">
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<paragraph id="8BF9D301BF49FFB20AC0FA1E9D7AAED8" blockId="112.[151,1436,1430,2030]" box="[151,289,1430,1455]" pageId="112" pageNumber="113">
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||
<heading id="D0B1646DBF49FFB20AC0FA1E9D7AAED8" bold="true" box="[151,289,1430,1455]" fontSize="10" level="2" pageId="112" pageNumber="113" reason="2">
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<emphasis id="B9320F13BF49FFB20AC0FA1E9D7AAED8" bold="true" box="[151,289,1430,1455]" pageId="112" pageNumber="113">Description</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="8BF9D301BF49FFB00AC0FA339F53AAB8" blockId="112.[151,1436,1430,2030]" lastBlockId="114.[151,1436,151,463]" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113">
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HL 0.63–0.66; HW 0.60–0.62; SL 0.43–0.44; EL 0.14;
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<collectingCountry id="F3519391BF49FFB20941FA339F62AEA5" box="[790,825,1467,1490]" name="Philippines" pageId="112" pageNumber="113">PH</collectingCountry>
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0.33–0.34; PW 0.42; WL 0.76; PSL 0.11; PTL 0.16; PTH 0.24; PTW 0.15; PPL 0.21; PPH 0.24; PPW 0.22–0.23; CI 95; SI 71–72; OI 23; DMI 55–56; LMI 43–45; PSLI 17; PeNI 35–36; LPeI 65–67; DPeI 94; PpNI 52–55; LPpI 88; DPpI 105–110; PPI 152–153 (two measured). Head weakly longer than wide (CI 95). Anterior clypeal margin with distinct median impression. Frontal carinae weakly developed and very fine, becoming weaker at anterior eye level, and fading out between posterior eye margin and posterior head margin. Antennal scrobes absent. Antennal scapes short, not reaching posterior head margin (SI 71–72). Eyes comparatively small to moderate (OI 23). Mesosomal outline in profile flat, weakly marginate from lateral to dorsal mesosoma, promesonotal suture and metanotal groove absent; mesosoma comparatively stout (LMI 43–45). Propodeum armed with short, triangular spines (PSLI 17). Propodeal lobes welldeveloped and broadly triangular, approximately of same height and volume as propodeal spines. Petiolar node in profile rectangular nodiform, approximately 1.3 times higher than long (LPeI 65–67), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal angles at about same height, dorsum not tapering backwards posteriorly; node in dorsal view weakly but distinctly longer than wide (DPeI 94). Postpetiole in profile rounded and approximately 1.2 times higher than long (LPpI 88), in dorsal view weakly wider than long (DPpI 105–110). Postpetiole in profile more voluminous than petiolar node, in dorsal view approximately 1.5 times wider than petiolar node (PPI 152–153). Mandibles partly with superficial, weak striation, mostly unsculptured, and general appearance smooth and shiny; clypeus with strong median longitudinal ruga and one or two very weak rugulae at each side; cephalic dorsum between frontal carinae with a pair of weakly developed, longitudinal, median rugae, area between median rugae and frontal carinae almost unsculptured, few traces of rugulae present anteriorly, median rugae of shorter than frontal carinae and ending at eye level; lateral head mostly unsculptured with weak irregular longitudinal rugulae anteriorly. Mesosoma laterally with few irregular rugae or rugulae, remainder unsculptured, mesosomal dorsum generally unsculptured with few traces of rugulae only; petiole, postpetiole, and gaster unsculptured, smooth and shining. Ground sculpture on whole body faint to absent. All dorsal surfaces of head, mesosoma, and waist segments with short, fine subdecumbent to erect pilosity, first gastral tergite with short, strongly appressed pubescence. Body uniformly brown.
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</paragraph>
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</subSubSection>
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||
<caption id="DF398389BF48FFB30AC0F83E9E68AC98" ID-DOI="http://doi.org/10.5281/zenodo.6039071" ID-Zenodo-Dep="6039071" httpUri="https://zenodo.org/record/6039071/files/figure.png" pageId="113" pageNumber="114" startId="113.[151,264,1974,1997]" targetBox="[181,1405,193,1953]" targetPageId="113">
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||
<paragraph id="8BF9D301BF48FFB30AC0F83E9E68AC98" blockId="113.[151,1435,1974,2031]" pageId="113" pageNumber="114">
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<emphasis id="B9320F13BF48FFB30AC0F83E9D2BACBA" bold="true" box="[151,368,1974,1997]" pageId="113" pageNumber="114">FIGURES 158–160.</emphasis>
|
||
<taxonomicName id="4C46A882BF48FFB30B2EF83F9D8BACBB" authorityName="Garcia & Fisher" authorityYear="2012" box="[377,464,1975,1996]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF48FFB30B2EF83F9D8BACBB" box="[377,464,1975,1996]" italics="true" pageId="113" pageNumber="114">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
, holotype—CASENT0426807 (Shannon Hartman 2011). 158. Body in profile. 159. Body in dorsal view. 160. Head in full-face view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35C808ABF4BFFB00AC0FD889D9CAF0B" pageId="114" pageNumber="115" type="discussion">
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FD889C81A96F" blockId="114.[151,1438,512,1148]" box="[151,218,512,536]" pageId="114" pageNumber="115">
|
||
<heading id="D0B1646DBF4BFFB00AC0FD889C81A96F" bold="true" box="[151,218,512,536]" fontSize="10" level="2" pageId="114" pageNumber="115" reason="2">
|
||
<emphasis id="B9320F13BF4BFFB00AC0FD889C81A96F" bold="true" box="[151,218,512,536]" pageId="114" pageNumber="115">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FDAC9963A89B" blockId="114.[151,1438,512,1148]" pageId="114" pageNumber="115">
|
||
The new species is only known from the
|
||
<typeStatus id="54FD6DA3BF4BFFB00836FDAC9ECAA94B" box="[609,657,548,572]" pageId="114" pageNumber="115">type</typeStatus>
|
||
locality, and only from
|
||
<specimenCount id="9D401888BF4BFFB009CBFDAC981FA94B" box="[924,1092,548,572]" pageId="114" pageNumber="115" type="generic">two specimens</specimenCount>
|
||
, which is similar to the limited material known for
|
||
<taxonomicName id="4C46A882BF4BFFB00BD7FDC19DB9A928" authorityName="Garcia & Fisher" authorityYear="2012" box="[384,482,584,608]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tyrion">
|
||
<emphasis id="B9320F13BF4BFFB00BD7FDC19DB9A928" box="[384,482,584,608]" italics="true" pageId="114" pageNumber="115">T. tyrion</emphasis>
|
||
</taxonomicName>
|
||
. However, in contrast to the latter, more bionomic information is available for
|
||
<taxonomicName id="4C46A882BF4BFFB00FD0FDC19C83A9F3" authorityName="Garcia & Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00FD0FDC19C83A9F3" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<specimenCount id="9D401888BF4BFFB00B4EFDE49DE4A9F3" box="[281,447,620,644]" pageId="114" pageNumber="115" type="generic">two specimens</specimenCount>
|
||
were sampled from a tropical dry forest on tsingy, and, interestingly, found in a ground nest of
|
||
<taxonomicName id="4C46A882BF4BFFB00ABCFD189F76A9D0" authority="Forel. Unfortunately" authorityName="Forel. Unfortunately" box="[235,813,655,680]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00ABCFD189E65A9DF" box="[235,574,656,680]" italics="true" pageId="114" pageNumber="115">Aphaenogaster swammerdami</emphasis>
|
||
Forel. Unfortunately
|
||
</taxonomicName>
|
||
, no additional information exists about the relationship between these two species. Nevertheless,
|
||
<taxonomicName id="4C46A882BF4BFFB0083DFD3C9F70A9BB" box="[618,811,692,716]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB0083DFD3C9F70A9BB" box="[618,811,692,716]" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
is a much larger species compared to
|
||
<taxonomicName id="4C46A882BF4BFFB00E81FD3D9968A9BB" authorityName="Garcia & Fisher" authorityYear="2012" box="[1238,1331,693,716]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00E81FD3D9968A9BB" box="[1238,1331,693,716]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
. Also,
|
||
<taxonomicName id="4C46A882BF4BFFB00FD3FD3C9D62A987" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00FD3FD3C9D62A987" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
is a very common and abundant species known to have an omnivorous diet consisting of arthropods and seeds (Boehing-Gaese
|
||
<emphasis id="B9320F13BF4BFFB00B81FD759E56A863" box="[470,525,764,788]" italics="true" pageId="114" pageNumber="115">et al</emphasis>
|
||
., 1996, 1999;
|
||
<bibRefCitation id="EFD7AEF0BF4BFFB008E9FD749F09A863" author="Fisher, B. L." box="[702,850,764,788]" pageId="114" pageNumber="115" pagination="811 - 819" refId="ref86873" refString="Fisher, B. L. (2003) Ants (Formicidae: Hymenoptera). In: S. M. Goodman & J. P. Benstead (Eds.), The natural history of Madagascar. University of Chicago Press, Chicago, pp. 811 - 819." type="book chapter" year="2003">Fisher, 2003</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFD7AEF0BF4BFFB0093CFD749853A863" author="Fisher, B. L." box="[875,1032,764,788]" pageId="114" pageNumber="115" pagination="811 - 819" refId="ref86873" refString="Fisher, B. L. (2003) Ants (Formicidae: Hymenoptera). In: S. M. Goodman & J. P. Benstead (Eds.), The natural history of Madagascar. University of Chicago Press, Chicago, pp. 811 - 819." type="book chapter" year="2003">Fisher (2003)</bibRefCitation>
|
||
also reported that the nests of
|
||
<taxonomicName id="4C46A882BF4BFFB00FD3FD749D62A84F" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00FD3FD749D62A84F" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
are famous for being shared with snakes (
|
||
<taxonomicName id="4C46A882BF4BFFB00948FCA898C7A840" authority="Gunther" authorityName="Gunther" box="[799,1180,799,824]" class="Reptilia" family="Lamprophiidae" genus="Leioheterodon" kingdom="Animalia" order="Squamata" pageId="114" pageNumber="115" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="B9320F13BF4BFFB00948FCA89869A84F" box="[799,1074,800,824]" italics="true" pageId="114" pageNumber="115">Leioheterodon modestus</emphasis>
|
||
Günther
|
||
</taxonomicName>
|
||
). The large nests of
|
||
<taxonomicName id="4C46A882BF4BFFB00FD3FCA89D62A82B" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00FD3FCA89D62A82B" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
appear to provide a variety of resources which could be used by
|
||
<taxonomicName id="4C46A882BF4BFFB00E19FCCD98F4A82B" authorityName="Garcia & Fisher" authorityYear="2012" box="[1102,1199,837,860]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00E19FCCD98F4A82B" box="[1102,1199,837,860]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
. It could be that
|
||
<taxonomicName id="4C46A882BF4BFFB00FD3FCCC9D62A8F7" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00FD3FCCC9D62A8F7" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
ignores or tolerates the much smaller
|
||
<taxonomicName id="4C46A882BF4BFFB008A2FCE19F0DA808" authorityName="Garcia & Fisher" authorityYear="2012" box="[757,854,872,896]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB008A2FCE19F0DA808" box="[757,854,872,896]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
although this is highly speculative at the moment. However, a lifestyle carried out within another ant colony could account for the scarcity of
|
||
<taxonomicName id="4C46A882BF4BFFB00E84FC05996EA8D3" authorityName="Garcia & Fisher" authorityYear="2012" box="[1235,1333,909,932]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00E84FC05996EA8D3" box="[1235,1333,909,932]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
material sampled so far. Nevertheless, more material and especially more nest collections from
|
||
<taxonomicName id="4C46A882BF4BFFB00E21FC389962A8BF" box="[1142,1337,944,968]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis id="B9320F13BF4BFFB00E21FC389962A8BF" box="[1142,1337,944,968]" italics="true" pageId="114" pageNumber="115">A. swammerdami</emphasis>
|
||
</taxonomicName>
|
||
or other ants including
|
||
<taxonomicName id="4C46A882BF4BFFB00B6BFC5D9DC1A89B" authorityName="Garcia & Fisher" authorityYear="2012" box="[316,410,981,1004]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00B6BFC5D9DC1A89B" box="[316,410,981,1004]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
are needed to come to any final conclusion about the lifestyle of the latter species.
|
||
</paragraph>
|
||
<paragraph id="8BF9D301BF4BFFB00A90FC709D9CAF0B" blockId="114.[151,1438,512,1148]" pageId="114" pageNumber="115">
|
||
<taxonomicName id="4C46A882BF4BFFB00A90FC709DF9AF78" authorityName="Garcia & Fisher" authorityYear="2012" box="[199,418,1016,1039]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00A90FC709DF9AF78" box="[199,418,1016,1039]" italics="true" pageId="114" pageNumber="115">Tetramorium tsingy</emphasis>
|
||
</taxonomicName>
|
||
is not likely to be confused with
|
||
<taxonomicName id="4C46A882BF4BFFB0094EFC719F23AF78" authorityName="Garcia & Fisher" authorityYear="2012" box="[793,888,1016,1040]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tyrion">
|
||
<emphasis id="B9320F13BF4BFFB0094EFC719F23AF78" box="[793,888,1016,1040]" italics="true" pageId="114" pageNumber="115">T. tyrion</emphasis>
|
||
</taxonomicName>
|
||
since both differ significantly in gastral pilosity/ pubescence and development of the propodeal spines/teeth. In
|
||
<taxonomicName id="4C46A882BF4BFFB00933FB959F9FAF43" authorityName="Garcia & Fisher" authorityYear="2012" box="[868,964,1053,1076]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00933FB959F9FAF43" box="[868,964,1053,1076]" italics="true" pageId="114" pageNumber="115">T. tsingy</emphasis>
|
||
</taxonomicName>
|
||
one can observe moderately short spines and strongly appressed pubescence on the first gastral tergite, whereas
|
||
<taxonomicName id="4C46A882BF4BFFB00993FBC9987FAF20" authorityName="Garcia & Fisher" authorityYear="2012" box="[964,1060,1088,1112]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tyrion">
|
||
<emphasis id="B9320F13BF4BFFB00993FBC9987FAF20" box="[964,1060,1088,1112]" italics="true" pageId="114" pageNumber="115">T. tyrion</emphasis>
|
||
</taxonomicName>
|
||
has only minute propodeal teeth and short but erect pilosity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C808ABF4BFFB00AC0FB24997DAE58" pageId="114" pageNumber="115" type="etymology">
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FB249D43AFB3" blockId="114.[151,1436,1196,1327]" box="[151,280,1196,1220]" pageId="114" pageNumber="115">
|
||
<heading id="D0B1646DBF4BFFB00AC0FB249D43AFB3" bold="true" box="[151,280,1196,1220]" fontSize="10" level="2" pageId="114" pageNumber="115" reason="2">
|
||
<emphasis id="B9320F13BF4BFFB00AC0FB249D43AFB3" bold="true" box="[151,280,1196,1220]" pageId="114" pageNumber="115">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FB47997DAE58" blockId="114.[151,1436,1196,1327]" pageId="114" pageNumber="115">
|
||
The epithet
|
||
<taxonomicName id="4C46A882BF4BFFB00B4BFB589D04AF90" authorityName="Garcia & Fisher" authorityYear="2012" box="[284,351,1232,1255]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B9320F13BF4BFFB00B4BFB589D04AF90" box="[284,351,1232,1255]" italics="true" pageId="114" pageNumber="115">tsingy</emphasis>
|
||
</taxonomicName>
|
||
is taken from the Malagasy language, and means "where one cannot walk barefoot". It is used to describe the karstic, vertical limestone formations encountered in the region. The new species appears to be endemic to this spectacular tsingy habitat. The species epithet is a noun in apposition and thus invariant.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C808ABF4BFFB00AC0FAD79C9FAE93" pageId="114" pageNumber="115" type="materials_examined">
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FAD79D22AE0F" blockId="114.[151,1436,1375,1508]" box="[151,377,1375,1400]" pageId="114" pageNumber="115">
|
||
<heading id="D0B1646DBF4BFFB00AC0FAD79D22AE0F" bold="true" box="[151,377,1375,1400]" fontSize="10" level="2" pageId="114" pageNumber="115" reason="2">
|
||
<emphasis id="B9320F13BF4BFFB00AC0FAD79D22AE0F" bold="true" box="[151,377,1375,1400]" pageId="114" pageNumber="115">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF9D301BF4BFFB00AC0FA0B9C9FAE93" blockId="114.[151,1436,1375,1508]" pageId="114" pageNumber="115">
|
||
<emphasis id="B9320F13BF4BFFB00AC0FA0B9D3FAEEB" bold="true" box="[151,356,1411,1436]" pageId="114" pageNumber="115">
|
||
<collectingCountry id="F3519391BF4BFFB00AC0FA0B9D3FAEEB" box="[151,356,1411,1436]" name="Madagascar" pageId="114" pageNumber="115">MADAGASCAR</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="49821DE3BF4BFFB00B2FFA0C9DACAEEB" box="[376,503,1412,1436]" country="Madagascar" name="Mahajanga" pageId="114" pageNumber="115">Mahajanga</collectingRegion>
|
||
, Parc National Tsingy de Bemaraha,
|
||
<quantity id="4CBE7EE4BF4BFFB009E2FA0C985CAEEB" box="[949,1031,1412,1436]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="114" pageNumber="115" unit="km" value="2.5">2.5 km</quantity>
|
||
62° ENE Bekopaka, Ankidrodroa River,
|
||
<geoCoordinate id="EE72B5C6BF4BFFB00AB4FA2F9D3DAEC8" box="[227,358,1447,1471]" degrees="19.13222" direction="south" orientation="latitude" pageId="114" pageNumber="115" precision="1" value="-19.13222">19.13222 S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE72B5C6BF4BFFB00B24FA2F9DA3AEC8" box="[371,504,1447,1471]" degrees="44.81467" direction="east" orientation="longitude" pageId="114" pageNumber="115" precision="1" value="44.81467">44.81467 E</geoCoordinate>
|
||
,
|
||
<quantity id="4CBE7EE4BF4BFFB00852FA2F9E0BAEC8" box="[517,592,1447,1471]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="114" pageNumber="115" unit="m" value="100.0">100 m</quantity>
|
||
, tropical dry forest on tsingy,
|
||
<date id="FFF8F5C1BF4BFFB009F0FA2F980DAEC8" box="[935,1110,1447,1471]" pageId="114" pageNumber="115" value="2001-11-11" valueMax="2001-11-15" valueMin="2001-11-11">11.–15.XI.2001</date>
|
||
(
|
||
<emphasis id="B9320F13BF4BFFB00E3FFA219CEFAE93" italics="true" pageId="114" pageNumber="115">B.L. Fisher, C. Griswold et al.</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |