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<document id="9D2E739EBE7E5031D9280A6C2F876710" ID-CLB-Dataset="2946" ID-DOI="10.11646/zootaxa.5082.1.3" ID-GBIF-Dataset="c601c282-b593-47ff-bd7f-0b901cbf7b6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783154" ID-ZooBank="06372651-6702-4741-AA55-2B302F34D21F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564268802" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="03EF120EFFBFBB5F098DB8B0FA040494" docLanguage="en" docName="zootaxa.5082.1.3.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudholophylla Blackburn 1911" docType="treatment" docVersion="5" lastPageNumber="39" masterDocId="FFD66A76FFBEBB56091ABB22FF980629" masterDocTitle="Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): a sixth species of Pseudholophylla Blackburn, 1911, and notes on the other known species" masterLastPageNumber="40" masterPageNumber="30" pageNumber="31" updateTime="1699472537074" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B0CE02A22627A76EF8D225F1F698D2A0">Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): a sixth species of Pseudholophylla Blackburn, 1911, and notes on the other known species</mods:title>
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<mods:namePart id="87DC6735BEE5331DA36152A4C8B56BAA">Hutchinson, Paul M.</mods:namePart>
<mods:affiliation id="6D232DE661FA19A53ECBD327340070F1">Department of Primary Industries and Regional Development WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia.</mods:affiliation>
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<mods:namePart id="6C61A6EFDEF04B708CD3C6095FB91FAB">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="0CD0968348FD21BE0A72DC967DEB5E39">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:identifier id="C0C8AB635FD47D0B9D0F4E789D9B2765" type="DOI">10.11646/zootaxa.5082.1.3</mods:identifier>
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<treatment id="03EF120EFFBFBB5F098DB8B0FA040494" ID-DOI="http://doi.org/10.5281/zenodo.5783146" ID-GBIF-Taxon="191647339" ID-Zenodo-Dep="5783146" LSID="urn:lsid:plazi:treatment:03EF120EFFBFBB5F098DB8B0FA040494" httpUri="http://treatment.plazi.org/id/03EF120EFFBFBB5F098DB8B0FA040494" lastPageId="9" lastPageNumber="39" pageId="1" pageNumber="31">
<subSubSection id="C35CF093FFBFBB57098DB8B0FDAE0585" box="[151,566,914,941]" pageId="1" pageNumber="31" type="nomenclature">
<paragraph id="8BF9A318FFBFBB57098DB8B0FDAE0585" blockId="1.[151,566,914,941]" box="[151,566,914,941]" pageId="1" pageNumber="31">
<heading id="D0B11474FFBFBB57098DB8B0FDAE0585" bold="true" box="[151,566,914,941]" fontSize="11" level="1" pageId="1" pageNumber="31" reason="1">
<taxonomicName id="4C46D89BFFBFBB57098DB8B0FDAE0585" authority="Blackburn, 1911" authorityName="Blackburn" authorityYear="1911" box="[151,566,914,941]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB57098DB8B0FDAE0585" bold="true" box="[151,566,914,941]" pageId="1" pageNumber="31">
<emphasis id="B9327F0AFFBFBB57098DB8B0FEF90584" bold="true" box="[151,353,914,941]" italics="true" pageId="1" pageNumber="31">Pseudholophylla</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570873B8B1FDAE0585" author="Blackburn, T." box="[361,566,914,941]" pageId="1" pageNumber="31" pagination="173 - 203" refId="ref5686" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn, 1911</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBFBB57098DB8F5FDFE0243" pageId="1" pageNumber="31" type="reference_group">
<paragraph id="8BF9A318FFBFBB57098DB8F5FE710227" blockId="1.[151,1436,983,1130]" pageId="1" pageNumber="31">
<treatmentCitationGroup id="AB568436FFBFBB57098DB8F5FE710227" pageId="1" pageNumber="31">
<treatmentCitation id="0AE78509FFBFBB57098DB8F5FE6A05C6" author="Burmeister, H. C. C." box="[151,498,983,1007]" page="426" pageId="1" pageNumber="31" year="1855">
<taxonomicName id="4C46D89BFFBFBB57098DB8F5FE6A05C6" authority="Burmeister, 1855: 426" authorityName="Burmeister" authorityPageNumber="426" authorityYear="1855" box="[151,498,983,1007]" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB57098DB8F5FE9105C6" box="[151,265,983,1007]" italics="true" pageId="1" pageNumber="31">Holophylla</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB57080AB8F5FE6A05C6" author="Burmeister, H. C. C." box="[272,498,983,1007]" pageId="1" pageNumber="31" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855: 426</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(junior homonym of
<taxonomicName id="4C46D89BFFBFBB570BD4B8F5FC7C05C6" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[718,996,983,1007]" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB570BD4B8F5FCD805C6" box="[718,832,983,1007]" italics="true" pageId="1" pageNumber="31">Holophylla</emphasis>
Erichson, 1847
</taxonomicName>
).
<typeStatus id="54FD1DBAFFBFBB570AEFB8FAFBBF05D9" box="[1013,1063,984,1008]" pageId="1" pageNumber="31">Type</typeStatus>
species:
<taxonomicName id="4C46D89BFFBFBB570D9EB8F5FECA0227" authority="Burmeister, 1855" authorityName="Burmeister" authorityYear="1855" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFBFBB570D9EB8F5FAF005C6" box="[1156,1384,983,1007]" italics="true" pageId="1" pageNumber="31">Holophylla furfuracea</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570C74B8FAFECA0227" author="Burmeister, H. C. C." pageId="1" pageNumber="31" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
</taxonomicName>
, by monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF9A318FFBFBB57098DBF36FBD40205" blockId="1.[151,1436,983,1130]" box="[151,1100,1044,1069]" pageId="1" pageNumber="31">
<treatmentCitationGroup id="AB568436FFBFBB57098DBF36FBD40205" box="[151,1100,1044,1069]" pageId="1" pageNumber="31">
<treatmentCitation id="0AE78509FFBFBB57098DBF36FDBB0204" author="Blackburn, T." box="[151,547,1044,1069]" page="183" pageId="1" pageNumber="31" year="1911">
<taxonomicName id="4C46D89BFFBFBB57098DBF36FDBB0204" authority="Blackburn, 1911: 183" authorityName="Blackburn" authorityPageNumber="183" authorityYear="1911" box="[151,547,1044,1069]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB57098DBF36FED90205" box="[151,321,1044,1068]" italics="true" pageId="1" pageNumber="31">Pseudholophylla</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570852BF36FDBB0204" author="Blackburn, T." box="[328,547,1044,1069]" pageId="1" pageNumber="31" pagination="173 - 203" refId="ref5686" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn, 1911: 183</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(replacement name for
<taxonomicName id="4C46D89BFFBFBB570A0FBF36FBD80205" authority="Burmeister, 1855" authorityName="Burmeister" authorityYear="1855" box="[789,1088,1044,1068]" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB570A0FBF36FC1F0205" box="[789,903,1044,1068]" italics="true" pageId="1" pageNumber="31">Holophylla</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570A97BF36FBD80205" author="Burmeister, H. C. C." box="[909,1088,1044,1068]" pageId="1" pageNumber="31" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF9A318FFBFBB57098DBF11FDFE0243" blockId="1.[151,1436,983,1130]" pageId="1" pageNumber="31">
<treatmentCitationGroup id="AB568436FFBFBB57098DBF11FDFE0243" pageId="1" pageNumber="31">
<treatmentCitation id="0AE78509FFBFBB57098DBF11FD9E0262" author="Blackburn, T." box="[151,518,1075,1099]" page="196" pageId="1" pageNumber="31" year="1911">
<taxonomicName id="4C46D89BFFBFBB57098DBF11FD9E0262" authority="Blackburn, 1911: 196" authorityName="Blackburn" authorityPageNumber="196" authorityYear="1911" box="[151,518,1075,1099]" class="Insecta" family="Melolonthidae" genus="Paralepidiota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBFBB57098DBF11FEBC0262" box="[151,292,1075,1099]" italics="true" pageId="1" pageNumber="31">Paralepidiota</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570833BF11FD9E0262" author="Blackburn, T." box="[297,518,1075,1099]" pageId="1" pageNumber="31" pagination="173 - 203" refId="ref5686" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn, 1911: 196</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AE78509FFBFBB570B0ABF11FD2D0262" author="Britton, E. B." box="[528,693,1075,1099]" page="33" pageId="1" pageNumber="31" year="1978">
<bibRefCitation id="EFD7DEE9FFBFBB570B0ABF11FD2D0262" author="Britton, E. B." box="[528,693,1075,1099]" pageId="1" pageNumber="31" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton 1978: 33</bibRefCitation>
</treatmentCitation>
(synonym).
<typeStatus id="54FD1DBAFFBFBB570A2FBF16FCFF0265" box="[821,871,1076,1100]" pageId="1" pageNumber="31">Type</typeStatus>
species:
<taxonomicName id="4C46D89BFFBFBB570AD8BF11FAE10262" ID-CoL="LP" authority="Blackburn, 1912" authorityName="Blackburn" authorityYear="1912" box="[962,1401,1075,1099]" class="Insecta" family="Melolonthidae" genus="Paralepidiota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="species" species="lepidoptera">
<emphasis id="B9327F0AFFBFBB570AD8BF11FB520262" box="[962,1226,1075,1099]" italics="true" pageId="1" pageNumber="31">Paralepidiota lepidoptera</emphasis>
<bibRefCitation id="EFD7DEE9FFBFBB570DD5BF11FAE10262" author="Blackburn, T." box="[1231,1401,1075,1099]" pageId="1" pageNumber="31" pagination="40 - 75" refId="ref5726" refString="Blackburn, T. (1912) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLII. Transactions of the Royal Society of South Australia, 36, 40 - 75." type="journal article" year="1912">Blackburn, 1912</bibRefCitation>
</taxonomicName>
, by subsequent monotypy (
<bibRefCitation id="EFD7DEE9FFBFBB5708A8BF70FDC20243" author="Blackburn, T." box="[434,602,1106,1130]" pageId="1" pageNumber="31" pagination="40 - 75" refId="ref5726" refString="Blackburn, T. (1912) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLII. Transactions of the Royal Society of South Australia, 36, 40 - 75." type="journal article" year="1912">Blackburn 1912</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBFBB57098DBFBAFE0603A3" pageId="1" pageNumber="31" type="diagnosis">
<paragraph id="8BF9A318FFBFBB57098DBFBAFE0603A3" blockId="1.[151,1437,1176,1419]" pageId="1" pageNumber="31">
<emphasis id="B9327F0AFFBFBB57098DBFBAFE8A029B" bold="true" box="[151,274,1176,1202]" pageId="1" pageNumber="31">Diagnosis.</emphasis>
Dorsal surface, pronotal hypomera, and anterior faces of profemora and protibiae bearing elliptical or elongate-pointed white scales; surface of clypeus, pronotum, and scutellum without long, backwardly directed setae; labrum deeply excavated anteriorly; antennae of males with antennomeres 410 or 510 lamellate; clypeus with anterior face moderately deep, greatest width 5.57.5 times mid depth, outline broadly rounded, usually slightly depressed in middle of anterior edge; upper surface of clypeus strongly concave; protarsal claws with a prominent tooth; anterior edge of metafemur not concave near base; lower spur of metatibiae narrow, thick and tapered, upper spur broadly expanded.
</paragraph>
</subSubSection>
<caption id="DF39F390FFBCBB54098DBDC4FE8C010A" ID-DOI="http://doi.org/10.5281/zenodo.5783156" ID-Zenodo-Dep="5783156" httpUri="https://zenodo.org/record/5783156/files/figure.png" pageId="2" pageNumber="32" startId="2.[151,264,1766,1790]" targetBox="[181,1409,213,1742]" targetPageId="2">
<paragraph id="8BF9A318FFBCBB54098DBDC4FE8C010A" blockId="2.[151,1436,1766,1827]" pageId="2" pageNumber="32">
<emphasis id="B9327F0AFFBCBB54098DBDC4FEA200D7" bold="true" box="[151,314,1766,1790]" pageId="2" pageNumber="32">FIGURES 13.</emphasis>
<taxonomicName id="4C46D89BFFBCBB54085ABDC5FC8C00D6" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[320,788,1767,1791]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="hurai" status="sp. nov.">
<emphasis id="B9327F0AFFBCBB54085ABDC5FDB000D6" box="[320,552,1767,1791]" italics="true" pageId="2" pageNumber="32">Pseudholophylla hurai</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<emphasis id="B9327F0AFFBCBB540A04BDC5FC0300D7" bold="true" box="[798,923,1766,1791]" pageId="2" pageNumber="32">
<taxonomicNameLabel id="A201C271FFBCBB540A04BDC5FC0300D7" box="[798,923,1766,1791]" pageId="2" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B9327F0AFFBCBB540D5EBDC4FBC900D7" bold="true" box="[1092,1105,1766,1790]" pageId="2" pageNumber="32">1</emphasis>
, Dorsal view;
<emphasis id="B9327F0AFFBCBB540DFFBDC4FB6A00D7" bold="true" box="[1253,1266,1766,1790]" pageId="2" pageNumber="32">2</emphasis>
, ventral view;
<emphasis id="B9327F0AFFBCBB540C93BDC4FA0E00D7" bold="true" box="[1417,1430,1766,1790]" pageId="2" pageNumber="32">3</emphasis>
, lateral view.
</paragraph>
</caption>
<caption id="DF39F390FFBDBB55098DBC1BFC92015C" ID-DOI="http://doi.org/10.5281/zenodo.5783158" ID-Zenodo-Dep="5783158" httpUri="https://zenodo.org/record/5783158/files/figure.png" pageId="3" pageNumber="33" startId="3.[151,264,1849,1873]" targetBox="[151,1436,835,1807]" targetPageId="3">
<paragraph id="8BF9A318FFBDBB55098DBC1BFC92015C" blockId="3.[151,1436,1849,1909]" pageId="3" pageNumber="33">
<emphasis id="B9327F0AFFBDBB55098DBC1BFEA30178" bold="true" box="[151,315,1849,1873]" pageId="3" pageNumber="33">FIGURES 47.</emphasis>
<taxonomicName id="4C46D89BFFBDBB550858BC1BFC810178" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[322,793,1849,1873]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="hurai" status="sp. nov.">
<emphasis id="B9327F0AFFBDBB550858BC1BFDB20178" box="[322,554,1849,1873]" italics="true" pageId="3" pageNumber="33">Pseudholophylla hurai</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<emphasis id="B9327F0AFFBDBB550A3EBC1BFC390178" bold="true" box="[804,929,1849,1873]" pageId="3" pageNumber="33">
<taxonomicNameLabel id="A201C271FFBDBB550A3EBC1BFC390178" box="[804,929,1849,1873]" pageId="3" pageNumber="33" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B9327F0AFFBDBB550D56BC1BFBC10178" bold="true" box="[1100,1113,1849,1873]" pageId="3" pageNumber="33">4</emphasis>
, Frontal view;
<emphasis id="B9327F0AFFBDBB550DECBC1BFA9B0178" bold="true" box="[1270,1283,1849,1873]" pageId="3" pageNumber="33">5</emphasis>
, head and pronotum, dorso-lateral view;
<emphasis id="B9327F0AFFBDBB5508B0BC7FFE2F015C" bold="true" box="[426,439,1885,1909]" pageId="3" pageNumber="33">6</emphasis>
, posterior view;
<emphasis id="B9327F0AFFBDBB550B78BC7FFDF7015C" bold="true" box="[610,623,1885,1909]" pageId="3" pageNumber="33">7</emphasis>
, antennal club.
</paragraph>
</caption>
<caption id="DF39F390FFBABB52098DBDDBFCC70154" ID-DOI="http://doi.org/10.5281/zenodo.5783160" ID-Zenodo-Dep="5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="4" pageNumber="34" startId="4.[151,264,1785,1809]" targetBox="[168,1426,702,1754]" targetPageId="4">
<paragraph id="8BF9A318FFBABB52098DBDDBFCC70154" blockId="4.[151,1437,1785,1917]" pageId="4" pageNumber="34">
<emphasis id="B9327F0AFFBABB52098DBDDBFED00138" bold="true" box="[151,328,1785,1809]" pageId="4" pageNumber="34">FIGURES 813.</emphasis>
<taxonomicName id="4C46D89BFFBABB520854BDDBFE600138" authorityName="Blackburn" authorityYear="1911" box="[334,504,1785,1809]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B9327F0AFFBABB520854BDDBFE600138" box="[334,504,1785,1809]" italics="true" pageId="4" pageNumber="34">Pseudholophylla</emphasis>
</taxonomicName>
aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right.
<emphasis id="B9327F0AFFBABB520C93BDDBFA0E0138" bold="true" box="[1417,1430,1785,1809]" pageId="4" pageNumber="34">8</emphasis>
,
<taxonomicName id="4C46D89BFFBABB52098DBC3FFD16011C" authority="(Macleay, 1888)" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[151,654,1821,1845]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B9327F0AFFBABB52098DBC3FFE47011C" box="[151,479,1821,1845]" italics="true" pageId="4" pageNumber="34">Pseudholophylla castaneipennis</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBABB5208F6BC3FFD1E011C" author="Macleay, W." box="[492,646,1821,1845]" pageId="4" pageNumber="34" pagination="897 - 924" refId="ref6071" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. Part II. The lamellicornes. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 897 - 924. https: // doi. org / 10.5962 / bhl. part. 29226" type="journal article" year="1888">Macleay, 1888</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B9327F0AFFBABB520B82BC3FFD3D011C" bold="true" box="[664,677,1821,1845]" pageId="4" pageNumber="34">9</emphasis>
,
<taxonomicName id="4C46D89BFFBABB520BA8BC3CFC30011C" authority="(Lea, 1919)" baseAuthorityName="Lea" baseAuthorityYear="1919" box="[690,936,1821,1845]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="B9327F0AFFBABB520BA8BC3CFCB0011C" box="[690,808,1821,1845]" italics="true" pageId="4" pageNumber="34">P. cavifrons</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBABB520A2CBC3FFC38011C" author="Lea, A. M." box="[822,928,1821,1845]" pageId="4" pageNumber="34" pagination="166 - 261" refId="ref6033" refString="Lea, A. M. (1919) Notes on some miscellaneous Coleoptera with descriptions of new species. Part V. Transactions of the Royal Society of South Australia, 43, 166 - 261." type="journal article" year="1919">Lea, 1919</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B9327F0AFFBABB520AA8BC3FFC53011C" bold="true" box="[946,971,1821,1845]" pageId="4" pageNumber="34">10</emphasis>
,
<taxonomicName id="4C46D89BFFBABB520AC2BC3CFABE011C" authority="(Burmeister, 1855)" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[984,1318,1821,1845]" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFBABB520AC2BC3CFBC4011C" box="[984,1116,1821,1845]" italics="true" pageId="4" pageNumber="34">P. furfuracea</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBABB520D70BC3FFA85011C" author="Burmeister, H. C. C." box="[1130,1309,1821,1845]" pageId="4" pageNumber="34" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B9327F0AFFBABB520C2ABC3FFAD0011C" bold="true" box="[1328,1352,1821,1845]" pageId="4" pageNumber="34">11</emphasis>
,
<taxonomicName id="4C46D89BFFBABB520C4FBC3CFE340170" authority="(Blackburn, 1912)" baseAuthorityName="Blackburn" baseAuthorityYear="1912" class="Insecta" family="Melolonthidae" genus="Paralepidiota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="lepidoptera">
<emphasis id="B9327F0AFFBABB520C4FBC3CFF7E0170" italics="true" pageId="4" pageNumber="34">P. lepidoptera</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBABB5209EEBC63FE3C0170" author="Blackburn, T." box="[244,420,1857,1881]" pageId="4" pageNumber="34" pagination="40 - 75" refId="ref5726" refString="Blackburn, T. (1912) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLII. Transactions of the Royal Society of South Australia, 36, 40 - 75." type="journal article" year="1912">Blackburn, 1912</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B9327F0AFFBABB5208ADBC63FE480170" bold="true" box="[439,464,1857,1881]" pageId="4" pageNumber="34">12</emphasis>
,
<taxonomicName id="4C46D89BFFBABB5208C4BC60FD590170" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" box="[478,705,1857,1881]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="B9327F0AFFBABB5208C4BC60FDB60170" box="[478,558,1858,1881]" italics="true" pageId="4" pageNumber="34">P. soror</emphasis>
<bibRefCitation id="EFD7DEE9FFBABB520B2FBC63FD590170" author="Britton, E. B." box="[565,705,1857,1881]" pageId="4" pageNumber="34" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
</taxonomicName>
;
<emphasis id="B9327F0AFFBABB520BD6BC63FD7D0170" bold="true" box="[716,741,1857,1881]" pageId="4" pageNumber="34">13</emphasis>
,
<taxonomicName id="4C46D89BFFBABB520BE9BC60FBAD0170" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[755,1077,1857,1881]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="hurai" status="sp. nov.">
<emphasis id="B9327F0AFFBABB520BE9BC60FCDC0170" box="[755,836,1857,1881]" italics="true" pageId="4" pageNumber="34">P. hurai</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<emphasis id="B9327F0AFFBABB520D5ABC63FB260170" bold="true" box="[1088,1214,1857,1881]" pageId="4" pageNumber="34">
<taxonomicNameLabel id="A201C271FFBABB520D5ABC63FB260170" box="[1088,1214,1857,1881]" pageId="4" pageNumber="34" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype male. Figs. 812 reproduced from
<bibRefCitation id="EFD7DEE9FFBABB52089ABC47FD890154" author="Britton, E. B." box="[384,529,1893,1917]" pageId="4" pageNumber="34" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
(CSIRO, used with permission).
</paragraph>
</caption>
<caption id="DF39F390FFBBBB53098DB8CCFADE022F" ID-DOI="http://doi.org/10.5281/zenodo.5783162" ID-Zenodo-Dep="5783162" box="[151,1350,1006,1031]" httpUri="https://zenodo.org/record/5783162/files/figure.png" pageId="5" pageNumber="35" startId="5.[151,250,1006,1030]" targetBox="[151,1436,237,983]" targetPageId="5">
<paragraph id="8BF9A318FFBBBB53098DB8CCFADE022F" blockId="5.[151,1350,1006,1031]" box="[151,1350,1006,1031]" pageId="5" pageNumber="35">
<emphasis id="B9327F0AFFBBBB53098DB8CCFEB8022F" bold="true" box="[151,288,1006,1030]" pageId="5" pageNumber="35">FIGURE 14.</emphasis>
Type locality of
<taxonomicName id="4C46D89BFFBBBB5308D6B8CCFC39022F" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[460,929,1006,1031]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="hurai" status="sp. nov.">
<emphasis id="B9327F0AFFBBBB5308D6B8CCFD2C022F" box="[460,692,1006,1030]" italics="true" pageId="5" pageNumber="35">Pseudholophylla hurai</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<emphasis id="B9327F0AFFBBBB530AB1B8CDFBB0022F" bold="true" box="[939,1064,1006,1031]" pageId="5" pageNumber="35">
<taxonomicNameLabel id="A201C271FFBBBB530AB1B8CDFBB0022F" box="[939,1064,1006,1031]" pageId="5" pageNumber="35" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Photograph by Mark Hura.
</paragraph>
</caption>
<caption id="DF39F390FFB8BB50098DBC56FBA00199" ID-DOI="http://doi.org/10.5281/zenodo.5783164" ID-Zenodo-Dep="5783164" httpUri="https://zenodo.org/record/5783164/files/figure.png" pageId="6" pageNumber="36" startId="6.[151,264,1908,1932]" targetBox="[166,1435,1036,1863]" targetPageId="6">
<paragraph id="8BF9A318FFB8BB50098DBC56FBA00199" blockId="6.[151,1437,1908,1968]" pageId="6" pageNumber="36">
<emphasis id="B9327F0AFFB8BB50098DBC56FECC01A5" bold="true" box="[151,340,1908,1932]" pageId="6" pageNumber="36">FIGURES 1516.</emphasis>
<taxonomicName id="4C46D89BFFB8BB500841BC56FC9D01A5" authority="(Blackburn)" baseAuthorityName="Blackburn" baseAuthorityYear="1912" box="[347,773,1908,1932]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="lepidoptera">
<emphasis id="B9327F0AFFB8BB500841BC56FD1901A5" box="[347,641,1908,1932]" italics="true" pageId="6" pageNumber="36">Pseudholophylla lepidoptera</emphasis>
(Blackburn)
</taxonomicName>
, female.
<emphasis id="B9327F0AFFB8BB500A7BBC56FCE201A5" bold="true" box="[865,890,1908,1932]" pageId="6" pageNumber="36">13</emphasis>
, Dorsal view;
<emphasis id="B9327F0AFFB8BB500D0BBC56FBB201A5" bold="true" box="[1041,1066,1908,1932]" pageId="6" pageNumber="36">14</emphasis>
, antenna and maxillary palp. Photographs from the Queensland Museum as QMDIU_03317 and QMDIU_03318, respectively.
</paragraph>
</caption>
<caption id="DF39F390FFB6BB5E098DBEF8FD210013" ID-DOI="http://doi.org/10.5281/zenodo.5783166" ID-Zenodo-Dep="5783166" httpUri="https://zenodo.org/record/5783166/files/figure.png" pageId="8" pageNumber="38" startId="8.[151,250,1498,1522]" targetBox="[151,1436,345,1474]" targetPageId="8">
<paragraph id="8BF9A318FFB6BB5E098DBEF8FD210013" blockId="8.[151,1436,1498,1594]" pageId="8" pageNumber="38">
<emphasis id="B9327F0AFFB6BB5E098DBEF8FEB903DB" bold="true" box="[151,289,1498,1522]" pageId="8" pageNumber="38">FIGURE 17.</emphasis>
Known distributions of
<taxonomicName id="4C46D89BFFB6BB5E0B05BEF8FC8203DB" box="[543,794,1498,1522]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9327F0AFFB6BB5E0B05BEF8FD5103DB" box="[543,713,1498,1522]" italics="true" pageId="8" pageNumber="38">Pseudholophylla</emphasis>
species
</taxonomicName>
across northern Australia. Upper map shows distribution relative to topography; lower map shows Köppen-Geiger classifications for comparison with climates.
<taxonomicName id="4C46D89BFFB6BB5E0D9BBEDCFECF0013" authority="(Burmeister, 1855)" baseAuthorityName="Burmeister" baseAuthorityYear="1855" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="38" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFB6BB5E0D9BBEDCFA04003F" box="[1153,1436,1534,1558]" italics="true" pageId="8" pageNumber="38">Pseudholophylla furfuracea</emphasis>
(
<bibRefCitation id="EFD7DEE9FFB6BB5E0984BD00FEC80013" author="Burmeister, H. C. C." box="[158,336,1570,1594]" pageId="8" pageNumber="38" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
is recorded only from “Australia”.
</paragraph>
</caption>
<subSection id="E2C9BAF3FFB6BB5F098DBDAEFA040494" lastPageId="9" lastPageNumber="39" pageId="8" pageNumber="38" type="key">
<subSubSection id="C35CF093FFB6BB5E098DBDAEFD24008F" box="[151,700,1676,1702]" pageId="8" pageNumber="38" type="description">
<paragraph id="8BF9A318FFB6BB5E098DBDAEFD24008F" blockId="8.[151,700,1676,1702]" box="[151,700,1676,1702]" pageId="8" pageNumber="38">
<heading id="D0B11474FFB6BB5E098DBDAEFD24008F" bold="true" box="[151,700,1676,1702]" fontSize="11" level="1" pageId="8" pageNumber="38" reason="1">
<emphasis id="B9327F0AFFB6BB5E098DBDAEFD24008F" bold="true" box="[151,700,1676,1702]" pageId="8" pageNumber="38">Key to males (females are largely unknown)</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFB6BB5F098DBDF3FA040494" lastPageId="9" lastPageNumber="39" pageId="8" pageNumber="38" type="key">
<key id="FFD7F429FFB6BB5F098DBDF3FA040494" lastPageId="9" lastPageNumber="39" pageId="8" pageNumber="38">
<keyStep id="30B2BBBDFFB6BB5E098DBDF3FA04012D" pageId="8" pageNumber="38">
<paragraph id="8BF9A318FFB6BB5E098DBDF3FA0400CF" blockId="8.[151,1437,1745,2031]" box="[151,1436,1745,1767]" pageId="8" pageNumber="38">
<keyLead id="30B7002DFFB6BB5E098DBDF3FA0400CF" box="[151,1436,1745,1767]" pageId="8" pageNumber="38">
1 Each paramere with a transverse pre-apical process (
<figureCitation id="137DBF9DFFB6BB5E0BC1BDF3FCD400CE" box="[731,844,1745,1767]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="8" pageNumber="38">Figs. 11, 13</figureCitation>
)............................................... 2
</keyLead>
</paragraph>
<paragraph id="8BF9A318FFB6BB5E098DBDCDFA04012D" blockId="8.[151,1437,1745,2031]" box="[151,1436,1774,1797]" pageId="8" pageNumber="38">
<keyLead id="30B7002DFFB6BB5E098DBDCDFA04012D" box="[151,1436,1774,1797]" pageId="8" pageNumber="38">
- Each paramere without a transverse pre-apical process (
<figureCitation id="137DBF9DFFB6BB5E0BE3BDCCFC1D012C" box="[761,901,1774,1797]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="8" pageNumber="38">Figs. 810, 12</figureCitation>
).......................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B2BBBDFFB6BB5F098DBC2EFA040687" lastPageId="9" lastPageNumber="39" pageId="8" pageNumber="38">
<paragraph id="8BF9A318FFB6BB5E098DBC2EFA05019D" blockId="8.[151,1437,1745,2031]" pageId="8" pageNumber="38">
<keyLead id="30B7002DFFB6BB5E098DBC2EFA05019D" pageId="8" pageNumber="38">
2 Antenna with the lamella of antennomere 4 approximately two-fifths to half length of lamella of antennomere 5; scales on head, pronotum, elytra, metepisternum, abdominal ventrites, and legs elongate and acutely pointed; scales on anterior surface of profemora and pronotal hypomera very elongate to setae-like; sternum, metepisternum, metepimeron, and metacoxae densely clothed with long, pale setae; pygidium with sparse, evenly distributed white microsetae; body colour yellow brown, length approximately 27.430.0 mm (
<figureCitation id="137DBF9DFFB6BB5E0B11BCA3FDF001BE" box="[523,616,1921,1943]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 47" captionStartId-0="2.[151,264,1766,1790]" captionStartId-1="3.[151,264,1849,1873]" captionTargetBox-0="[181,1409,213,1742]" captionTargetBox-1="[151,1436,835,1807]" captionTargetId-0="figure-19@2.[151,1435,181,1742]" captionTargetId-1="figure-298@3.[151,1436,835,1807]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 13. Pseudholophylla hurai Hutchinson &amp; Allsopp, new species, holotype male. 1, Dorsal view; 2, ventral view; 3, lateral view." captionText-1="FIGURES 47. Pseudholophylla hurai Hutchinson &amp; Allsopp, new species, holotype male. 4, Frontal view; 5, head and pronotum, dorso-lateral view; 6, posterior view; 7, antennal club." figureDoi-0="http://doi.org/10.5281/zenodo.5783156" figureDoi-1="http://doi.org/10.5281/zenodo.5783158" httpUri-0="https://zenodo.org/record/5783156/files/figure.png" httpUri-1="https://zenodo.org/record/5783158/files/figure.png" pageId="8" pageNumber="38">Figs. 17</figureCitation>
); parameres with apices rounded and recurved, as in
<figureCitation id="137DBF9DFFB6BB5E0D7FBCA3FB3301BF" box="[1125,1195,1921,1943]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="8" pageNumber="38">Fig. 13</figureCitation>
;
<collectingRegion id="49826DFAFFB6BB5E0DACBCA3FAFF01BE" box="[1206,1383,1921,1943]" country="Australia" name="Northern Territory" pageId="8" pageNumber="38">Northern Territory</collectingRegion>
.......................................................
<taxonomicName id="4C46D89BFFB6BB5E0A7BBCBCFA84019D" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[865,1308,1950,1972]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="38" phylum="Arthropoda" rank="species" species="hurai" status="sp. nov.">
<emphasis id="B9327F0AFFB6BB5E0A7BBCBCFBA4019A" box="[865,1084,1950,1971]" italics="true" pageId="8" pageNumber="38">Pseudholophylla hurai</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
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<taxonomicNameLabel id="A201C271FFB6BB5E0C3CBCBDFA05019D" box="[1318,1437,1950,1972]" pageId="8" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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- Antenna with antennomere 4 angulate but not lamellate, antennal club of 6 nearly equal lamellae; dorsal surface with sparse, elliptical white scales, abdominal ventrites with dense white scales; body colour reddish brown, length approximately
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; parameres with apices elongate, as in
<figureCitation id="137DBF9DFFB7BB5F0B57BBB5FD0A0684" box="[589,658,151,173]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="9" pageNumber="39">Fig. 11</figureCitation>
; northeastern
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......
<taxonomicName id="4C46D89BFFB7BB5F0ACABBB5FA040687" ID-CoL="LP" authority="(Blackburn, 1912)" baseAuthorityName="Blackburn" baseAuthorityYear="1912" box="[976,1436,151,174]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="lepidoptera">
<emphasis id="B9327F0AFFB7BB5F0ACABBB5FB7F0685" box="[976,1255,151,172]" italics="true" pageId="9" pageNumber="39">Pseudholophylla lepidoptera</emphasis>
(
<bibRefCitation id="EFD7DEE9FFB7BB5F0DE9BBB5FA0D0687" author="Blackburn, T." box="[1267,1429,151,174]" pageId="9" pageNumber="39" pagination="40 - 75" refId="ref5726" refString="Blackburn, T. (1912) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLII. Transactions of the Royal Society of South Australia, 36, 40 - 75." type="journal article" year="1912">Blackburn, 1912</bibRefCitation>
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3 Antenna with lamella of antennomere 4 approximately one-third as long as lamella of antennomere 5; terminal palpomere of maxillary palp with flat elliptical area on upper surface; aedeagus as in
<figureCitation id="137DBF9DFFB7BB5F0A95BBF0FC5106CE" box="[911,969,210,232]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="9" pageNumber="39">Fig. 8</figureCitation>
; northwestern
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........................................................................
<taxonomicName id="4C46D89BFFB7BB5F0AD9BBCDFA04072F" authority="(Macleay, 1888)" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[963,1436,239,262]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B9327F0AFFB7BB5F0AD9BBCDFB62072C" box="[963,1274,239,261]" italics="true" pageId="9" pageNumber="39">Pseudholophylla castaneipennis</emphasis>
(
<bibRefCitation id="EFD7DEE9FFB7BB5F0C1CBBCDFA0D072F" author="Macleay, W." box="[1286,1429,239,262]" pageId="9" pageNumber="39" pagination="897 - 924" refId="ref6071" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. Part II. The lamellicornes. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 897 - 924. https: // doi. org / 10.5962 / bhl. part. 29226" type="journal article" year="1888">Macleay, 1888</bibRefCitation>
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- Antenna with lamella of antennomere 4 more than half as long as lamella of antennomere 5; terminal palpomere of maxillary palp with an elliptical concavity of the upper surface; northern
<collectingCountry id="F351E388FFB7BB5F0A24BA08FC0E0769" box="[830,918,298,320]" name="Australia" pageId="9" pageNumber="39">Australia</collectingCountry>
.......................................... 4
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<keyLead id="30B7002DFFB7BB5F098DBA6AFA0407FA" pageId="9" pageNumber="39">
4 Scales on frons and pronotum broadly elliptical, almost circular; antenna with lamella of antennomere 4 approximately twothirds as long as lamella of antennomere 5, pronotal hypomera and anterior face of profemora with a dense clothing of large white scales; body colour pale yellowish brown, length approximately
<quantity id="4CBE0EFDFFB7BB5F0A82BAA0FC4507B1" box="[920,989,386,408]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="39" unit="mm" value="21.0">21 mm</quantity>
(
<bibRefCitation id="EFD7DEE9FFB7BB5F0AF1BAA0FB1507BE" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." box="[1003,1165,386,408]" pageId="9" pageNumber="39" pagination="1819" refId="ref6289" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Melolonthinae Leach, 1819. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 467 - 507." type="book chapter" year="2019">
Weir
<emphasis id="B9327F0AFFB7BB5F0D38BAA1FBCD07BE" box="[1058,1109,386,408]" italics="true" pageId="9" pageNumber="39">et al.</emphasis>
2019
</bibRefCitation>
, Plate 64K;
<bibRefCitation id="EFD7DEE9FFB7BB5F0C1DBAA0FA0107B1" author="Allsopp, P. G." box="[1287,1433,386,408]" pageId="9" pageNumber="39" pagination="451 - 486" refId="ref5561" refString="Allsopp, P. G. (2020 b) Clarification of the status of the types of Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae) described before 1950. Zootaxa, 4885 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4885.4.1" type="journal article" year="2020">Allsopp 2020b</bibRefCitation>
, Figs. 6062); parameres apical third very constricted in ventral view, as in
<figureCitation id="137DBF9DFFB7BB5F0AAABABDFC72079C" box="[944,1002,415,437]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="9" pageNumber="39">Fig. 9</figureCitation>
; northeastern
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...................................................................................
<taxonomicName id="4C46D89BFFB7BB5F0D3DBA9FFA0407FA" authority="(Lea, 1919)" baseAuthorityName="Lea" baseAuthorityYear="1919" box="[1063,1436,445,467]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="B9327F0AFFB7BB5F0D3DBA9FFABE07FB" box="[1063,1318,445,466]" italics="true" pageId="9" pageNumber="39">Pseudholophylla cavifrons</emphasis>
(
<bibRefCitation id="EFD7DEE9FFB7BB5F0C29BA9FFA0D07FA" author="Lea, A. M." box="[1331,1429,445,467]" pageId="9" pageNumber="39" pagination="166 - 261" refId="ref6033" refString="Lea, A. M. (1919) Notes on some miscellaneous Coleoptera with descriptions of new species. Part V. Transactions of the Royal Society of South Australia, 43, 166 - 261." type="journal article" year="1919">Lea, 1919</bibRefCitation>
)
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</paragraph>
<paragraph id="8BF9A318FFB7BB5F098DBAF9FA040402" blockId="9.[151,1437,151,701]" pageId="9" pageNumber="39">
<keyLead id="30B7002DFFB7BB5F098DBAF9FA040402" pageId="9" pageNumber="39">- Scales on frons and pronotum elongate and acutely pointed; antenna with lamella of antennomere 4 as long as lamella of antennomere 5; scales on pronotal hypomera and anterior face of profemora acutely pointed, almost seta-like; parameres with apices linear or tapering but not constricted in ventral view.......................................................... 5</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BF9A318FFB7BB5F098DB910FA0404AA" blockId="9.[151,1437,151,701]" pageId="9" pageNumber="39">
<keyLead id="30B7002DFFB7BB5F098DB910FA0404AA" pageId="9" pageNumber="39">
5 Antenna with antennomere 3 broadly triangular; upper surface of clypeus very sparsely punctate; posterior margin of pronotum with a sparse fringe of fine, erect, pale setae; parameres as in
<figureCitation id="137DBF9DFFB7BB5F0A2AB96DFCEE044C" box="[816,886,591,613]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="9" pageNumber="39">Fig. 10</figureCitation>
; unknown distribution.........................................................................................
<taxonomicName id="4C46D89BFFB7BB5F0ACEB94FFA0404AA" authority="(Burmeister, 1855)" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[980,1436,621,643]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFB7BB5F0ACEB94FFB7804AB" box="[980,1248,621,642]" italics="true" pageId="9" pageNumber="39">Pseudholophylla furfuracea</emphasis>
(
<bibRefCitation id="EFD7DEE9FFB7BB5F0DF7B94FFA0D04AA" author="Burmeister, H. C. C." box="[1261,1429,621,643]" pageId="9" pageNumber="39" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BF9A318FFB7BB5F098DB9A9FA040494" blockId="9.[151,1437,151,701]" pageId="9" pageNumber="39">
<keyLead id="30B7002DFFB7BB5F098DB9A9FA040494" pageId="9" pageNumber="39">
- Antenna with antennomere 3 elongate; upper surface of clypeus densely punctate; posterior margin of pronotum without fringe of erect setae; parameres as in
<figureCitation id="137DBF9DFFB7BB5F0B12B985FDD60494" box="[520,590,679,701]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="9" pageNumber="39">Fig. 12</figureCitation>
; northwestern
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.............
<taxonomicName id="4C46D89BFFB7BB5F0D26B985FA040494" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" box="[1084,1436,679,701]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="B9327F0AFFB7BB5F0D26B985FA8E0494" box="[1084,1302,679,701]" italics="true" pageId="9" pageNumber="39">Pseudholophylla soror</emphasis>
<bibRefCitation id="EFD7DEE9FFB7BB5F0C06B985FA040494" author="Britton, E. B." box="[1308,1436,679,701]" pageId="9" pageNumber="39" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
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